LEE KONG CHIAN NATURAL HISTO RY MUSEUM
2021
THE MARINE AND
ESTUARINE MOLLUSCS
OF THAILAND
FRED E. WELLS, KITITHORN SANPANICH, SIONG KIAT TAN &
TEERAPONG DUANGDEE
LEE KONG CHIAN NATURAL HISTORY MUSEUM
2021
THE MARINE AND
ESTUARINE MOLLUSCS
OF THAILAND
FRED E. WELLS, KITITHORN SANPANICH, SIONG KIAT TAN &
TEERAPONG DUANGDEE
The Marine and Estuarine Molluscs of Thailand
By Fred E. Wells1,2, Kitithorn Sanpanich3, Siong Kiat Tan4 & Teerapong Duangdee5
Published by:
Lee Kong Chian Natural History Museum
National University of Singapore
2 Conservatory Drive
Singapore 117377
REPUBLIC OF SINGAPORE
Website: https://lkcnhm.nus.edu.sg/
Email: nhmvisit@nus.edu.sg
Editor:
Copyeditor:
Typesetter:
Ng Ting Hui
Cai Peixuan
Cai Peixuan
ISBN:
DOI:
978-981-14-7613-6 (online)
10.26107/LKCNHM-EBOOK-2021-0001
© 2021 Lee Kong Chian Natural History Museum
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in
any form, or by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior permission
of the copyright holder. For information regarding permission(s), please write to: nhmvisit@nus.edu.sg
Cover photograph © Kitithorn Sanpanich
1School
of Molecular and Life Sciences, Curtin University, GPO Box U1987, Perth, Western Australia 6845
Integrative Research Center, Field Museum of Natural History, Chicago, Illinois 60605 USA
3Institute of Marine Science, Burapha University, Tambon Saensook, Amphur Moengchonburi, Chonburi, 20131 Thailand
4Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377,
Republic of Singapore
5Department of Marine Science, Faculty of Fisheries, Kasetsart University, Bangkok, 10900 Thailand
Email: fred.wells@curtin.edu.au (FEW); kitithorn2504@gmail.com (KS); nhmtsk@nus.edu.sg (SKT) & tdku@
hotmail.com (TD)
2Negaunee
II
CONTENTS
DEDICATION ............................................................................................................................................................................................................. 1
ABSTRACT.................................................................................................................................................................................................................. 2
INTRODUCTION ...................................................................................................................................................................................................... 3
MATERIAL AND METHODS ................................................................................................................................................................................ 4
THE MARINE AND ESTUARINE MOLLUSCS OF THAILAND ................................................................................................................ 6
LIST OF THAI MARINE AND ESTUARINE MOLLUSCS ......................................................................................................................... 16
CLASS BIVALVIA ............................................................................................................................................................................................. 16
CLASS CEPHALOPODA ................................................................................................................................................................................. 68
CLASS GASTROPODA .................................................................................................................................................................................... 75
CLASS POLYPLACOPHORA ..................................................................................................................................................................... 164
CLASS SCAPHOPODA ................................................................................................................................................................................. 164
LITERATURE CITED ........................................................................................................................................................................................ 166
ACKNOWLEDGEMENTS ................................................................................................................................................................................. 186
HOW TO CITE THIS BOOK ............................................................................................................................................................................ 186
APPENDIX ............................................................................................................................................................................................................ 187
III
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
DEDICATION
We are pleased to dedicate this book to the late Jørgen Hylleberg of the University of Aarhus, Denmark, for the
outstanding service he provided to Thai malacology during his quarter of a century in Thailand as the Danish
Advisor to the Phuket Marine Biological Center and as the organiser of the Tropical Marine Mollusc Programme.
1
The Marine and Estuarine Molluscs of Thailand
ABSTRACT
Thailand is a large Southeast Asian country on the margin of the biodiverse coral triangle. It has a 3,299 km
coastline on both the Gulf of Thailand in the Pacific Ocean and the Andaman Sea in the Indian Ocean. An extensive
literature and internet search documented over 15,000 records of 3,359 species of Thai marine and estuarine
molluscs. Records of 137 species are considered to be incorrect or dubious as their known ranges do not include
Southeast Asia. The remaining 3,222 species include 1,963 gastropods, 1,116 bivalves, 129 cephalopods, seven
polyplacophorans and seven scaphopods; no records were found of monoplacophorans or aplacophorans. Most
records cannot be verified through specimens lodged in museums. The taxonomy of all species has been updated
to their current status. Previous names used, occurrence on the Gulf of Thailand and/or Andaman Sea coast, and
literature records and notes (if any) are provided for each species. Two invasive species, the mussels Mytilopsis
sallei (Récluz, 1849) and Mytella strigata (Hanley, 1843), are known to have been introduced to Thailand, but there
may be more introduced species that have not yet been detected. Strategies to further document the marine and
estuarine molluscs of Thailand are proposed.
2
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
INTRODUCTION
The Indo-West Pacific is the largest marine biogeographical province in the world. It spans the tropical oceans
from about 30 degrees north of the equator to about 30 degrees south, and extends across the vast regions of the
Indian and Pacific Oceans from Hawaii to the east coast of Africa. Within the Indo-Pacific, the coral triangle, from
the Philippines and Malaysia to Papua New Guinea, is the most megadiverse area of the world oceans (HoeghGuldberg, 2009). Molluscs are the “insects of the sea” with by far the greatest biodiversity of the animal groups.
While there is no complete inventory of the marine molluscs of any area, the Philippines have received the most
scientific attention. The Philippine Islands have a rich diversity of marine habitats and an accordingly broad
diversity of marine mollusc species that have long been studied, with 5,835 marine mollusc species having been
documented in the country (Poppe, 2017).
In contrast to the Philippines, the marine and estuarine molluscs of Thailand have been relatively poorly studied,
but there have been several compilations of the phylum. Early records include: Morlet (1889) with 167 species;
Lynge (1909) with 316 bivalves; and Suvatti (1937, 1938) with 270 species. Nateewathana et al. (1981) reported
548 species in the Phuket Marine Biological Center (PMBC) collection at that time; the number has grown
significantly since then. The largest compilation of Thai marine and estuarine molluscs was Nabhitabhata (2009),
with 1,267 species. Recently, BEDO (2017a–e) listed 1,270 species. Having been published over a span of nearly
130 years, there have been considerable taxonomic changes that make direct comparisons of the above lists
difficult. Clearly the marine molluscan fauna of Thailand, with a broad diversity of habitats on both the Pacific and
Indian Oceans, is much greater than indicated in the above compilations.
The goals of the present book are to first compile as exhaustive a list as is possible of the scientific information on
the marine and estuarine molluscs of Thailand, then to update the scientific names of the species recorded. The
study will provide a basis for further scientific investigations of the group in Thailand. It is emphasised that in the
absence of voucher material of most of the species in publicly accessible locations, the present paper provides only
an introduction to marine and estuarine species recorded from Thailand and identifications of many of the species
listed may be incorrect. Dayrat (2011) emphasised the risk of assuming such unverified literature records were
correct. 16 species of the nudibranch genus Discodoris were recorded in the literature from the Mediterranean Sea
until a revision of the group demonstrated that the genus is in fact absent in the area. However, the present paper
provides an entry point into our understanding of the marine and estuarine molluscs of Thailand.
3
The Marine and Estuarine Molluscs of Thailand
MATERIAL AND METHODS
An extensive literature search, both in English and Thai, was undertaken of the marine and estuarine molluscs of
Thailand, including books, journals and revisions of family and genera that might include records of species from
Thailand. The Phuket Marine Biological Center (PMBC) publishes both a Research Bulletin and Special Publication
series; all issues of each series were examined. In addition, all volumes of the Tropical Marine Mollusc Programme
are available on the PMBC website and were searched. Molluscan journals, such as Indo -Pacific Mollusca,
Monographs of Marine Mollusca, Molluscan Research and The Veliger, were also consulted. Available shell books
were also examined. An extensive internet search was conducted, including Google Scholar and open repositories,
i.e., ResearchGate and Academia.edu, using terms such as ‘Thailand’, and phrases associated with molluscs and the
various classes. Literature cited sections of papers reporting Thai marine and estuarine molluscs were used to
locate further papers. To be as inclusive as possible, grey literature was also used. The literature search
concentrated on revisions of particular families and genera and faunal surveys. Other literature, such as ecological
and physiological studies, are not cited unless they add to list of species recorded in Thailand. A recent
development is the trend for international museums to database their collections and place the information on the
web in sites such as the Global Biodiversity Information Facility (GBIF; https://www.gbif.org/). GBIF was searched
to obtain currently available Thai records. As this and other facilities develop, more information will become
available on the marine and estuarine molluscs of Thailand.
Only taxa identified to species are included in this analysis; those identified only by family or genus are disregarded
as there is no way of knowing whether species A of one author is the same as species A of another paper. The
internet search turned up numerous records of Thai shells for sale by dealers; these are transitory, unverified and
were not used.
Every species listed in all of the publications was checked against the World List of Marine Species (WoRMS;
WoRMS Editorial Board [2018]) to determine the current valid name and family placement. If a species could not
be located in WoRMS Editorial Board (2019), alternative spellings, such as changing the ending from ‘-a’ to ‘-us’ or
vice versa, and other generic placements were tried. Other information provided by WoRMS Editorial Board
(2019) was included, such as species listed as uncertain, nomen dubium, taxon inquirendum, etc. Where this was
unsuccessful, other sources were used, such as Ruhoff (1980) and other internet databases such as the Atlas of
Living Australia (ALA, 2019). Over 15,000 citations to the scientific literature on Thai molluscs are provided in the
list below.
Some freshwater species can tolerate low salinity brackish water. This is a continuum, with each species having
its own salinity tolerances. Species listed by WoRMS Editorial Board (2019) as only occurring in freshwater are
not included, but freshwater species that also inhabit brackish water are listed.
The higher-level systematics of molluscs are currently in a state of flux as new data, particularly genetic
information, is added to our knowledge. The higher taxonomy used in this book follows WoRMS Editorial Board
(2019). Because of inconsistent use across the phylum, subgenera and subspecies are not used. However,
cephalopods are an exception as many papers list cephalopod subgenera.
The following information is provided for each species: current scientific name including genus, species, author
and date; names under which the species was previously recorded in Thailand; whether the species was recorded
from the Gulf of Thailand and/or Andaman Sea coastline of Thailand; and literature references to the species in
date order and notes (if any). Most changes in species taxonomy have been transfers of a species from one genus
to another. Alternative names used for a species are shown so future researchers can trace the species through the
various publications. Misspellings in earlier references are included for the same reason. Very minor changes, such
as changes in the ending of a name from ‘-a’ or ‘-us’ or vice versa, after the gender of a species changed upon a
move to another genus, or differences in the year the species was described (i.e., 1841 versus 1842), are not shown.
However, if the time difference is substantial (i.e., 1841 versus 1860), it is included. In some cases, the author
shown in the Thai reference is not the actual author of the species; this is also indicated. Also included are cases
where the Thai reference did not give an author and date. However, full citation of each species is provided as
recorded by WoRMS Editorial Board (2019). “Reported as” indicates that the species was not reported with the
currently accepted name. “Also reported as” indicates that the species has been reported from Thailand using the
current name but has also been reported under different names.
Until very recently, the original scientific literature was essentially unavailable in Thailand, and there were few
shell books available. In most cases species were, out of necessity, identified by Thai researchers based on
photographs and descriptions in the shell books, often black-and-white photocopies of colour illustrations, and
4
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
other secondary sources. Without a knowledge of closely related species, this sometimes resulted in species being
incorrectly identified as those figured in the books. We have listed these misidentifications where they were
obvious in Appendix 1, but there are undoubtedly others. PMBC has a small but growing collection of species that
can be examined. The Thai Natural History Museum (THNHM) is also small, and there is a mollusc collection at
Kasetsart University. The rapid development of the internet and literature databases, such as the Biodiversity
Heritage Library (https://www.biodiversitylibrary.org/), has recently made large portions of the original
taxonomic literature much more available to all researchers. Future identifications should be based on the original
descriptions, if available. Revisions of families or genera are the second preference, followed by books on Thai
marine and estuarine molluscs. If books on overseas localities are used, care should be taken to ensure that the
geographic range of each species includes Thailand.
5
The Marine and Estuarine Molluscs of Thailand
THE MARINE AND ESTUARINE MOLLUSCS OF THAILAND
MARINE AND ESTUARINE HABITATS OF THAILAND
Thailand is a large country in Southeast Asia with a land area of 513,115 km 2 and a coastline of 3,299 km. It is a
peninsular country lying 5–20°N and 97–106°E. Thailand has two extensive coastlines, the eastern coast located
along the Gulf of Thailand in the Pacific Ocean, and the western coast facing the Andaman Sea in the Indian Ocean
(Fig. 1). The Thai Exclusive Economic Zone (EEZ) in the Gulf of Thailand has an area of 304,000 km2, and the EEZ
along the Andaman coast is 116,280 km2 (Chavanich et al., 2010a).
The climate of coastal Thailand is tropical, with an average annual temperature of 27.6°C, ranging from a mean of
25.5°C in January to 29.9°C in April. While the average annual rainfall is 1,556 mm, it is seasonal and there is little
rain during winter and summer. Average rainfall is highest in September (378.3 mm) and lowest in December (4.6
mm). The average annual relative humidity is 76.1%, varying from a low of 70.0% in January to a high of 81.4% in
October (Pumijumnong, 2014).
The intertidal and shallow water environments of the two coasts are similar, with extensive rocky shores, sand
beaches, tidal flats, mudflats, estuaries, coral reefs, seagrass beds and mangroves occurring on both coasts (Fig. 2).
There is an increasing number and variety of artificial habitats such as jetties, bridges, roads and aquaculture
facilities, particularly extensive prawn farms on both coasts. Bays and islands are more numerous on the Andaman
coast (Chavanich et al., 2010a). There are about 545 islands on the Andaman coast, 100 in the eastern Gulf of
Thailand and 150 in the western Gulf (Phongsuwan et al., 2013).
Fig. 1. Map of Thailand showing the locations of the Gulf of Thailand and the Andaman Sea coasts. Basemap is from Google®,
adapted by Kitithorn Sanpanich.
6
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Prao Beach, Kood Island, Trat Province
Ma Jor Bay, Sattahip, Chonburi Province
Klong Kon, Samutsongkram Province
Klouy Bay, Kood Island, Trat Province
Koh Jed, Similan Islands, Phang-nga Province
Ship port, Kuraburi, Phang-nga Province
Fig. 2. The most common marine habitats in Thailand (Photographs by: Kitithorn Sanpanich).
