Moose

Naming

"Alces alces" is called a "moose" in North American English, but an "elk" in British English. The word "elk" in North American English refers to a completely different species of deer, "Cervus canadensis", also called the wapiti. A mature male moose is called a bull, a mature female a cow, and an immature moose of either sex a calf.

Distribution

Europe and Asia:
⤷ "Finland" : In 2009, there was a summer population of 115,000.
⤷ "Norway" : In 2009, there were a winter population of around 120,000. In 2015 31,131 moose were shot. In 1999, a record number of 39,422 moose were shot.
⤷ "Latvia" : in 2015, there were 21,000.
⤷ "Estonia" : 13,260
⤷ "Lithuania" : around 14,000 in 2016
⤷ "Poland" : 2,800
⤷ "Czech Republic" : maximum of 50
⤷ "Russia" : In 2007, there were approximately 600,000.
⤷ "Sweden" : Summer population is estimated to be 300,000–400,000. Around 100,000 are shot each fall. About 10,000 are killed in traffic accidents yearly.

Habitat

Moose require habitat with adequate edible plants, cover from predators, and protection from extremely hot or cold weather. Moose travel among different habitats with the seasons to address these requirements. Moose are cold-adapted mammals with thickened skin, dense, heat-retaining coat, and a low surface:volume ratio, which provides excellent cold tolerance but poor heat tolerance. Moose survive hot weather by accessing shade or cooling wind, or by immersion in cool water. In hot weather, moose are often found wading or swimming in lakes or ponds. When heat-stressed, moose may fail to adequately forage in summer and may not gain adequate body fat to survive the winter. Also, moose cows may not calve without adequate summer weight gain. Moose require access to both young forest for browsing and mature forest for shelter and cover. Forest disturbed by fire and logging promotes the growth of fodder for moose. Moose also require access to mineral licks, safe places for calving and aquatic feeding sites.

Moose avoid areas with little or no snow as this increases the risk of predation by wolves and avoid areas with deep snow, as this impairs mobility. Thus, moose select habitat on the basis of trade-offs between risk of predation, food availability, and snow depth. With reintroduction of bison into boreal forest, there was some concern that bison would compete with moose for winter habitat, and thereby worsen the population decline of moose. However, this does not appear to be a problem. Moose prefer sub-alpine shrublands in early winter, while bison prefer wet sedge valley meadowlands in early winter. In late winter, moose prefer river valleys with deciduous forest cover or alpine terrain above the tree line, while bison preferred wet sedge meadowlands or sunny southern grassy slopes.

Reproduction

Moose are mostly diurnal. They are generally solitary with the strongest bonds between mother and calf. Although moose rarely gather in groups, there may be several in close proximity during the mating season.

Rutting and mating occurs in September and October. During the rut, mature bulls will cease feeding completely for a period of approximately two weeks; this fasting behavior has been attributed to neurophysiological changes related to redeployment of olfaction for detection of moose urine and moose cows. The males are polygamous and will seek several females to breed with. During this time both sexes will call to each other. Males produce heavy grunting sounds that can be heard from up to 500 meters away, while females produce wail-like sounds. Males will fight for access to females. Initially, the males assess which of them is dominant and one bull may retreat, however, the interaction can escalate to a fight using their antlers.

Female moose have an eight-month gestation period, usually bearing one calf, or twins if food is plentiful, in May or June. Twinning can run as high as 30% to 40% with good nutrition Newborn moose have fur with a reddish hue in contrast to the brown appearance of an adult. The young will stay with the mother until just before the next young are born. The life span of an average moose is about 15–25 years. Moose populations are stable at 25 calves for every 100 cows at 1 year of age. With availability of adequate nutrition, mild weather, and low predation, moose have a huge potential for population expansion.

Calves nursing in spring.
File:Cowcalflyingdown.JPG|Calves stay near their mothers at all times.
File:Ninemomoose.JPG|This calf is almost ready to leave its mother.
File:Mainstmoose.JPG|This yearling was probably recently chased away by its pregnant mother.

Food

The moose is a browsing herbivore and is capable of consuming many types of plant or fruit. The average adult moose needs to consume 40.9 megajoules per day to maintain its body weight. Much of a moose's energy is derived from terrestrial vegetation, mainly consisting of forbs and other non-grasses, and fresh shoots from trees such as willow and birch. These plants are rather low in sodium, and moose generally need to consume a good quantity of aquatic plants. While much lower in energy, aquatic plants provide the moose with its sodium requirements, and as much as half of their diet usually consists of aquatic plant life. In winter, moose are often drawn to roadways, to lick salt that is used as a snow and ice melter. A typical moose, weighing 360 kg, can eat up to 32 kg of food per day.