A key difference between the two coasts is that the Gulf of Thailand is shallow, with a mean depth of 45 m and a
maximum of 80 m. The bottom is largely mud. The Upper Gulf is a square-shaped area at the northern end, with
four major rivers emptying into it. The Chao Phraya River is the largest, with an annual average flow of 13,220 km3.
The Lower Gulf has two ridges on its southern margin that restrict water flow into and out of the South China Sea
(Wattayakorn, 2006). The Gulf is influenced by the southwest monsoon from May to September and the northeast
monsoon from October to February (Phongsuwan et al., 2013).
In contrast, there is a narrow continental shelf on the Andaman coast, up to 100 km wide, that gives way to a steep
continental slope that reaches a depth of up to 2,000 m. Intertidal rocky shores and sand beaches are found on
both coasts.
7
The Marine and Estuarine Molluscs of Thailand
There is an estimated 153 km2 of coral reefs in Thailand, with three reef types occurring: coral communities with
no true reef structure; developing fringing reefs; and early formation of fringing reefs (Yeemin et al., 2001). In the
Andaman Sea, coral reefs usually occur as fringes in clear water on the eastern sides of islands where they are
sheltered from the southwest monsoon. In the eastern Gulf of Thailand, coral reefs are most common on the east
and north coasts of islands where they are sheltered from the southwest monsoon. The northeast monsoon is more
influential in the western Gulf, and reefs there are most common on the west and south coasts. Rocky shores
dominate on the windward side of most islands (Phongsuwan et al., 2013).
As with many other parts of the world, Thai coral reefs have been heavily stressed by human activity, which
includes direct destruction of reefs, and indirect effects such as pollution and nutrient enrichment. The effects of
tourism, with trampling and inadvertent destruction of fragile reef structures, are particularly evident in Thailand.
There have been several severe bleaching events, with a substantial bleaching occurring in 2010. Several
outbreaks of the crown of thorns starfish have occurred on reefs in the Andaman Sea, particularly in the 1980s.
The Indian Ocean tsunami on 26 December 2004 significantly affected coral reefs on the Andaman coast; while
about 40% of surveyed reefs had no visible damage, 47% had low-to-moderate impacts and 13% were severely
damaged. However, surveys taken two years after the tsunami showed a slightly higher coverage of live coral than
surveys two years before the tsunami (Phongsuwan et al., 2013).
Extensive, diverse, and biologically productive (74 known mangrove species) mangrove forests are a key intertidal
habitat in Thailand. Mangroves are common on the muddy tidal flats at river mouths. There are extensive
mangroves on the Andaman coast, particularly in Phang Nha province, which has 18.5% of the total mangrove area
of Thailand. Mangroves are common along the east coast of the Gulf of Thailand and along the Chao Phraya River
delta near Bangkok. The mangrove forests often have two layers. The upper storey grows to a height of 20 m or
more, with large trees of Rhizophora, Bruguiera, Heritiera and Xylocarpus. Common understorey species are in the
genera Bruguiera and Ceriops. The palm Nypa fruticans is common along the edges of intertidal creeks
(Pumijumnong, 2014).
There has been a considerable loss of mangroves in Thailand for a variety of reasons, including logging, pollution,
etc. The primary cause of losses is the removal of mangroves to create space for shrimp farms. Levees are built to
a height of 1–1.5 m to increase the water volume available for the shrimp. Unfortunately, the ponds become
unviable after about five years and are abandoned. The levees change water flow patterns and restrict
recolonisation of the ponds by mangroves, even if they are actively planted. There were about 320,000 ha of
mangroves in Thailand in 1975, but this was halved by 1996. Conservation and rehabilitation increased the
mangrove area to about 240,000 ha by 2004 (Pumijumnong, 2014).
Seagrass habitats in Thailand have been classified into three categories: mangrove–associated beds, shallow
sandy–bottom beds and coral reef-associated beds (UNEP et al., 2005). There are 12 species of seagrasses in
Thailand in the genera Halophila, Cymodocea, Halodule, Enhalus, Thalassia, Syringodium and Ruppia. 11 species
have been recorded from both coasts; Ruppia maritima is known only from the Gulf of Thailand. The area of
seagrasses is much larger and denser on the Andaman coast (9,448 ha) than in the Gulf of Thailand (5,489 ha)
(Adulyanukosol & Poovachiranon, 2007). There is a distinct seasonality of seagrasses in Thailand. For example,
Khogkhaoa et al. (2017) studied the spatial distribution of seagrasses and their biomass in relation to a number of
physical factors near the mouth of the Trang River on the Andaman Coast. 10 species with a total area of 61 km2
were found during the dry season, nearly half (29 km 2) of which was Halophila decipiens. The other common
species were Enhalus acoroides and Halophila ovalis. Only eight species with a total area of 24 km2 were found in
the rainy season. The most important limiting factors were water depth and light intensity.
The marine algal flora is diverse, with about 326 species having been recorded: 98 chlorophytes, 56 phaeophytes
and 172 rhodophytes. The number of blue green algae has not been determined (Coppejans et al., 2010, 2017).
Marine algae are abundant in a wide range of shallow water habitats, including rocky shores and artificial hard
surfaces, and as epiphytes on seagrasses and mangroves.
HISTORY OF MOLLUSC EXPLORATION IN THAILAND
According to Hylleberg (2013), the first extensive sampling of benthic species in the Gulf of Thailand was undertaken
in early 1900 by the Danish Expedition to Siam, funded by the Danish government and the Carlsberg Foundation.
There were only two scientists, the botanist Johs Schmidt and the echinoderm specialist Theodor Mortensen. Koh
Chang Island in the eastern Gulf of Thailand was used as a base with Mortensen undertaking boat trips of up to a
week from Koh Chang for dredging and intertidal collecting. Unfortunately, with the large number of rivers in the
region, the bottom was predominately mud and Mortensen had considerable difficulty in locating suitable habitats
for dredging. Specimens collected were deposited in the Zoological Museum, Copenhagen. Bergh (1902) published
on opisthobranchs and Lynge (1909) reported on bivalve molluscs collected during the expedition.
8
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Three additional Thai–Danish expeditions to Thailand were undertaken in the following decades, but these were
primarily botanical. The Fifth Thai–Danish Expedition in January to March 1966 was originally conceived as a
botanical expedition, but in 1961, Germany had initiated successful trawl fisheries in the Gulf of Thailand.
Therefore, the Thai Department of Fisheries was interested in exploring the fisheries resources of the Andaman
coast and made their new research vessel M/S DHANARAJATA available for the expedition. The survey evolved
into a combined terrestrial botanical and marine biological expedition that included 15 Thai scientists and
students from the departments of Fisheries and Agriculture in Chulalongkorn University, and four Danish
scientists and representatives from the UNESCO office in Singapore, Singapore Botanic Gardens and the Allan
Hancock Foundation in California. The eminent Danish scientist Gunnar Thorson led the marine biological
component of the expedition. Its goals were: 1) research work on fish-food animals on sediment bottoms from a
quantitative point of view; 2) training of groups of young Thai marine biologists; and 3) the creation of a broad
reference collection of marine fauna for use by the Thai Marine Fisheries Laboratory in the future education of
marine and fishery biologists (Seidenfaden et al., 1968).
A key result of the expedition was the development of a concept for the creation of the Phuket Marine Biological
Center. The Thai–Danish team examined 10 sites on Phuket Island for the possible construction of a marine
laboratory. The variety of habitats that could be reached by boat or vehicle within 30 minutes, including coral
reefs, rocky shores, open and sheltered sandy beaches and mangroves, led to the present site of PMBC being
chosen. An agreement for the development of PMBC was signed by the Royal Thai and Danish governments in
October 1968 (Seidenfaden et al., 1968).
The Thai government paid for the buildings and other infrastructure while the Danish contribution was scientific
equipment, staff training, and expert assistance. Jørgen Hylleberg was appointed as the Danish Advisor, a role that
he fulfilled from 1979 to 1994. Hylleberg then became project director for the new Tropical Marine Mollusc
Programme (TMMP) and continued to advise PMBC until TMMP funding was ended by the Danish International
Development Agency (DANIDA) in 2004 (Hylleberg, 2013).
The creation of PMBC opened a new era in Thai marine research that was primarily focused on discovering the
biodiversity of marine and estuarine biota on the Andaman coast; much of the work was on molluscs. A new
building was opened in 1983 to house the rapidly growing Reference Collection. PMBC started two journals, the
Research Bulletin and Special Publications, both of which contain a significant number of papers on molluscs. The
two series provided venues for PMBC and other scientists, from both Thailand and overseas, to publish scientific
information on the Thai marine environment. The journals are sent in exchange for publications from other
national and international institutions, thus enhancing the PMBC library (Hylleberg, 2013).
Facilitating the education of young scientists has been a key PMBC outcome. A number of field courses have been
presented at PMBC and PMBC staff have been able to participate in overseas opportunities. Particularly successful
has been the ability of Thai scientists to obtain advanced degrees in Denmark. Mollusc workers include Charatsee
Aungtonya, Anuwat Nateewathana and Vararin Vongpanich with MSc degrees from the University of Aarhus;
Anuwat Nateewathana continued on to get a PhD from the same university. Charatsee Aungtonya obtained a PhD
at the University of Copenhagen. The transfer of students worked both ways, with students from the University of
Aarhus undertaking their field studies at PMBC. The mollusc workers among these include Michael Bech, Britt
Gribsholt and Peter Middlefart (Hylleberg, 2013).
The TMMP, coordinated by Jørgen Hylleberg, was established in 1991 as a joint research activity between Thailand
and other Southeast Asian countries. Annual workshops were held, which allowed participants to present papers
and interact with other participants in a week-long conference, then to undertake field excursions. Denmark, India
and Thailand participated in the first year of the TMMP. The first conference was devoted to the possible
aquaculture of the large Murex ramosus. Indonesia joined in 1992. Over time the number of participants and the
range of scientific topics grew rapidly. The quantity of biodiversity information presented increased so much that
from 1997 onwards an entire volume of the workshop proceedings was devoted to biodiversity studies. Each
annual workshop was held in a different Southeast Asian country, increasing the ability of scientists in the region
to participate. The opportunity to meet at a workshop and exchange scientific information led to numerous
cooperative projects among the participants. By the time the programme was terminated in 2004, the TMMP had
published 11 sets of proceedings and two monographs. Mollusc researchers from 57 institutions in 22 countries
had participated in the various workshops. Research presented concentrated on the Indian Ocean and South China
Sea in Thailand, Cambodia, Vietnam, India and Indonesia. A number of mollusc experts from other Asian countries,
Europe, South Africa, United States and Australia also participated in the TMMP, considerably broadening the
TMMP outlook (Hylleberg, 2013).
An ambitious project, Biodiversity of the Andaman Sea Shelf (BIOSHELF), was undertaken from 1996 to 2000 on
the Andaman coast of Thailand to survey marine biodiversity along the entire Thai coastline from the Myanmar
9
The Marine and Estuarine Molluscs of Thailand
border in the north to Malaysia in the south. Extensive benthic surveys were made using a variety of grabbing and
trawling techniques down to 1,000 m and open water species to deeper dept hs. A total of 98 stations were
conducted on 12 transects between 40 and 900 m, and an additional 16 stations in the Thai EEZ. 19 nearshore
stations were made (Aungtonya et al., 2000; Tendal et al., 2014). Specimens collected were deposited in the
PMBC Reference Collection. Extensive reports were compiled on the crustaceans (Bruce et al., 2002) and
polychaetes (Eibye–Jacobsen, 2002), but only limited information on molluscs was published: Schwabe (2006) on
chitons and Jensen (2007) on opisthobranchs.
In addition to the Danish work in Thailand cited above, there have been a number of major oceanographic
expeditions that have worked in Thai waters for at least part of their extensive voyages. Again, Danes played a
leading role. The Thai component of these expeditions was summarised by Jensen & Knudsen (2001). The DANA
circumnavigated the world from 1928 to 1930. The expedition was focused on plankton, fish and oceanography.
It was in Thai waters in April and May 1929. The expedition was led by the Danish scientists Johs Schmidt and
Anton Bruun. The GALATHEA expedition, undertaken from 1950 to 1952, was led again by Anton Bruun. It worked
in the Gulf of Thailand in June 1951. Unfortunately, the focus of the expedition was on the deep sea, so the Gulf of
Thailand molluscs were not included. The Naga Expedition, also led by Anton Bruun, worked in Thailand in 1959.
Included among the scientists was the prominent malacologist Jørgen Knudsen.
More recently, a workshop on the systematics, anatomy and natural history of shallow water marine bivalves was
held in Kungkrabaen, Chanthaburi Province, in August and September 2005 (Bieler et al., 2008a). The primary
goal was to increase scientific knowledge of marine bivalves in the eastern Gulf of Thailand, but associated work
was also undertaken on gastropods. Thai and international scientists conducted field studies in Kungkrabaen for
two weeks, then returned to their home institutions to complete the associated laboratory studies and write their
papers describing the results. A special volume of the Raffles Bulletin of Zoology contains the workshop results in
17 papers (Bieler et al., 2008b).
Since the completion of the Kungkrabaen workshop, primary development of scientific information on Thai marine
molluscs has resulted from short-term studies of particular areas such as marine parks. Most of the work has been
undertaken by Kitithorn Sanpanich of Burapha University and Teerapong Duangdee of Kasetsart University.
OVERVIEW OF SPECIES RECORDED
The present compilation presents records of 3,359 marine and estuarine species in Thailand. 137 species are
rejected or considered dubious: 105 on geographic grounds because the species does not occur in the Southeast
Asian marine biogeographic region (Appendix 1); 26 on a taxonomic basis; four for being fossils; and two for being
freshwater species. The remaining 3,222 species (Table 1; Fig. 3) are considered valid by WoRMS Editorial Board
(2019) and other sources. The compilation provides an up-to-date understanding of our knowledge of the marine
and estuarine molluscs of Thailand, and a basis for future research on the phylum in Thailand. However, individual
records should be treated with scepticism as most cannot be verified by examination of the original specimens and
compared with other material. The PMBC collections are an exception as they are available, as are the collections
at Kasetsart University and THNHM.
Table 1. Number of marine and estuarine species in each class of mollusc recorded from Thailand.
Class/Subclass
Subclass Heterodonta
Subclass Protobranchia
Subclass Pteriomorpha
Total Class Bivalvia
Subclass Nautiloidea
Subclass Coeleoidea
Total Class Cephalopoda
Subclass Caenogastropoda
Subclass Heterobranchia
Subclass Neritimorpha
Subclass Patellogastropoda
Subclass Vetigastropoda
Total Class Gastropoda
Total Class Polyplacophora
Total Class Scaphopoda
Total Phylum Mollusca
Gulf of Thailand
only
246
11
88
345
0
12
12
264
81
8
2
43
398
0
5
760
Andaman Sea
only
130
2
64
196
0
36
36
463
111
13
3
48
638
5
1
875
10
Both coasts
358
4
181
543
1
80
81
621
195
22
9
52
899
2
1
1526
No specific
locality
24
0
8
32
0
0
0
9
13
4
1
1
28
0
0
60
Total species
758
17
341
1116
1
128
129
1357
400
47
15
144
1963
7
7
3222
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Cypraea tigris Linnaeus, 1758:
Kood Island, Trat Prov.
Cassis cornuta (Linnaeus, 1758):
Koh Huyong, Similan Islands National Marine Park
Spondylus varius G. B. Sowerby I, 1827:
Koh Kata, Tarutao National Marine Park
Chicoreus torrefactus (G. B. Sowerby II, 1841):
Koh Rorknai, Lanta Islands, Krabi Prov.
Epidendrium billeeanum (DuShane & Bratcher, 1965):
Koh Ha Yai, Lanta Islands, Krabi Prov.
Lambis scorpius (Linnaeus, 1758):
Koh Pa chumba, Surin Islands National Park, Ranong Prov.
Pteria penguin (Röding, 1798):
Koh Kumpun, Trat Prov.
Phenacovolva rosea (A. Adams, 1854):
Hin Plai Nam, Chang Islands, Trat Prov.
Fig. 3. Examples of marine mollusc species recorded in Thailand (Photographs by: Kitithorn Sanpanich).
11
The Marine and Estuarine Molluscs of Thailand
The 3,222 species recorded from Thailand are obviously an incomplete representation of the phylum in Thailand.
Most of the species recorded are gastropods (1,963) and bivalves (1,116) (Table 1), but even in these groups, there
are undoubtedly many species that have yet to be recorded. Because of their economic importance, cephalopods
are the best-studied group in the country; we found literature references to 129 species. However, there are still
more species to be recorded. Polyplacophorans (7) and scaphopods (7) are poorly studied; no records were
encountered of monoplacophorans or aplacophorans.
INTRODUCED SPECIES OF MARINE AND ESTUARINE MOLLUSCS
There is an increasing awareness worldwide of the numbers of marine species being introduced from one part of
the oceans to another. For example, over 250 species are known to have been introduced into Australia (NIMPIS,
2017). Most of these species are apparently innocuous, causing no harm in their new environment. However, a
small proportion become invasive marine species (IMS) that can cause considerable environmental and economic
damage by changing ecosystems, outcompeting native species, threatening fisheries and aquaculture, introducing
diseases and interfering with industrial facilities (Johnson & Chapman, 2007; Molnar et al., 2008; Wells et al., 2009;
Crowe & Frid, 2015). Of the 250 species introduced into Australian waters, only about a dozen are regarded as IMS
(NIMPCG, 2009a, 2009b; NIMPIS, 2017).
There has been no comprehensive examination of introduced marine and estuarine species in Thailand.
Understanding the Thai situation is made more difficult because of the many centuries that wooden boats were
used to transport goods and people within Southeast Asia and beyond. Species may have been introduced
hundreds of years ago that now appear to be native. The problem has been exacerbated in recent decades by the
globalisation of shipping, which allows vessels to rapidly move from one area of the world’s oceans to another.
Thailand has two major ports that can serve as primary recipient areas for introduced marine species. Both are in
the inner Gulf of Thailand. Laem Chabang is the larger. It handled 13,313 vessels in 2017 and has extensive support
facilities such as drydocks and a floating drydock (LCP, 2018). In fiscal year 2016, Bangkok Port handled 3,067
vessels with 21 million tonnes of cargo (PAT, 2020). There are also other ports that handle smaller numbers of
international vessels.
In a survey of introduced marine species in Southeast Asia, Chavanich et al. (2010a) recorded only seven
introduced species in Thailand, but emphasised that there has been no survey conducted in the country. The low
number known from Thailand is probably a reflection of the lack of study, but also may reflect a relatively low
number of introductions.
Only one of the seven species reported in Thailand by Chavanich et al. (2010a) was a mollusc, the false mussel
Mytilopsis sallei (Récluz, 1849) (reported as Mytilopsis adamsi Morrison, 1946), which has been recorded in the
Songkla Lakes in the south of the country (Wangkulangkul & Lheknim, 2008; Wangkulangkul, 2018). Mytilopsis
sallei has been reported in numerous Asian countries, including India (Kalyanasundaram, 1975), Japan (Habe,
1980; Furuse & Hasegawa, 1984), Hong Kong (Morton, 1980, 1989; Huang & Morton, 1983), Taiwan (Chang, 1985),
Singapore (Tan & Morton, 2008) and China (Cai et al., 2014). Wangkulangkul (2018) recently surveyed the
population of Mytilopsis sallei in the Songkla Lakes and found the species to be well established. However, it is
most abundant in the rivers flowing into the lake where the water is about 15 parts per thousand (ppt). It is much
less common in the higher-salinity parts of the lakes system. The fact that Mytilopsis sallei became established in
Songkla Lakes emphasises the point that concern about IMS in Thailand is not restricted to the major ports of
Bangkok and Laem Chabang. Sanpanich & Wells (2019) recently reported that the species has now been found in
the inner Gulf of Thailand, again in lower-salinity areas.
Chavanich et al. (2010a) did not record the Central American mussel Mytella strigata (Hanley, 1843) (previously
known as Mytella charruana [d’Orbigny, 1842]) in any Southeast Asian country. However, it has recently become
invasive in the Philippines (Rice et al., 2016; Mediodia et al., 2017; Vallejo, 2017), Singapore (Lim et al., 2018) and
India (Jayachandran et al., 2019). Sanpanich & Wells (2019) surveyed the inner Gulf of Thailand and found the
species to be widespread, with densities of up to 40,000 m −2. In Singapore, Lim et al. (2018) expressed concern
that the species might be outcompeting the economically important Asian green mussel, Perna viridis (Linnaeus,
1758), a concern shared for the Thailand populations by Sanpanich & Wells (2019). Sanpanich & Wells (2019)
reported anecdotal information that Perna viridis is being grown in aquaculture farms in the Phuket area on the
Andaman coast. The species is uncommon in the region, and seed stock are being transferred from Sri Racha on
the Gulf of Thailand to Phuket, providing a direct mechanism for the possible introduction of Mytella strigata to
the Andaman coast.
Two other bivalve species have been introduced to Asia that should be mentioned; neither has yet been recorded
in Thailand. The mussel Brachidontes striatulus (Hanley, 1843) was introduced to Singapore over a decade ago.
12
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Densities of up to 75 per 100 cm2 were found in salinities ranging from 12 to 20 ppt (Morton & Tan, 2006). The
Australian mussel Xenostrobus securis (Lamarck, 1819) has been introduced to European waters and also Japan
(Kohama et al., 2001; Iwasaki & Yamamoto, 2014) and Hong Kong (Morton & Leung, 2015).
Brachidontes pharaonis (Fischer, 1870) is a small mytilid mussel whose native range is the western Indian Ocean
and Red Sea. However, in 1876, just seven years after the Suez Canal was opened, the species was found at Port
Said on the Mediterranean coast of Egypt (Dogan et al., 2007). Brachidontes pharaonis was first discovered along
the Israeli coast in 1937, and was still rare in the early 1970s. However, a survey 30 years later found dense
populations on rocks, where it was the dominant species (Galil, 2000). It has become invasive in many parts of the
Mediterranean (Terranova et al., 2006; Cilia & Deidun, 2012). The taxonomy of the species has been confused by
the close resemblance of the shells of Brachidontes pharaonis to those of the Indo-Pacific Brachidontes variabilis
(Krauss, 1848). Lamy (1936) and Huber (2010) considered the two to be conspecific and recognised the proper
name as Brachidontes pharaonis. This was followed in Thailand by Swennen et al. (2001) and Nabhitabhata (2009)
(see Appendix 1). Brachidontes variabilis has also been reported in Thailand by Nateewathana et al. (1981),
Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009) and BEDO (2017b). However, GBIF (2019)
regards the two as separate species and Brachidontes variabilis as the native Thai species.
Two of the species in Appendix 1 warrant further consideration for possibly occurring in Thailand. The eastern
American mud snail Tritia obsoleta (Say, 1822) has been introduced into California, where it is very abundant in
San Francisco Bay. It was possibly introduced in association with the transplantation of oysters for aquaculture
(Carlton, 1992; GISD, 2015a). There is no evidence of Tritia obsoleta in Thailand, and it is likely to have been
misidentified.
The European oyster Ostrea edulis (Linnaeus, 1758) has been introduced into a number of countries, including:
Australia, Canada, Fiji, Israel, Japan, Mauritius, Namibia, New Zealand, South Africa, Tonga and the United States
(Carlton, 1992; GISD, 2015b). It has been reported from Thailand by Suvatti (1937, 1938) and Duangdee et al.
(2011) (Appendix 1), but was not included by Bussarawit & Cedhagen (2010) in their revision of Thai oysters. The
species has been reported from southwestern Australia using genetic techniques (Morton et al., 2003). A genetic
analysis would be required to demonstrate Ostrea edulis is present in Thailand.
FISHERIES IMPORTANCE OF THAI MOLLUSCS
Thailand is one of the largest producers of seafood products in the world. It is an important industry for the
country. Over 30 species of molluscs, predominately cephalopods, are important in the commercial catch. The
production can be divided into wild-caught fisheries and aquaculture.
Derrick et al. (2017) reported that Thai catches from Thailand’s EEZ increased from approximately 400,000 tonnes
in 1950 to 2.6 million tonnes in 1987. The catch then declined to 1.7 million tonnes in 2014. However, Derrick et al.
(2017) analysed the catch statistics and concluded that because of under reporting the actual catch may have been
three times greater.
A survey in 2000 reported that 58,119 fishing boats were in use in Thailand. Most (80%) were small local boats;
many were at or near the subsistence level. 20% of the boats were >5 tonnes and capable of undertaking voyages
ranging upwards from a full day to several weeks. About 60% of the catch was made in Thai waters (41% from the
Gulf of Thailand and 19% from the Andaman Sea). The total catch in 2007 was 2.2 million tonnes valued at 63
billion Baht (about USD $2 billion). Cephalopods contributed 6% of the total 2007 fisheries catch, while other
molluscs were 1% (FAO, 2019). Not included in this survey was hand collecting along the shoreline, much of which
was for molluscs, particularly bivalves.
Ngamprasertkit (2018) reported that wild-caught squid and cuttlefish in 2017 were worth USD $345 million.
Other molluscs added another USD $374 million. The total wild catch in 2015 was 1.32 million metric tonnes, with
squid contributing 91,000 tonnes and other molluscs 16,700 tonnes. Squids, cuttlefishes and octopuses are all
commercially exploited in Thailand. The key squid are four species of Loligo, Loliolus sumatrensis and Sepioteuthis
lessoniana. The most important cuttlefish are five species of Sepia and Sepiella inermis. The dominant octopus are
Amphioctopus membranaceus, A. aegina and Cistopus indicus (FAO, 2019). There are also small wild-caught fisheries
for the short-necked clam Paratapes undulatus, scallops such as Amusium pleuronectes, and the razor clam Solen
strictus (Chalermwat et al., 2003).
Thai aquaculture is dominated by shrimp production, which constitutes over 90% of the total value, which was
USD $1.75 billion in 2007. Molluscan aquaculture in that year totalled 362,800 tonnes, 76% of which was the Asian
green mussel (Perna viridis) (FAO, 2019). The green mussel has been cultured in Thailand for over 60 years using
three methods: bamboo poles, rope culture and raft culture (Keawtawee et al., 2018). Chalermwat et al. (2003)
13
The Marine and Estuarine Molluscs of Thailand
divided Thai molluscan aquaculture into two methods: species produced through traditional aquaculture
techniques without associated hatcheries and species obtaining stock from hatcheries.
Yang et al. (2019) reported that Thai aquaculture production is among that of the top 10 nations worldwide, with
1.45% of global production. In 2012, Thai molluscan aquaculture totalled 205,192 metric tonnes. The most
important species were: green mussels (Perna viridis), 107,229 tonnes; blood cockles (Anadara granosa), 71,450
tonnes; oysters (Crassostrea spp.), 26,508 tonnes; European flat oysters (Ostrea edulis), 4 tonnes; and abalone
(Haliotis spp.), 5 tonnes. Small quantities of horse mussels (Modiolus spp.) were also grown but not reported (Yang
et al., 2019). According to Chalermwat et al. (2003), the key species of oysters are Saccostrea cucullata, Crassostrea
belcheri and Crassostrea iredalei.
Ngamprasertkit (2018) reported that total marine aquaculture in Thailand in 2017 was 508,700 tonnes. The
dominant molluscs in that year were Asian green mussels (Perna viridis) at 115,500 tonnes, blood cockles (Anadara
granosa) at 59,000 tonnes, and oysters at 19,900 tonnes. Green mussels and blood cockles typically constitute over
80% of the total cultured shellfish harvest by weight, but the relatively small oyster harvest is much more valuable
and provides almost 50% of the total annual farmed shellfish crop value (Chalermwat et al., 2003).
The horse mussel Arcuatula arcuatula is produced in small quantities for animal feed and, at the time of the report,
aquaculture for the abalone Haliotis asinina and the spotted babylon Babylonia areolata were being developed
(Keawtawee et al., 2018).
COMPARISON OF THE MOLLUSCS OF THE GULF OF THAILAND AND THE ANDAMAN SEA
The total number of species recorded from the Gulf of Thailand (2,494) was slightly more than the 2,400 recorded
from the Andaman Sea. 61 species have been recorded from Thailand without either coastline being specified.
About 44% of the species (1,524) have been recorded from both coastlines, demonstrating a considerable overlap
of widespread Indo-Pacific species. The high degree of overlap in mollusc species found on the two coasts is
consistent with the finding of Wells (2002), who recorded a very high overlap between the molluscs of the eastern
Indian Ocean and the western Pacific Ocean. Further collecting in Thailand would be expected to increase the number
of species in common on the two coasts as many species have been recorded in only one or two papers.