Moose lack upper front teeth, but have eight sharp incisors on the lower jaw. They also have a tough tongue, lips and gums, which aid in the eating of woody vegetation. Moose have six pairs of large, flat molars and, ahead of those, six pairs of premolars, to grind up their food. A moose's upper lip is very sensitive, to help distinguish between fresh shoots and harder twigs, and is prehensile, for grasping their food. In the summer, moose may use this prehensile lip for grabbing branches and pulling, stripping the entire branch of leaves in a single mouthful, or for pulling forbs, like dandelions, or aquatic plants up by the base, roots and all. A moose's diet often depends on its location, but they seem to prefer the new growths from deciduous trees with a high sugar content, such as white birch, trembling aspen and striped maple, among many others. To reach high branches, a moose may bend small saplings down, using its prehensile lip, mouth or body. For larger trees a moose may stand erect and walk upright on its hind legs, allowing it to reach branches up to 4.26 meters or higher above the ground.

Moose also eat many aquatic plants, including lilies and pondweed. Moose are excellent swimmers and are known to wade into water to eat aquatic plants. This trait serves a second purpose in cooling down the moose on summer days and ridding itself of black flies. Moose are thus attracted to marshes and river banks during warmer months as both provide suitable vegetation to eat and water to wet themselves in. Moose have been known to dive over 5.5 metres to reach plants on lake bottoms, and the complex snout may assist the moose in this type of feeding. Moose are the only deer that are capable of feeding underwater. As an adaptation for feeding on plants underwater, the nose is equipped with fatty pads and muscles that close the nostrils when exposed to water pressure, preventing water from entering the nose. Other species can pluck plants from the water too, but these need to raise their heads in order to swallow.

Moose are not grazing animals but browsers. Like giraffes, moose carefully select foods with less fiber and more concentrations of nutrients. Thus, the moose's digestive system has evolved to accommodate this relatively low-fiber diet. Unlike most hooved, domesticated animals, moose cannot digest hay, and feeding it to a moose can be fatal. The moose's varied and complex diet is typically expensive for humans to provide, and free-range moose require a lot of forested hectarage for sustainable survival, which is one of the main reasons moose have never been widely domesticated.Moose are hunted as a game species in many of the countries where they are found. Moose meat tastes, wrote Henry David Thoreau in "The Maine Woods", "like tender beef, with perhaps more flavour; sometimes like veal". While the flesh has protein levels similar to those of other comparable red meats, it has a low fat content, and the fat that is present consists of a higher proportion of polyunsaturated fats than saturated fats.

Predators

A full-grown moose has few enemies except Siberian tigers which regularly prey on adult moose, but a pack of gray wolves can still pose a threat, especially to females with calves. Brown bears are also known to prey on moose of various sizes and are the only predator besides the wolf to attack moose both in Eurasia and North America. However, brown bears are more likely to take over a wolf kill or to take young moose than to hunt adult moose on their own. American black bears and cougars can be significant predators of moose calves in May and June and can, in rare instances, prey on adults. Wolverine are most likely to eat moose as carrion but have killed moose, including adults, when the large ungulates are weakened by harsh winter conditions. Killer whales are the moose's only known marine predator as they have been known to prey on moose swimming between islands out of North America's Northwest Coast, however, there is at least one recorded instance of a moose preyed upon by a Greenland shark.

In some areas, moose are the primary source of food for wolves. Moose usually flee upon detecting wolves. Wolves usually follow moose at a distance of 100 to 400 meters, occasionally at a distance of two to three kilometers. Attacks from wolves against young moose may last seconds, though sometimes they can be drawn out for days with adults. Sometimes, wolves will chase moose into shallow streams or onto frozen rivers, where their mobility is greatly impeded. Moose will sometimes stand their ground and defend themselves by charging at the wolves or lashing out at them with their powerful hooves. Wolves typically kill moose by tearing at their haunches and perineum, causing massive blood loss. Occasionally, a wolf may immobilise a moose by biting its sensitive nose, the pain of which can paralyze a moose. Wolf packs primarily target calves and elderly animals, but can and will take healthy, adult moose. Moose between the ages of two and eight are seldom killed by wolves. Though moose are usually hunted by packs, there are cases in which single wolves have successfully killed healthy, fully-grown moose.

Research into moose predation suggests that their response to perceived threats is learned rather than instinctual. In practical terms this means moose are more vulnerable in areas where wolf or bear populations were decimated in the past but are now rebounding. These same studies suggest, however, that moose learn quickly and adapt, fleeing an area if they hear or smell wolves, bears, or scavenger birds such as ravens.

Moose are also subject to various diseases and forms of parasitism. In northern Europe, the moose botfly is a parasite whose range seems to be spreading.