The two Thai coastlines are very different, particularly in the absence of deep water in the Gulf of Thailand. Collecting
on the Andaman coast appears to have been more extensive through the long–term work of PMBC. On the other hand,
there are more institutions and diffuse collecting efforts in the Gulf of Thailand. As there is no way of quantifying
collecting effort, it is uncertain whether molluscan biodiversity in the two regions is similar. Because of their economic
importance, cephalopods have been the most thoroughly studied class. 94 species have been recorded from the Gulf
of Thailand and 106 from the Andaman coast (Table 1), suggesting the latter may have a slightly greater biodiversity.
SUGGESTIONS FOR FUTURE RESEARCH
The lack of detailed information on the marine and estuarine molluscs of Thailand prevents an accurate estimation
of the number of species in the country, or on either of its coasts. This lack of basic biodiversity information is the
most pressing research requirement for our understanding of Thai marine and estuarine molluscs. Fortunately,
there has been a great deal of survey work undertaken in recent decades. We have highlighted above studies that
have been undertaken but where the molluscs have not been published, such as the original Thai–Danish expedition
in 1900, the Fifth Expedition in 1966, and the 1996–2000 BIOSHELF expedition. Specimens from these studies
are deposited either in PMBC or the Statens Naturhistoriske Museum, Copenhagen, Denmark. Identification and
publication of the materials in these collections would contribute substantially to our knowledge of Thai marine
molluscs in a cost-effective manner.
Once this material is collated and published, additional field surveys could be undertaken to further document the
group in Thailand. The Kungkrabaen workshop (Bieler et al., 2008a) and similar workshops in Singapore (e.g., Tan
& Goh, 2016) and Australia (e.g., Wells et al., 2003) provide a format for undertaking such surveys, and PMBC
provides a natural location for a workshop. The format is simple. A small local organising committee selects the
venue and organises facilities. Accommodation and meals could be provided either at PMBC or in a nearby venue;
these would need to be inexpensive. Basic laboratory facilities, vehicles, boats, samp ling equipment, research
permits, etc. would be provided by PMBC. International and Thai scientists would be invited to participate in the
workshop. The scientists would concentrate on taxonomy, though basic field ecology and related subjects could also
be researched. Thai graduate students could be selected to work with the team at the field location. The field
workshop would be held for about 18 days and be targeted to coincide with two spring low tide periods. The
organising committee would be responsible for all logistical aspects. Once the venue and timing have been organised,
the international researchers can be invited.
14
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
The international researchers would be expected to arrange their own transport to the site and contribute to the
cost of the workshop through a registration fee that would cover the costs of accommodation, meals and some of
the other workshop expenses. In return, the international researchers would be afforded the opportunity to
conduct research in their area of expertise in an overseas location with local scientists, and have all logistical
aspects organised for them, including the necessary collecting and export permits. Holotypes and a selection of
paratypes of any new species described and representative samples of species collected would be deposited in
PMBC. If sufficient paratypes and other specimens are available, they could be deposited in THNHM and the
taxonomist’s home institution. Two years should be allowed for the international scientists to obtain funding for
their participation.
All participants would be expected to publish one or more papers reporting their results in a joint volume, such as
the PMBC Special Publication series. Alternatively, papers could be published in international journals and cite the
workshop as part of a numbered series. Three years should be allowed for completion of laboratory studies,
writing up of papers, and the reviewing process. Thus, the workshop would be a five-year project. PMBC is an
obvious location for a workshop, but there are a number of other possible venues in Thailand. Grant support can
be sought to lower the cost to participants and enhance the workshop and/or publication.
It should be noted that the workshop format would work equally well for other taxa, either as independent
workshops or as multidisciplinary surveys.
Much of the impetus for the Fifth Thai–Danish Expedition 1966 was a requirement by the Thai Department of
Fisheries to understand the food chains that support commercially valuable fisheries (Seidenfaden et al., 1968).
As molluscs are the most diverse phylum in the marine environment and many species are ecologically and/or
economically important, a better understanding of the biodiversity and distribution of molluscs in Thailand is
required to support fisheries science.
15
The Marine and Estuarine Molluscs of Thailand
LIST OF THAI MARINE AND ESTUARINE MOLLUSCS
Class Bivalvia
Subclass Heterodonta
Infraclass Archiheterodonta
Order Carditida
Superfamily Carditoidea
Family Carditidae
Beguina semiorbiculata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989),
Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Gemert (2003), Duangdee
(2004, 2007), Kasetsart University (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Hylleberg & Aungtonya (2013), BEDO (2017a).
Cardita crassicosta Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO
(2017a).
Cardita leana Dunker, 1860. Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Aungtonya et al.
(1999), Nabhitabhata (2009), BEDO (2017a).
Cardita variegata Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1938),
Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Bussarawit (1995), Sanpanich (1998,
2011), Aungtonya et al. (1999), Robba et al. (2002), Gemert (2003), Duangdee (2004, 2007), Wells (2005),
Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a), ONEP (2007),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Hylleberg & Aungtonya (2013), Printrakoon et al.
(2016), BEDO (2017a), GBIF (2019a, 2019b).
Cardites bicolor (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998).
Cardites floridanus (Conrad, 1838). Reported as: Carditamera floridana (Conrad, 1838). Andaman Sea. Reported
by: Nabhitabhata (2009), BEDO (2017a).
Family Condylocardiidae
Carditellona pulchella (Lynge, 1909). Reported as: Carditella (Carditellona) pulchella Lynge, 1909. Gulf of
Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009).
Crassacuna pusilla (Lynge, 1909). Reported as: Choniocardia (Carditellopsis) pusilla (Lynge, 1909). Gulf of
Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009), Huber (2010).
Superfamily Crassatelloidea
Family Crassatellidae
Bathytormus radiatus (G. B. Sowerby I, 1825). Also reported as: Crassatella radiata (Sowerby, 1825); and
Crassatellites radiata (Sowerby). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Kurozumi et al. (1989), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Bathytormus ziczac (Reeve, 1842). Reported as: Crassatella ziczac Reeve, 1842; and Talabrica ziczac (Reeve,
1842). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Chattina picta (Adams & Reeve, 1850). Reported as: Crassatina picta (Adams & Reeve, 1850); and Talabrica cf.
picta (A. Adams & Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009).
Infraclass Euheterodonta
Superorder Anomalodesmata
Superfamily Clavagelloidea
Family Clavagellidae
16
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Bryopa lata (Broderip, 1834). Reported as: Clavagella (Dacosta) lata Broderip in Owen, 1835. Gulf of Thailand,
Andaman Sea. Reported by: Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Family Penicillidae
Verpa penis (Linnaeus, 1758). Reported as: Brechites dichotomous Chenu, 1843; and Brechites (Verpa) aquarius
(E. J. Burrow, 1815). Gulf of Thailand, Andaman Sea. Reported by: Smith (1976), Lynge (1909), Nabhitabhata
(2009), Huber (2010), GBIF (2019a).
Verpa philippinensis (Chenu, 1843). Reported as: Brechites philippinensis (Chenu, 1843). Gulf of Thailand,
Andaman Sea. Reported by: Sanpanich & Duangdee (2009), Sanpanich (2011).
Superfamily Cuspidarioidea
Family Cuspidariidae
Cardiomya singaporensis (Hinds, 1843). Reported as: Neaera singaporensis Hinds, 1843; and Cardiomya
(Cardiomya) singaporensis (Hind, 1843). Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993),
Scott (1995), Nabhitabhata (2009).
Cuspidaria gigantea Prashad, 1932. Reported as: Cuspidaria kawamurai Kuroda, 1948. Andaman Sea. Reported
by: Nabhitabhata (2009), BEDO (2017a).
Cuspidaria rosea (Hinds, 1843). Reported as: Neaera rosea Hinds, 1843. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009).
Cuspidaria steindachneri Sturany, 1899. Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Plectodon cf. ligula (Yokoyama, 1922). Gulf of Thailand. Reported by: Scott (1995).
Pseudoneaera semipellucida (Kuroda, 1948). Reported as: Austroneaera cf. semipellucida (Kuroda, 1948). Gulf
of Thailand. Reported by: Scott (1995).
Rengea caduca (E. A. Smith, 1894). Reported as: Cuspidaria caduca (E. A. Smith, 1894). Andaman Sea. Reported
by: Nabhitabhata (2009), BEDO (2017a).
Superfamily Pandoroidea
Family Lyonsiidae
Entodesma navicula (Adams & Reeve, 1850). Reported as: Agriodesma navicula (Adams & Reeve, 1850). Gulf of
Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a).
Family Pandoridae
Pandora aversa (Hedley, 1913). Reported as: Coelodon elongatus Carpenter, 1864; Pandorella elongata
(Carpenter, 1864); and Pandora (Frenamya) elongata (Carpenter, 1864). Gulf of Thailand. Reported by: Lynge
(1909), Bernard et al. (1993), Robba et al. (2002), Nabhitabhata (2009).
Superfamily Thracioidea
Family Laternulidae
Exolaternula liautaudi (Mittre, 1844). Reported as: Laternula cf. corrugata (Reeve, 1863). Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Exolaternula spengleri (Gmelin, 1791). Reported as: Laternula rostrata (G. B. Sowerby II, 1839). Gulf of Thailand.
Reported by: GBIF (2019b).
Laternula anatina (Linnaeus, 1758). Also reported as: Laternula (Laternula) anatina (Linnaeus, 1758). Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Kasetsart University (2006),
Nabhitabhata (2009), Taylor (2009), Huber (2010), Sanpanich (2011).
Laternula attenuata (Reeve, 1863). Gulf of Thailand. Reported by: Nabhitabhata (2009).
Family Periplomatidae
Cochlodesma praetenue (Pulteney, 1799). Reported as: Laternula truncata (Lamarck, 1818). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999),
Prezant et al. (2008), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Periploma indicum Melvill, 1898. Reported as: Periploma (Periploma) indicum Melvill, 1898. Gulf of Thailand.
Reported by: Robba et al. (2002), Nabhitabhata (2009).
17
The Marine and Estuarine Molluscs of Thailand
Family Thraciidae
Thracia concinna Reeve, 1859. Reported as: Eximiothracia cf. concinna (Gould, 1861). Gulf of Thailand. Reported
by: Scott (1995).
Thracia modesta Angas, 1868. Reported as: Thracia jacksonensis Smith, 1786. Gulf of Thailand. Reported by:
Sanpanich (2011).
Superfamily Verticordioidea
Family Euciroidae
Euciroa eburnea (Wood-Mason & Alcock, 1891). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Family Verticordiidae
Spinosipella deshayesiana (P. Fischer, 1862). Also reported as: Verticordia deshayesiana (Fischer, 1862). Andaman
Sea. Reported by: Bernard et al. (1993), Simone & Cunha (2008), Nabhitabhata (2009), BEDO (2017a).
Superorder Imparidentia
Order Adapedonta
Superfamily Hiatelloidea
Family Hiatellidae
Hiatella arctica (Linnaeus, 1767). Reported as: Hiatella orientalis (Yokoyama, 1920). Gulf of Thailand, Andaman
Sea. Reported by: Gemert (2003).
Superfamily Solenoidea
Family Pharidae
Cultellus attenuatus Dunker, 1861. Also reported as: Phaxas attenuatus (Dunker, 1862); and Phaxus attenuatus
(Dunker, 1862). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Amornjaruchit
(1988), Nateewathana (1995), Scott (1995), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a).
Cultellus maximus (Gmelin, 1791). Also reported as: Cultellus lacteus (Spengler, 1794). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Robba et al. (2002), Nabhitabhata (2009),
Negri et al. (2014), BEDO (2017a), GBIF (2019a).
Cultellus ovalis Dunker, 1861. Reported as: Cultellus (Pharella) ovalis Dunker, 1861. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009). Reported by WoRMS Editorial Board (2019) as a taxon inquirendum.
Cultellus philippianus Dunker, 1877. Reported as: Ensiculus philipanus (Dunker). Gulf of Thailand. Reported by:
Duangdee et al. (2007).
Cultellus subellipticus Dunker, 1862. Reported as: Solecurtus scarprum Gould and Cultellus scalprum (Gould,
1851). Gulf of Thailand. Reported by: Lynge (1909), Suvatti (1937, 1938), Habe (1965), Swennen et al. (2001),
Nabhitabhata (2009), GBIF (2019b).
Elizia tricarinata. Unable to confirm. Gulf of Thailand. Reported by: Suvatti (1938), Nabhitabhata (2009).
Ensiculus cultellus (Linnaeus, 1758). Reported as: Cultellus (Ensiculus) cultellus Linnaeus, 1758; Phaxas cultellus
(Linnaeus, 1758) and Phaxas (Ensiculus) cultellus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Habe (1965), Nateewathana et al. (1981), Kurozumi et al. (1989), Scott (1995), Aungtonya
et al. (1999), Swennen et al. (2001), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Ensiculus marmoratus (Dunker, 1862). Also reported as: Phaxas marmoratus (Dunker, 1861). Gulf of Thailand,
Andaman Sea. Reported by: Habe (1965), Kilburn & Hylleberg (1998), Nabhitabhata (2009).
Neosiliqua winteriana (Dunker, 1852). Also reported as: Siliqua winteriana (Dunker, 1853). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009), BEDO (2017a).
Orbicularia orbiculata (W. Wood, 1828). Reported as: Elizia orbicularis Wood. Gulf of Thailand. Reported by:
Suvatti (1938), Nabhitabhata (2009).
Pharella acutidens (Broderip & G. B. Sowerby I, 1829). Gulf of Thailand. Reported by: Swennen et al. (2001).
Pharella javanica (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Taylor et al. (2007), Nabhitabhata (2009), GBIF (2019a).
18
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Siliqua fasciata (Spengler, 1794). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Siliqua grayana (Dunker, 1862). Reported as: Siliqua (Poortenia) grayana (Dunker, 1862). No Thai locality
provided. Reported by: Huber (2010).
Siliqua japonica (Dunker, 1861). Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999),
Nabhitabhata (2009), BEDO (2017a).
Siliqua minima (Gmelin, 1791). Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993), Swennen
et al. (2001), Robba et al. (2002), Nabhitabhata (2009), Negri et al. (2014).
Siliqua radiata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909),
Suvatti (1937, 1938), Nateewathana et al. (1981), Aungtonya et al. (1999), Kasetsart University (2000, 2006),
Nabhitabhata (2009), BEDO (2017a), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Sinonovacula constricta (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Chaitiamwong et al.
(1971), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Sinonovacula rugosa (Linnaeus, 1767). Reported as: Sinovacula rugosa (Linnaeus, 1767). Gulf of Thailand,
Andaman Sea. Reported by: Aungtonya et al. (1999), Nabhitabhata (2009). This species cannot be confirmed
through WoRMS Editorial Board (2019), but is discussed by Morton (2009).
Family Solenidae
Solen ceylonensis Leach, 1814. Reported as: Solen truncata Wood, 1815. Gulf of Thailand, Andaman Sea. Reported
by: Scott (1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Solen corneus Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Kurozumi et al. (1989),
Carpenter & Niem (1998), Tuaycharoen (1999), Tuaycharoen & Matsukuma (2001), Robba et al. (2002), Cosel
(2002), Nabhitabhata (2009), Saeedi et al. (2013), Negri et al. (2014), Coan et al. (2015), GBIF (2019a).
Solen curtus Des Moulins, 1832. Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009). This
species cannot be confirmed through WoRMS Editorial Board (2019), but is reported by Swennen et al. (2001).
Solen delesserti Sowerby, 1874. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Solen exiguus Dunker, 1862. Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Tuaycharoen
(1999), Tuaycharoen & Matsukuma (2001), Nabhitabhata (2009), Simone (2009).
Solen grandis Dunker, 1862. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Chaitiamwong et al.
(1971), Nateewathana et al. (1981), Bernard et al. (1993), Carpenter & Niem (1998), Aungtonya et al. (1999),
Tuaycharoen & Matsukuma (2001), Kasetsart University (2006), Duangdee et al. (2009), Nabhitabhata (2009),
Sanpanich (2011), BEDO (2017a).
Solen linearis Spengler, 1794. Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993), Nabhitabhata
(2009).
Solen malaccensis Dunker, 1862. Also reported as: Ensis malaccensis Dunker, 1861. Gulf of Thailand. Reported
by: Suvatti (1938), Tuaycharoen & Matsukuma (2001), Nabhitabhata (2009).
Solen regularis Dunker, 1862. Gulf of Thailand. Reported by: Tuaycharoen & Matsukuma (2001), Nabhitabhata
(2009).
Solen roseomaculatus Pilsbry, 1901. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Carpenter & Niem (1998), Aungtonya et al. (1999), ONEP (2007), Nabhitabhata (2009).
Solen sarawakensis Cosel, 2002. Also reported as: Solen timorensis Dunker, 1852. Gulf of Thailand. Reported by:
Swennen et al. (2001), Cosel (2002), Nabhitabhata (2009), Saeedi et al. (2013).
Solen strictus Gould, 1861. Gulf of Thailand, Andaman Sea. Reported by: Amornjaruchit (1988), Nateewathana
(1995), Aungtonya et al. (1999), Tuaycharoen (1999), Tuaycharoen & Matsukuma (2001), Nabhitabhata (2009),
BEDO (2017a), GBIF (2019b).
Solen thailandicus Cosel, 2002. Reported as: Solen (Solen) thailandicus R. von Cosel, 2002. No Thai locality
provided. Reported by: Cosel (2002), Huber (2010), Saeedi et al. (2013).
Solen thachi Cosel, 2002. Reported as: Solen (Solen) thachi R. von Cosel, 2002. No Thai locality provided. Reported
by: Cosel (2002), Huber (2010).
Solen timorensis Dunker, 1852. Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Solen vagina Linnaeus, 1758. Also reported as: Solen brevis Gray, 1832; Solen abbreviatus Philippi; and Solen
lamarckii Deshayes, 1839. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Carpenter & Niem
(1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University (2006), Nabhitabhata (2009),
BEDO (2017a), GBIF (2019b).
Solen vaginoides Lamarck, 1818. Also reported as: Solen correctus Iredale, 1937. Gulf of Thailand. Reported by:
Lynge (1909), Swennen et al. (2001), ONEP (2007), Nabhitabhata (2009).
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The Marine and Estuarine Molluscs of Thailand
Order Cardiida
Superfamily Cardioidea
Family Cardiidae
Acrosterigma attenuatum (G. B. Sowerby II, 1841). Reported as: Laevicardium attenuatum (Sowerby, 1841).
Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Aungtonya et al. (1999), Vidal (1999b),
Hylleberg (2004), Nabhitabhata (2009), Poorten (2009), BEDO (2017a).
Acrosterigma biradiatum (Bruguière, 1789). Andaman Sea. Reported by: BEDO (2017a).
Acrosterigma cygnorum (Deshayes, 1855). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich (2011), Duangdee et al. (2013a), Sanpanich & Duangdee (2018).
Acrosterigma impolitum (G. B. Sowerby II, 1834). Also reported as: Acrosterigma rosemariensis Wilson &
Stevenson, 1977; Trachycardium impolitum Sowerby, 1840; Trachycardium impolitum (Sowerby, 1841); and
Vasticardium impolitum (Sowerby, 1833). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard
et al. (1993), Vidal (1999b), Swennen et al. (2001), Robba et al. (2002), Hylleberg (2004, 2009), Nabhitabhata
(2009), Sanpanich (2011), GBIF (2019b).
Acrosterigma maculosum (W. Wood, 1815). Also reported as: Trachycardium maculosum (Wood, 1815);
Acrosterigma arenicolum Reeve, 1845; and Trachycardium arenicola Reeve, 1843. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Nateewathana et al. (1981), Bernard et al. (1993), Bussarawit (1995), Aungtonya
et al. (1999), Vidal (1999b), Hylleberg (2004, 2009), Nabhitabhata (2009), Poorten (2009), Sanpanich (2011),
BEDO (2017a).
Acrosterigma oxygonum (G. B. Sowerby II, 1834). Gulf of Thailand, Andaman Sea. Reported by: Vidal (1999a,
1999b), Hylleberg (2004), Nabhitabhata (2009).
Acrosterigma simplex (Spengler, 1799). Also reported as: Acrosterigma unicolor (G. B. Sowerby II, 1834) and
Acrosterigma nebulosum Reeve, 1845. Gulf of Thailand, Andaman Sea. Reported by: Vidal (1999b), Wells (2005),
Duangdee et al. (2009), Hylleberg (2009), Nabhitabhata (2009), Sanpanich (2011), Sanpanich & Duangdee (2018).
Acrosterigma transcendens (Melvill & Standen, 1899). Also reported as: Vasticardium transcendens (Melvill &
Standen, 1899). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2018), Sanpanich (2011).
Afrocardium exochum (Melvill in Melvill & Standen, 1907). Andaman Sea. Reported by: Nabhitabhata (2009),
BEDO (2017a).
Afrocardium richardi (Audouin, 1826). Also reported as: Acrosterigma richardi; Fragum carditaeforme Reeve,
1845; and Afrocardium carditaeforme Reeve, 1845. Gulf of Thailand. Reported by: Lynge (1909), Robba et al.
(2002), Nabhitabhata (2009).
Corculum cardissa (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Morris & Purchon (1981), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998),
Aungtonya et al. (1999), Hylleberg (2004, 2009), Nabhitabhata (2009), Poorten (2009), BEDO (2017a).
Ctenocardia fornicata (G. B. Sowerby II, 1840). Also reported as: Ctenocardia adamsi (A. Adams & Reeve, 1850).
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Gemert
(2003), Hylleberg (2004, 2009), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Ctenocardia translata (Prashad, 1932). Gulf of Thailand. Reported by: Poorten (2007).
Ctenocardia virgo (Reeve, 1846). Also reported as: Ctendcardia perornata; Ctenocardium imbricatum; and
Fragum peronatum Iredale, 1929. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al.
(1993), Hidalgo et al. (1999), Hylleberg (2004, 2009), Poorten (2007, 2009), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Discors multipunctatum (G. B. Sowerby I in Broderip & G. B. Sowerby I, 1833). Reported as: Laevicardium
multipunctatum (G. B. Sowerby I in Broderip & G. B. Sowerby I, 1833). Gulf of Thailand, Andaman Sea. Reported
by: Gemert (2003), Hylleberg (2009), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Fragum carinatum (Lynge, 1909). Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979),
Nateewathana et al. (1981), Aungtonya et al. (1999), Hylleberg (2004, 2009), Nabhitabhata (2009).
Fragum erugatum (Tate, 1889). Also reported as: Afrocardium erugatum (Tate, 1889). Gulf of Thailand, Andaman
Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Fragum fragum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Duangdee (2004), Hylleberg (2004, 2009),
Wells (2005), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009, 2013a),
Nabhitabhata (2009), Sanpanich (2011).
Fragum nivale (Reeve, 1845). Thai locality not stated. Reported by: Hylleberg (2009).
Fragum scruposum (Deshayes, 1855). Also reported as: Fragum bannoi Otuka, 1937. Gulf of Thailand, Andaman
Sea. Reported by: Gemert (2003), Hylleberg (2009), Huber (2010), BEDO (2017a).
20
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Fragum sueziense (Issel, 1869). Reported as: Cardium (Acanthocardium) sueziense Issel, 1869; and Parvicardium
suezensis (Issel, 1869). Gulf of Thailand. Reported by: Lynge (1909), Aungtonya et al. (1999), Gemert (2003),
Hylleberg (2004, 2009), Nabhitabhata (2009).
Fragum unedo (Linnaeus, 1758). Also reported as: Cardium unedo Linnaeus; and Hemicardium unedo Linnaeus,
1758. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Nielsen
(1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998),
Aungtonya et al. (1999), Wells (2005), Poorten (2007), Hylleberg (2009), Nabhitabhata (2009), Sanpanich
(2011), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019a).
Frigidocardium centumliratum (Melvill, 1907). Reported as: Frigidocardium centumliratum (Melvill in Melvill
& Standen, 1907). Andaman Sea. Reported by: Poorten & Gemert (2016).
Frigidocardium exasperatum (G. B. Sowerby II, 1839). Reported as: Microcardium exasperatum (Sowerby, 1834);
and Nemocardium exasperatum (Sowerby, 1841). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana
et al. (1981), Aungtonya & Hylleberg (1998), Aungtonya et al. (1999), Hylleberg (2004, 2009), Nabhitabhata
(2009), BEDO (2017a).
Frigidocardium torresi (E. A. Smith, 1885). Reported as: Nemocardium torresi (Smith, 1885). Andaman Sea.
Reported by: Nabhitabhata (2009), Poorten & Gemert (2016), BEDO (2017a).
Fulvia aperta (Bruguière, 1789). Reported as: Cardium (Papyridea) rugatum Gronovics, 1781; Cardium (Fulvia)
apertum; and Fulvia (Fulvia) aperta (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: Lynge
(1909), Suvatti (1937, 1938), Kurozumi et al. (1989), Hylleberg (1994, 2004, 2009), Vidal (1994), Sanpanich
(1998, 2011), Swennen et al. (2001), Kasetsart University (2006), ONR & EPP (2006), Duangdee (2007),
Duangdee et al. (2007, 2011), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018).
Fulvia australis (G. B. Sowerby II, 1834). Also reported as: Laevicardium australis (Sowerby, 1841); and Fulvia
striata (Splengler, 1799). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al.
(1981), Bussarawit (1995), Aungtonya & Hylleberg (1998), Aungtonya et al. (1999), Swennen et al. (2001),
Hylleberg (2004, 2009), Kasetsart University (2006), ONR & EPP (2006), Duangdee (2007), ONEP (2007),
Nabhitabhata (2009), Duangdee et al. (2011), GBIF (2019a).
Fulvia hungerfordi (G. B. Sowerby III, 1901). Also reported as: Fulvia (Laevifulvia) hungerfordi (G. B. Sowerby
III, 1901); and Cardium (Papyridea) hungerfordi Sowerby, 1901. Gulf of Thailand. Reported by: Lynge (1909),
Bernard et al. (1993), Higo et al. (1999), Robba et al. (2002), Hylleberg (2004, 2009), Nabhitabhata (2009),
Poorten (2009).
Fulvia laevigata (Linnaeus, 1758). Reported as: Fulvia papyracea (Bruguière, 1789); and Cardium (Papyridea)
papyraceum Chemnitz, 1782. Gulf of Thailand. Reported by: Lynge (1909), Carpenter & Niem (1998), Hylleberg
(1994, 2004), Duangdee et al. (2007), Nabhitabhata (2009).
Fulvia mutica (Reeve, 1844). Reported as: Cardium (Papyridea) annae Pilsbry, 1904. Gulf of Thailand. Reported
by: Lynge (1909), Hylleberg (2004), Nabhitabhata (2009).
Fulvia scalata Vidal, 1994. Gulf of Thailand, Andaman Sea. Reported by: Hylleberg (2009), Nabhitabhata (2009).
Hippopus hippopus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Poorten (2007), Nabhitabhata (2009).
Laevicardium lobulatum Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Hylleberg (2004, 2009),
Nabhitabhata (2009), Poorten (2009).
Laevicardium undatopictum (Pilsbry, 1904). Gulf of Thailand. Reported by: Hylleberg (2009), Sanpanich (2011).
Lunulicardia hemicardium (Linnaeus, 1758). Also reported as: Hemicardium hemicardium (Linnaeus, 1758);
Fragum hemicardium (Linnaeus, 1758); Fragum hemicardium carinata Lynge, 1909; Corculum hemicardium
(Linnaeus, 1758); and Cardium hemicardium Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by:
Suvatti (1937, 1938), Bernard et al. (1993), Carpenter & Niem (1998), Sanpanich (1998, 2011), Kasetsart
University (2000), Hylleberg (2004), Nabhitabhata (2009), Sanpanich & Duangdee (2018), J. D. Taylor et al.
(unpublished data). Poorten (2007) questions the Thai records.
Lunulicardia retusa (Linnaeus, 1767). Reported as: Lunulicardia retusa (Linnaeus, 1758); Lunulicardia subretusum
(Linnaeus, 1767); and Fragum subretusa (Sowerby). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Aungtonya et al. (1999), Swennen et al. (2001), Gemert (2003), Hylleberg (2004,
2009), Duangdee et al. (2007), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Lunulicardia retusa auricula (Niebuhr, 1775). Reported as: Lunulicardia auricula (Niebuhr in Forsskål, 1775).
Gulf of Thailand. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Lyrocardium lyratum (G. B. Sowerby II, 1840). Andaman Sea. Reported by: BEDO (2017a).
Maoricardium pseudolatum Voskuil & Onverwagt, 1991. Reported as: Plagiocardium pseudolatum Voskuil &
Onverwagt, 1991. Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Duangdee et al. (2009).
Maoricardium pseudolima (Lamarck, 1819). Andaman Sea. Reported by: GBIF (2019a).
Maoricardium setosum (Redfield, 1846). Reported as: Plagiocardium setosum (Redfield, 1848); Plagiocardium
(Maoricardium) setosum (Redfield, 1846); Vepricardium setosum (Redfield, 1846); and Cardium (Acanthocardium)
setosum Redfield, 1846. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bussarawit (1995),
Aungtonya et al. (1999), Swennen et al. (2001), Hylleberg (2004), Nabhitabhata (2009), BEDO (2017a).
Microcardium cf. aequiliratum Poutiers, 1981. Andaman Sea. Reported by: Nabhitabhata (2009).
21
The Marine and Estuarine Molluscs of Thailand
Nemocardium fulvum ter Poorten, 2013. Andaman Sea. Tentatively reported by: Poorten (2013).
Papyridea lata (Born, 1778). Reported as: Cardium latum Born, 1780. Gulf of Thailand, Andaman Sea. Reported
by: Morlet (1889), Suvatti (1938), Duangdee et al. (2009), Nabhitabhata (2009).
Papyridea soleniformis (Bruguière, 1789). Reported as: Laevicardium bullata (Linnaeus, 1758). Andaman Sea.
Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009).
Tridacna crocea Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Bussarawit (1995),
Chantrapornsyl et al. (1996), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999),
Upanoi & Banchungmanee (1999), Banchongmanee & Upanoi (2000), Duangdee (2004), Wells (2005),
Duangdee et al. (2007, 2009, 2013a, 2013b), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2018), BEDO (2017a), GBIF (2019a, 2019b).
Tridacna gigas (Linnaeus, 1758). Gulf of Thailand. Reported by: Tantanasiriwong (1979), Nateewathana et al.
(1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Hylleberg (2004), Nabhitabhata (2009), BEDO
(2017a).
Tridacna maxima (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1979),
Nateewathana et al. (1981), Bussarawit (1995), Chantrapornsyl et al. (1996), Carpenter & Niem (1998),
Aungtonya et al. (1999), Upanoi & Banchungmanee (1999), Banchongmanee & Upanoi (2000), Hylleberg
(2004), Kasetsart et al. (2004), Wells (2005), Duangdee et al. (2009), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Tridacna squamosa Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909),
Suvatti (1937, 1938), Tantanasiriwong (1979), Nateewathana et al. (1981), Amornjaruchit (1988), Bernard
et al. (1993), Bussarawit (1995), Chantrapornsyl et al. (1996), Carpenter & Niem (1998), Sanpanich (1998,
2011), Aungtonya et al. (1999), Upanoi & Banchungmanee (1999), Banchongmanee & Upanoi (2000),
Kittiwattanawong et al. (2001), Duangdee (2004, 2007), Hylleberg (2004), Kasetsart et al. (2004), Wells
(2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2018), BEDO (2017a), GBIF (2019a, 2019b).
Vasticardium angulatum (Lamarck, 1819). Also reported as: Acrosterigma angulata (Lamarck, 1819);
Laevicardium attenuatum (Sowerby); Acrosterigma attenuatum (Sowerby, 1841); Trachycardium angulatum
(Lamarck, 1819); and Vasticardium alternatum (Sowerby). Gulf of Thailand, Andaman Sea. Reported by:
Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya
et al. (1999), Duangdee (2004, 2007), Hylleberg (2004), Kasetsart University (2006), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011, 2013a), ONEP (2007), Nabhitabhata (2009), Poorten (2009), Sanpanich &
Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a).
Vasticardium compunctum Kira, 1959. Gulf of Thailand. Reported by: Kasetsart University (2006), Duangdee
(2007), Nabhitabhata (2009).
Vasticardium dupuchense (Reeve, 1845). Reported as: Acrosterigma dupuchense (Reeve, 1845); and Cardium
dupuchense Reeve, 1845. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Suvatti (1938),
Nabhitabhata (2009), Sanpanich (2011).
Vasticardium elongatum (Bruguière, 1789). Also reported as: Casticardium elongatum (J. C. Bruguière, 1789)
forma enode and forma okinawaense Huber (2010); Acrosterigma elongata (Bruguière, 1789); and Trachycardium
elongatum (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (1998, 2011), Swennen
et al. (2001), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Huber (2010), GBIF (2019a).
Vasticardium fidele (Vidal, 1992). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Hylleberg
(2004, 2009, 2015), Nabhitabhata (2009), BEDO (2017a).
Vasticardium flavum (Linnaeus, 1758). Reported as: Acrosterigma flava (Linnaeus, 1758); Acrosterigma flava
flava (Linnaeus, 1758); Acrosterigma gratiosa (Deshayes, 1855); Vasticardium flavum flavum (Linnaeus, 1758);
and Trachycardium flavum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Suvatti (1937, 1938), Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al.
(1981), Kurozumi et al. (1989), Bussarawit (1995), Sanpanich (1998, 2011), Gemert (2003), Duangdee (2004,
2007), Kasetsart University (2006), ONR & EPP (2006), Nabhitabhata (2009), Poorten (1997, 2007, 2009),
Duangdee et al. (2011, 2013b), Hylleberg & Aungtonya (2013), BEDO (2017a), Sanpanich & Duangdee (2018),
GBIF (2019a), J. D. Taylor et al. (unpublished data).
Vasticardium indioceanum (Vidal, 1993). Also reported as: Vasticardium elongatum indioceanum (Vidal, 1993).
Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Hylleberg (2004), GBIF (2019a, 2019b).
Vasticardium kenyanum (Cox, 1930). No Thai locality provided. Reported by: Poorten (2009).
Vasticardium luteomarginatum (Voskuil & Onverwagt, 1991). Reported as: Acrosterigma luteomarginata
(Voskuil & Onverwagt, 1991); Trachycardium luteomarginatum Voskuil & Onverwagt, 1991; and Vasticardium
luteomarginatum luteomarginatum (Voskuil & Onverwagt, 1991). Gulf of Thailand, Andaman Sea. Reported by:
Voskuil & Onverwagt (1991), Vidal (1997a), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018),
Sanpanich (2011), Duangdee et al. (2013a), Galli (2015), BEDO (2017a).
Vasticardium mendanaense (G. B. Sowerby III, 1897). Reported as: Acrosterigma mendanaense (G. B. Sowerby
III, 1897). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
22
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Vasticardium pectiniforme (Born, 1780). Also reported as: Cardium pectiniforme Born, 1780; Vasticardium
rugosum (Lamarck, 1819); and Trachycardium rugosum (Lamarck, 1819). Gulf of Thailand, Andaman Sea.
Reported by: Suvatti (1937, 1938), Vidal (1997b), Carpenter & Niem (1998), Gemert (2003), Duangdee (2004,
2007), Kasetsart University (2006), Duangdee et al. (2009), Hylleberg (2009), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2018), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Vasticardium rubicundum (Reeve, 1844). Reported as: Trachycardium rubicandum (Reeve, 1845). Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999),
Duangdee (2004), Hylleberg (2004), BEDO (2017a).
Vasticardium sewelli (Prashad, 1932). Reported as: Cardium laddi Abrard, 1946. No Thai locality given.
Reported by: Vidal (1999a), Poorten (2009).
Vasticardium subrugosum (Sowerby, 1840). Also reported as: Trachycardium subrugosum (Sowerby, 1840);
and Vasticardium flavum subrugosum (Sowerby, 1838). Gulf of Thailand, Andaman Sea. Reported by: Vidal
(1997b), Carpenter & Niem (1998), Aungtonya et al. (1999), Gemert (2003), Hylleberg (2004), BEDO (2017a).
Vasticardium wilsoni (Voskuil & Onverwagt, 1991). Reported as: Acrosterigma wilsoni (Voskuil & Onverwagt,
1991). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Vepricardium asiaticum (Bruguière, 1789). Also reported as: Trachycardium asiaticum (Bruguière, 1792); and
Cardium (Bucardium) asiaticum Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Bernard et al. (1993), Bussarawit (1995), Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al. (1999),
Vidal (2000), Swennen et al. (2001), Hylleberg (2004, 2009), ONEP (2007), Duangdee et al. (2009, 2011),
Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Vepricardium coronatum (Schröter, 1786). Reported as: Cardium coronatum Schröter, 1786; Cardium
(Bucardium) coronatum Spengler, 1786; Cardium fimbriatum Wood, 1815; and Cardium (Ringicardium)
fimbriatum (Wood). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937,
1938), Hylleberg (1994, 2004, 2009), Vidal (2000), Swennen et al. (2001), Robba et al. (2002), Gemert (2003),
Kasetsart University (2006), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Vepricardium incarnatum (Reeve, 1844). Also reported as: Vepricardium mirabile (Deshayes, 1854). Gulf of
Thailand, Andaman Sea. Reported by: Bernard et al. (1993), Hylleberg (2009), Sanpanich (2011).
Vepricardium multispinosum (G. B. Sowerby II, 1839). Also reported as: Cardium (Bucardium) multispinosum
Sowerby, 1840; and Trachycardium multispinosum (Sowerby, 1841). Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Morris & Purchon (1981), Nateewathana et al. (1981), Bernard et al. (1993), Bussarawit
(1995), Aungtonya et al. (1999), Higo et al. (1999), Swennen et al. (2001), Hylleberg (2004, 2009), Wells
(2005), Nabhitabhata (2009), Sanpanich (2011).
Vepricardium rubrohamatum Voskuil & Onverwagt, 1988. Reported as: Vepricardium albohamatum Hylleberg
& Vidal, 2000. Andaman Sea. Reported by: Vidal (2000), Hylleberg (2004), Nabhitabhata (2009), Huber (2010),
BEDO (2017a).
Vepricardium sinense (G. B. Sowerby II, 1840). Also reported as: Cardium (Ringicardium) sinense (Sowerby);
and Vepricardium sinense (Sowerby, 1840). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti
(1937, 1938), Bernard et al. (1993), Carpenter & Niem (1998), Aungtonya et al. (1999), Vidal (2000), Swennen
et al. (2001), Robba et al. (2002), Hylleberg (2004, 2009), Kasetsart University (2006), Duangdee (2007), ONEP
(2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a).
Superfamily Tellinoidea
Family Donacidae
Donax brazieri E. A. Smith, 1892. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1938), Nabhitabhata
(2009), Sanpanich (2011). Not recorded by Signorelli & Printrakoon (2020).
Donax cuneatus Linnaeus, 1758. Also reported as: Donax (Latona) cuneatus Linnaeus, 1758; and Donax bicolor
Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Nateewathana et al. (1981),
Bernard et al. (1993), Carpenter & Niem (1998), Aungtonya et al. (1999), Kasetsart University (2000), Swennen
et al. (2001), Gemert (2003), Wells (2005), ONEP (2007), Duangdee et al. (2009), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b), Signorelli & Printrakoon (2020).
Donax faba Gmelin, 1791. Also reported as: Donax (Latona) faba Gmelin, 1791; and Latona faba Gmelin. Gulf of
Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937, 1938), Nielsen (1976b),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Amornjaruchit (1988), Kurozumi
et al. (1989), Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011),
Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee (2004), Kasetsart
University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), BEDO (2017a), GBIF (2019a, 2019b), Signorelli & Printrakoon (2020), J. D. Taylor et al.
(unpublished data).
Donax incarnatus Gmelin, 1791. Reported as: Donax (Latona) incarnatus Gmelin, 1791; Donax (Dentilatona)
incarnatus Gmelin, 1791; and Donax dysoni Deshayes, 1854. Gulf of Thailand, Andaman Sea. Reported by: Lynge
23
The Marine and Estuarine Molluscs of Thailand
(1909), Suvatti (1938), Carpenter & Niem (1998), Subba Rao & Dey (2000), Swennen et al. (2001), Robba et al.
(2002), Nabhitabhata (2009), Negri et al. (2014), BEDO (2017a), GBIF (2019b), Signorelli & Printrakoon (2020),
J. D. Taylor et al. (unpublished data).
Donax introradiatus Reeve, 1854. Reported as: Donax (Paraserrula) introradiatus Reeve, 1854. Gulf of Thailand.
Reported by: Signorelli & Printrakoon (2020).
Donax scortum Linnaeus, 1758. Reported as: Hecuba scortum Gmelin, 1791; and Donax (Hecuba) scortum Linnaeus,
1758. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Amornjaruchit (1988), Aungtonya
et al. (1999), Kasetsart University (2000, 2006), BEDO (2017a), GBIF (2019a), Signorelli & Printrakoon (2020),
J. D. Taylor et al. (unpublished data).
Donax semigranosus (Dunker, 1877). Also reported as: Donax (Deltachion) semigranosus Dunker, 1877. Gulf of
Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Sanpanich (1998, 2011), Robba et al. (2002),
Nabhitabhata (2009), Huber (2010), GBIF (2019b), Signorelli & Printrakoon (2020).
Donax solidus Spengler, 1798. Reported as: Donax (Latona) solidus Spengler, 1798. Andaman Sea. Reported by:
Signorelli & Printrakoon (2020).
Donax spinosus Gmelin, 1791. Also reported as: Donax (Deltachion) spinosus Gmelin, 1791. Andaman Sea.
Reported by: Huber (2010), Signorelli & Printrakoon (2020), J. D. Taylor et al. (unpublished data).
Donax ticaonicus Hanley, 1845. Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009). Not
recorded by Signorelli & Printrakoon (2020).
Donax veruinus Hedley, 1913. Reported as: Donax nitidus Deshayes, 1855. Gulf of Thailand. Reported by: Lynge
(1909), Bernard et al. (1993), Robba et al. (2002), Nabhitabhata (2009), Negri et al. (2014). According to
Signorelli & Printrakoon (2020), Donax veruinus is the valid name for Donax nitidus. It has been recorded as
above in Thai waters, but Signorelli & Printrakoon (2020) did not find any specimens.
Donax victoris Fischer-Piette (1942). Reported as: Donax (Latona) victoris Fischer-Piette (1942). No Thai
locality provided. Reported by: Huber (2010). Not recorded by Signorelli & Printrakoon (2020).
Family Psammobiidae
Asaphis violascens (Forsskål in Niebuhr, 1775). Also reported as: Capsa rugosa Lamarck, 1801; Asaphis
dichotoma (Anton, 1838); Asaphis tahitensis Born; and Asaphis tahitensis Bernardi. Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937, 1938), Nielsen (1976a), Tantanasiriwong
(1979), Nateewathana et al. (1981), Kurozumi et al. (1989), Bernard et al. (1993), Willan (1993), Bussarawit
(1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University
(2000, 2006), Gemert (2003), Duangdee (2004, 2007), Duangdee et al. (2007, 2009, 2011), Taylor et al. (2007),
Nabhitabhata (2009), Sanpanich & Duangdee (2009), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF
(2019a, 2019b), J. D. Taylor et al. (unpublished data).
Gari ambigua (Reeve, 1857). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Gari amethystus (Wood, 1815). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich (2011).
Gari anomala (Deshayes, 1855). Also reported as: Psammobia anomala Deshayes, 1854; and Psammobia tenuis
Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Scott (1995), Nabhitabhata (2009).
Gari castrensis (Spengler, 1794). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Gari crassula (Deshayes, 1855). Reported as: Gari (Crassulobia) crassula (G. P. Deshayes, 1855). No Thai locality
provided. Reported by: Huber (2010).
Gari elongata (Lamarck, 1818). Also reported as: Gari (Psammotaena) elongata (Lamarck, 1818); Psammotaea
elongata (Lamarck, 1818); and Solenotellina violacea Lamarck. Gulf of Thailand, Andaman Sea. Reported by:
Suvatti (1938), Nateewathana et al. (1981), Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al.
(1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), ONEP (2007), Taylor et al. (2007),
Nabhitabhata (2009), BEDO (2017a), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Gari inflata (Bertin, 1880). Reported as: Gari (Psammotaena) inflata (Bertin, 1880). Gulf of Thailand. Reported
by: Gemert (2003).
Gari lessoni (Blainville, 1826). Reported as: Gari cf. lessoni (Blainville, 1826). Andaman Sea. Reported by:
Nabhitabhata (2009), GBIF (2019a).
Gari livida (Lamarck, 1818). Also reported as: Psammobia zonalis Lamarck, 1818. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Nabhitabhata (2009), Sanpanich (2011).
Gari maculosa (Lamarck, 1818). Also reported as: Gari (Gari) maculosa (Lamarck, 1818); Psammobia corrugata
Deshayes, 1855; and Psammobia scabra Chemnitz, 1782. Gulf of Thailand, Andaman Sea. Reported by: Lynge
(1909), Suvatti (1938), Nateewathana et al. (1981), Willan (1993), Aungtonya et al. (1999), Gemert (2003),
Kasetsart University (2006), Duangdee (2007), Duangdee et al. (2007, 2011, 2013a), Nabhitabhata (2009),
Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO (2017a), Sanpanich & Duangdee (2018).
Gari minor (Deshayes, 1854). Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999),
Gemert (2003), Swennen et al. (2001).
Gari oriens (Deshayes, 1855). Gulf of Thailand. Reported by: Sanpanich & Duangdee (2009), Sanpanich (2011).
24
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Gari pallida (Deshayes, 1831). Also reported as: Gari (Gari) pallida (Deshayes, 1855); and Gari weinkauffi
(Crosse, 1864). Gulf of Thailand, Andaman Sea. Reported by: Willan (1993), Nabhitabhata (2009), Sanpanich
(2011), BEDO (2017a), GBIF (2019a).
Gari pulcherrima (Deshayes, 1855). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (2011), BEDO (2017a).
Gari radiata (Dunker, 1845). Also reported as: Psammobia radiata Dunker, 1845. Gulf of Thailand, Andaman Sea.
Reported by: Nateewathana et al. (1981), Bernard et al. (1993), Aungtonya et al. (1999), Swennen et al. (2001),
Nabhitabhata (2009), BEDO (2017a).
Gari sibogai Prashad, 1932. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya
et al. (1999), Nabhitabhata (2009).
Gari squamosa (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Gari togata (Deshayes, 1855). Andaman Sea. Reported by: GBIF (2019a).
Gari truncata (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Kilburn
& Hylleberg (1998), Kasetsart University (2000), Swennen et al. (2001), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), Sanpanich (2011), BEDO (2017a).
Hiatula adamsii (Reeve, 1857). Reported as: Soletellina adamsii Reeve, 1857; and Sanguinolaria (Soletellina)
adamsi Deshayes. Gulf of Thailand. Reported by: Suvatti (1938), Swennen et al. (2001), Nabhitabhata (2009).
Hiatula atrata (Reeve, 1857). Reported as: Soletellina cf. atrata Reeve, 1857. Gulf of Thailand. Reported by:
Gemert (2003).
Hiatula chinensis (Mörch, 1853). Reported as: Solenotellina truncata Gmelin; and Hiatula planulata (Reeve, 1857).
Gulf of Thailand. Reported by: Suvatti (1938), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Hiatula connectens (Martens, 1865). Andaman Sea. Reported by: GBIF (2019a, 2019b).
Hiatula diphos (Linnaeus, 1771). Also reported as: Soletellina diphos (Linnaeus, 1771). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Robba
et al. (2002), Willan et al. (2007), Nabhitabhata (2009), BEDO (2017a).
Hiatula rosea (Gmelin, 1791). Reported as: Hiatula ruppelliana (Reeve, 1857). Gulf of Thailand. Reported by:
Nabhitabhata (2009), Sanpanich (2011).
Hiatula petalina (Deshayes, 1855). Reported as: Soletellina petalina (G. P. Deshayes, 1855). No Thai locality
provided. Reported by: Huber (2010).
Family Semelidae
Abra fragillima (Issel, 1869). Reported as: Semele fragillima (Issel, 1869). Gulf of Thailand. Reported by: Gemert
(2003).
Abra inanis Prashad, 1932. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Abra scotti M. Huber, 2010. Reported as: Tellina declivis Sowerby, 1868. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009).
Cumingia mutica G. B. Sowerby I, 1833. Reported as: Cumingia striata A. Adams, 1850. Gulf of Thailand. Reported
by: Printrakoon et al. (2016).
Ervilia bisculpta Gould, 1861. Reported as: Spondervilia bisculpta (Gould, 1861); and Ervillia bisculpta Deshayes,
1854. Gulf of Thailand. Reported by: Lynge (1909), Gemert (2003), Robba et al. (2004), Nabhitabhata (2009).
Leptomya gravida Hanley, 1879. Andaman Sea. Reported by: Duangdee et al. (2013a).
Leptomya psittacus Hanley, 1882. Andaman Sea. Reported by: Duangdee et al. (2013b).
Leptomya rostrata (H. Adams, 1868). Reported as: Leptomya spectabilis Deshayes, 1882; and Leptomya
subrostrata (Issel, 1869). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Gemert (2003), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Sanpanich (2011), BEDO (2017a).
Semele amabilis Adams, 1854. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nabhitabhata (2009),
Sanpanich (2011).
Semele australis (G. B. Sowerby I, 1832). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee
(2009), Sanpanich (2011).
Semele carnicolor (Hanley, 1845). Also reported as: Semele jukesii (Reeve, 1853). Gulf of Thailand, Andaman Sea.
Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Sanpanich (1998),
Aungtonya et al. (1999), Robba et al. (2002), Gemert (2003), Taylor et al. (2007), Simone & Wilkinson (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Duangdee et al. (2011), Sanpanich (2011), BEDO
(2017a), GBIF (2019a, 2019b).
Semele casta (Reeve, 1853). Reported as: Semele casta A. Adams, 1853. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Duangdee et al. (2011).
Semele cordiformis (Holten, 1802). Also reported as: Semele sinensis Adams, 1854. Gulf of Thailand, Andaman
Sea. Reported by: Suvatti (1937, 1938), Nateewathana et al. (1981), Sanpanich (1998, 2011), Aungtonya et al.
(1999), Robba et al. (2002), Simone & Wilkinson (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009),
BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Semele crenulata (Sowerby, 1853). Gulf of Thailand. Reported by: Nabhitabhata (2009).
Semele duplicata (Reeve, 1853). Gulf of Thailand. Reported by: Sanpanich (2011), Sanpanich & Duangdee
(2009, 2018).
Semele exarata (A. Adams & Reeve, 1850). Gulf of Thailand. Reported by: Duangdee et al. (2011).
Semele jucunda (Reeve, 1853). Reported as: Semele monilis Tate, 1891. Gulf of Thailand, Andaman Sea. Reported
by: Sanpanich (2011).
Semele lamellosa (Sowerby, 1830). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009,
2018), Sanpanich (2011), Duangdee et al. (2013a).
Semele scabra (Hanley, 1843). Reported as: Semele sponsa Adams. Gulf of Thailand. Reported by: Suvatti (1937,
1938), Kurozumi et al. (1989), Nabhitabhata (2009).
Semele zebuensis (Hanley, 1843). Reported as: Semele duplicata (Sowerby, 1833). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Theora lata (Hinds, 1843). Also reported as: Theora (Theora) lata (Hinds, 1843). Gulf of Thailand. Reported by:
Lynge (1909), Bernard et al. (1993), Robba et al. (2002, 2003), Nabhitabhata (2009), Negri et al. (2014), GBIF
(2019b).
Family Solecurtidae
Azorinus abbreviatus (Gould, 1861). Gulf of Thailand, Andaman Sea. Reported by: Scott (1995), Carpenter & Niem
(1998), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009).
Azorinus chamasolen (da Costa, 1778). Reported as: Solecurtus (Azor) emarginatus Spengler, 1782. Gulf of
Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Azorinus coarctatus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001),
Nabhitabhata (2009), BEDO (2017a).
Azorinus minutus (Dunker, 1862). Andaman Sea. Reported by: BEDO (2017a).
Azorinus scheepmakeri (Dunker, 1852). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001),
Nabhitabhata (2009), GBIF (2019a).
Solecurtus australis (Dunker, 1862). Andaman Sea. Reported by: BEDO (2017a).
Solecurtus consimilis Kuroda & Habe in Habe, 1961. Gulf of Thailand. Reported by: BEDO (2017a).
Solecurtus divaricatus (Lischke, 1869). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Aungtonya et al. (1999), ONEP (2007), Duangdee et al. (2009), Nabhitabhata (2009), GBIF (2019a).
Solecurtus exaratus (Philippi, 1849). Also reported as: Solecurtus wilsoni Tryon, 1870. Gulf of Thailand. Reported
by: Sanpanich (1998), Robba et al. (2002), Nabhitabhata (2009), Huber (2010).
Solecurtus philippinarum (Dunker, 1862). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001),
Nabhitabhata (2009), BEDO (2017a).
Solecurtus rhombus (Spengler, 1794). Reported as: Solecurtus quoyi Reeve, 1874; and Solecurtus (Macha) rhombus
Spengler, 1782. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nabhitabhata (2009), Sanpanich
(2011).
Solecurtus subcandidus Sturany, 1899. Gulf of Thailand. Reported by: Sanpanich (2011).
Solecurtus sulcatus (Dunker, 1862). Andaman Sea. Reported by: BEDO (2017a).
Family Tellinidae
Abranda casta (Hanley, 1844). Reported as: Arcopella casta (Hanley, 1844); Tellina (Arcopella) casta Hanley, 1844;
and Pinguitellina casta (Hanley, 1844). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard
et al. (1993), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a).
Abranda languida (E. A. Smith, 1885). Reported as: Tellina languida Smith, 1885; and Macoma (Pinguimacoma)
languida (Smith, 1885). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002, 2003), Nabhitabhata
(2009).
Angulinides opalinus (Gmelin, 1791). Reported as: Tellina (Tellinides) opalina Gmelin, 1791. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata
(2009).
Angulus lanceolatus (Gmelin, 1791). Reported as: Tellina lanceolata Gmelin, 1791; and Tellina (Angulus)
lanceolata Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Duangdee (2004),
Nabhitabhata (2009), Sanpanich (2011), Huber et al. (2015).
Apolymetis meyeri (Dunker, 1846). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO
(2017a), GBIF (2019b).
Arcopaginula inflata (Gmelin, 1791). Reported as: Tellina inflata Gmelin, 1791; and Tellina (Arcopaginula)
inflata (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981),
Bernard et al. (1993), Sanpanich (1998), Aungtonya et al. (1999), Swennen et al. (2001), D uangdee (2004,
2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
26
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Arcopella isseli (H. Adams, 1871). Reported as: Tellina (Acropagia) isseli H. Adams, 1870. Gulf of Thailand.
Reported by: Lynge (1909), Bernard et al. (1993), Gemert (2003), Nabhitabhata (2009).
Afsharius berghi (Lynge, 1909). Reported as: Tellina (Moerella) berghi Lynge, 1909. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Afsharius patagiatus (Prashad, 1932). Reported as: Tellina patagiatus (Prashad, 1932). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Cadella cf. delta (Yokoyama, 1922). Gulf of Thailand. Reported by: Gemert (2003).
Cadella diluta (E. A. Smith, 1885). Reported as: Tellina diluta Smith, 1885. Gulf of Thailand. Reported by:
Sanpanich (2011).
Cadella hainanensis Scarlato, 1965. No Thai locality provided. Reported by: Huber et al. (2015).
Cadella martensi (Lynge, 1909). Reported as: Tellina (Merisca) martensi Lynge, 1909. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Cadella nucleolus (Deshayes, 1855). Reported as: Tellina smithi Lynge, 1909; and Cadella smithi (Lynge, 1909).
Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993).
Cadella obtusalis (Deshayes, 1855). Reported as: Tellina (Moerella) obtusalis Deshayes, 1854. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Cadella semen Hanley, 1845. Reported as: Tellina (Cadella) semen Hanley, 1844; and Tellina (Moerella) semen
Hanley, 1844. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Robba et al.
(2002), Gemert (2003), Simone & Wilkinson (2008), Nabhitabhata (2009).
Clathrotellina carnicolor (Hanley, 1846). Reported as: Tellina (Acropagia) carnicolor Hanley, 1846; and Tellina
(Clathrotellina) carnicolor Hanley, 1846. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002),
Nabhitabhata (2009), Sanpanich & Duangdee (2009).
Clathrotellina elegantissima (E. A. Smith, 1885). Reported as: Tellina elegantissima E. A. Smith, 1885; and
Tellina (Acropagia) elegantissima Smith, 1885. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nabhitabhata (2009), Sanpanich (2011).
Clathrotellina pretium (Salisbury, 1934). Reported as: Arcopagia pretium (Salisbury, 1934); Tellina pretium
(Salisbury, 1934); and Tellina (Cyclotellina) prestiosa (Deshayes, 1855). Gulf of Thailand, Andaman Sea. Reported
by: Nateewathana et al. (1981), Aungtonya et al. (1999), Robba et al. (2004), Nabhitabhata (2009), BEDO (2017a).
Cyclotellina remies (Linnaeus, 1758). Reported as: Tellina remies Linnaeus, 1758; and Tellina (Cyclotellina)
remies Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976a), Tantanasiriwong (1979),
Morris & Purchon (1981), Nateewathana et al. (1981), Nateewathana (1995), Carpenter & Niem (1998),
Aungtonya et al. (1999), Gemert (2003), Duangdee (2004), Kasetsart University (2006), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich (2011), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019a).
Dallitellina deshayesii (Hanley, 1844). Reported as: Tellina (Tellinella) deshayesii Hanley, 1844. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009), Huber et al. (2015).
Dallitellina rostrata (Linnaeus, 1758). Also reported as: Tellina rostrata Linnaeus, 1758; Tellina (Pharaonella)
rostrata Linnaeus, 1758; Tellina spengleri Gmelin; Tellina (Tellinella) spengleri Gmelin, 1791; Tellina spengleri
(Gmelin, 1791); and Dallitellina spengleri (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Morlet
(1889), Lynge (1909), Suvatti (1938), Bernard et al. (1993), Kilburn & Hylleberg (1998), Aungtonya et al.
(1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Robba et al. (2002), Duangdee (2004), Wells
(2005), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Dallitellina winckworthi (Salisbury, 1934). Reported as: Tellina winckworthi Salisbury, 1934. Gulf of Thailand.
Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Exotica cygnus (Hanley, 1844). Reported as: Macoma cygnus (Hanley, 1844); Tellina cygnus Hanley, 1846; Tellina
(Peronaea) cygnus Hanley, 1844; and Macoma (Pinguimacoma) cygnus (Hanley, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Kilburn & Hylleberg (1998), Nabhitabhata (2009), Sanpanich (2011).
Exotica triradiata (H. Adams, 1871). Reported as: Tellina triradiata H. Adams, 1871. Gulf of Thailand. Reported
by: Sanpanich (2011).
Eurytellina angulosa (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017a).
Hanleyanus oblongus (Gmelin, 1791). Reported as: Tellina emarginata G. B. Sowerby I, 1825; Tellides carinata
(Spengler, 1798); Tellina (Tellides) carinata Spengler, 1798; and Tellina (Angulus) emarginata (Sowerby, 1825).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Swennen et al. (2001), Robba
et al. (2002), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011), GBIF (2019b).
Hanleyanus vestalis (Hanley, 1844). Reported as: Psammotreta solenella (Deshayes, 1854); Tellina vestalis
Hanley, 1844; Tellina (Tellinides) vestalis Hanley, 1844; and Tellina (Angulus) vestalis Hanley, 1844. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Robba et al. (2002), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a).
Herouvalia caelata (A. Adams, 1854). Reported as: Tellina fabrefacta Pilsbry, 1904. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Iridona iridescens (Benson, 1842). Reported as: Moerella iridescens (Benson, 1842); and Tellina (Angulus)
iridescens W.H. Benson, 1842. Gulf of Thailand. Reported by: Lynge (1909), Duangdee et al. (2007), Nabhitabhata
(2009).
27
The Marine and Estuarine Molluscs of Thailand
Jactellina clathrata (Deshayes, 1835). Reported as: Exotica clathrata Deshayes, 1835. Gulf of Thailand, Andaman
Sea. Reported by: Wells (2005), Sanpanich & Duangdee (2009, 2018), Sanpanich (2011), Duangdee et al. (2013b).
Jactellina secunda (Bertin, 1878). Reported as: Loxoglypta secunda (Bertin, 1878). Gulf of Thailand. Reported by:
Gemert (2003).
Jactellina subpallida (E. A. Smith, 1891). No Thai locality provided. Reported by: Huber et al. (2015).
Jactellina transcalpta (G. B. Sowerby III, 1915). Also reported as: Jactellina (Loxoglypta) transculpta (G. B.
Sowerby III, 1915). No Thai locality provided. Reported by: Kurozumi et al. (1989), Huber et al. (2015).
Jitlada bertini (Jousseaume, 1895). Reported as: Arcopagia yemenensis (Melvill, 1898). Gulf of Thailand. Reported
by: Robba et al. (2002, 2003), Nabhitabhata (2009).
Jitlada hanleyi M. Huber, Langleit & Kreipl, 2015. Reported as: Tellina (Angulus) corbuloides Hanley, 1844. Gulf
of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Jitlada philippinarum (Hanley, 1844). Reported as: Tellina australis Deshayes, 1854; and Tellina (Macomona)
australis Deshayes, 1854. Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002, 2003), Nabhitabhata
(2009), Sanpanich (2011), GBIF (2019b).
Leporimetis coarctata (Philippi, 1845). Reported as: Florimetis coarctata (Philippi, 1845); and Tellina (Metis)
coarctata Philippi, 1845. Gulf of Thailand. Reported by: Lynge (1909), Swennen et al. (2001), Nabhitabhata
(2009), J. D. Taylor et al. (unpublished data).
Leporimetis ephippium (Spengler, 1798). Reported as: Psammotreta ephippium (Spengler, 1798); and Apolymetis
eplippium (Spengler, 1798). Gulf of Thailand. Reported by: Carpenter & Niem (1998), Sanpanich (1998),
Nabhitabhata (2009).
Leporimetis papyracea (Gmelin, 1791). Reported as: Leporimetis papyracea Gmelin, 1791. Gulf of Thailand.
Reported by: Duangdee (2004).
Leporimetis spectabilis (Hanley, 1844). Also reported as: Apolymetis spectabilis (Hanley, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011).
Macalia bruguieri (Hanley, 1844). Reported as: Macalia cf. bruguiere (Hanley, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Gemert (2003), Nabhitabhata (2009),
Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Macomona deltoidalis (Lamarck, 1818). Reported as: Tellina deltoidalis Lamarck, 1818; and Tellina (Macomana)
deltoidalis Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University (2006), Duangdee
(2007), ONEP (2007), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich (2011).
Macomopsis chinensis (Hanley, 1845). Reported as: Abra hainanensis Scarlato, 1965. No Thai locality stated.
Reported by: Bernard et al. (1993).
Macomopsis moluccensis (Martens, 1865). Reported as: Tellina (Macoma) moluccensis Martens, 1865; Macoma
moluccensis Martens, 1868; and Macoma retrosa Sowerby, 1867. Gulf of Thailand, Andaman Sea. Reported by:
Lynge (1909), Nabhitabhata (2009), BEDO (2017a).
Moerella virgulata (Hanley, 1845). Reported as: Exotica virgulata (Hanley, 1844). Andaman Sea. Reported by:
Wells (2005), Duangdee et al. (2013b).
Nitidotellina lischkei M. Huber, Langleit & Kreipl, 2015. Reported as: Tellina (Fabulina) minuta Lischke, 1874.
Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya
et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Nitidotellina lux (Hanley, 1844). Reported as: Tellina (Angulus) lux Hanley, 1844; and Tellina (Angulus) rhomboides
Quoy & Gaimard, 1833. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Nitidotellina pallidula (Lischke, 1871). Reported as: Tellina (Moerella) pallidula Lischke, 1871. Gulf of Thailand.
Reported by: Robba et al. (2002), Nabhitabhata (2009), Negri et al. (2014).
Nitidotellina unifasciata (G. B. Sowerby II, 1867). Reported as: Tellina (Moerella) nitens Deshayes, 1854;
Nitidotellina nitens (Deshayes, 1855); Tellina cf. nitens Deshayes, 1854; and Moerella cf. nitens (Deshayes,
1854). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002), Gemert (2003), Meyer et al. (2008),
Simone & Wilkinson (2008), Nabhitabhata (2009).
Nitidotellina valtonis (Hanley, 1844). Reported as: Tellina valtonis (Hanley, 1844); Tellina (Moerella) valtonis
Hanley, 1844; Tellinides valtonis (Hanley, 1855); Tellina (Angulus) iridella Martens, 1865; and Moerella cf. iridella
Martens, 1865. Gulf of Thailand. Reported by: Lynge (1909), Scott (1995), Robba et al. (2002), Nabhitabhata
(2009), Negri et al. (2014), J. D. Taylor et al. (unpublished data).
Oudardia virgo (Hanley, 1844). Reported as: Tellina virgo Handy, 1844. Gulf of Thailand. Reported by: Swennen
et al. (2001), Nabhitabhata (2009).
Pharaonella pharaonis (Hanley, 1844). Also reported as: Tellina pharaonis Hanley, 1844. Gulf of Thailand,
Andaman Sea. Reported by: Sanpanich (2011).
Pharaonella sieboldii (Deshayes, 1855). Reported as: Tellina sieboldii Deshayes, 1855; and Tellina (Pharaonella)
perrieri (Bertin, 1878). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et
al. (1999), Nabhitabhata (2009).
Phylloda foliacea (Linnaeus, 1758). Reported as: Tellina (Phylloda) foliacea Linnaeus, 1758; and Tellina (Phyllodosa)
foliacea Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Aungtonya et
al. (1999), Duangdee (2004), Nabhitabhata (2009), GBIF (2019a).
28
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Phyllodina tenuisculpta (G. B. Sowerby II, 1868). Reported as: Tellina (Macoma) tenuisculta Sowerby, 1868. Gulf
of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Pinguitellina altissima (Smith, 1906). No Thai locality provided. Reported by: Huber et al. (2015).
Pinguitellina nux (Hanley, 1844). Reported as: Tellina (Acropagia) nux Hanley, 1844. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Pinguitellina pinguis (Hanley, 1844). Reported as: Tellina (Acropagia) pinguis Hanley, 1844; and Tellina
(Pinguitellina) pinguis Hanley, 1844. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Robba et al.
(2002), Gemert (2003), Nabhitabhata (2009), Taylor (2009), Negri et al. (2014), GBIF (2019a).
Pinguitellina pudica (Hanley, 1844). Reported as: Tellina pudica Hanley, 1844; and Arcopagia pudica (Hanley,
1844). Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993), Subba Rao & Dey (2000), Robba et al.
(2002, 2003), Nabhitabhata (2009), Negri et al. (2014).
Pinguitellina robusta (Hanley, 1844). Andaman Sea. Reported by: Nabhitabhata (2009).
Pristipagia radians (Deshayes, 1855). Reported as: Tellina radians Deshayes, 1855. Gulf of Thailand, Andaman
Sea. Reported by: Sanpanich & Duangdee (2009), Sanpanich (2011), Duangdee et al. (2013b), BEDO (2017a).
Psammacoma candida (Lamarck, 1818). Reported as: Macoma (Psammacoma) candida (Lamarck, 1818); and
Macoma candida (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Bernard et al. (1993), Aungtonya
et al. (1999), Swennen et al. (2001), Nabhitabhata (2009).
Psammacoma fallax (Bertin, 1878). Reported as: Tellina (Macoma) fallax Bertin, 1878; and Macoma
(Psammacoma) fallax Bertin, 1878. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002),
Nabhitabhata (2009), Negri et al. (2014).
Psammacoma gubernaculum (Hanley, 1844). Reported as: Tellina (Macoma) papyracea Spengler, 1798;
Macoma gubernaculums (Hanley, 1844); and Psammotreta (Pseudometis) praerupta (Salisbury, 1934). Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009), GBIF (2019a,
2019b).
Psammotreta maluccensis (Martens, 1865). No Thai locality provided. Reported by: Bernard et al. (1993).
Psammotreta psammotella (Lamarck, 1818). Reported as: Tellina (Tellinides) psammotella Lamarck, 1818; and
Tellina psammotella Lamarck, 1818. Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993),
Nabhitabhata (2009).
Pseudopsammobia simplex (G. B. Sowerby III, 1894). Also reported as: Gari (Gari) simplex (Sowerby, 1894);
and Psammobia simplex (Sowerby, 1894). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002),
Nabhitabhata (2009), Negri et al. (2014), GBIF (2019b).
Pulvinus micans (Hanley, 1844). Reported as: Tellina (Peronaea) micans Hanley, 1844; and Psammotreta micans
(Hanley, 1844). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Subba Rao &
Dey (2000), Nabhitabhata (2009).
Quadrans bougei (G. B. Sowerby III, 1909). Reported as: Tellina bougei Sowerby, 1909. Andaman Sea. Reported
by: Sanpanich & Duangdee (2009), Sanpanich (2011).
Quadrans gargadia (Linnaeus, 1758). Reported as: Tellina gargadia (Linnaeus, 1758); and Tellina (Quadrans)
gargadia Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Duangdee (2004), Wells
(2005), Nabhitabhata (2009).
Quadrans spinosus (Hanley, 1844). Reported as: Tellina (Quadrans) minor V. Bertin, 1878. Gulf of Thailand.
Reported by: Lynge (1909), Kurozumi et al. (1989), Nabhitabhata (2009).
Quidnipagus palatam Iredale, 1929. Reported as: Tellina palatum (Iredale, 1929); Tellina (Quidnipagus) palatum
(Iredale, 1929); Tellina (Tellinella) rugosa Born, 1778; and Tellina (Arcopagia) rugosa Born, 1778. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937), Nielsen (1976b), Tantanasiriwong (1979),
Morris & Purchon (1981), Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999),
Gemert (2003), Duangdee (2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Sanpanich (2011),
Hylleberg & Aungtonya (2013), BEDO (2017a).
Salmacoma vappa Iredale, 1929. Andaman Sea. Reported by: GBIF (2019a).
Scissulina dispar (Conrad, 1837). Also reported as: Tellina (Angulus) dispar Conrad, 1837; and Macoma dispar
(Conrad, 1837). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Scutarcopagia linguafelis (Linnaeus, 1758). Reported as: Tellina linguafelis Linnaeus, 1758. Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009).
Scutarcopagia pulcherrima (G. B. Sowerby I, 1825). Reported as: Tellina pulcherrima Sowerby, 1825; and
Tellina (Tellinella) pulcherrima Sowerby, 1825. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya & Hylleberg (1998), Aungtonya et al. (1999),
Nabhitabhata (2009), BEDO (2017a), GBIF (2019b).
Scutarcopagia scobinata (Linnaeus, 1758). Also reported as: Tellina (Scutarcopagia) scobinata Linnaeus, 1758.
Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon
(1981), Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Wells (2005),
Nabhitabhata (2009), Duangdee et al. (2013b), Hylleberg & Aungtonya (2013), BEDO (2017a).
29
The Marine and Estuarine Molluscs of Thailand
Scutarcopagia verrucosa (Hanley, 1844). Reported as: Tellina botanica (Hedley, 1918); and Tellina verrucosa
(Hanley, 1844). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011), BEDO
(2017a).
Semelangulus liratus M. Huber, Langleit & Kreipl, 2015. Gulf of Thailand. Reported by: Huber et al. (2015).
Semelangulus tenuiliratus (G. B. Sowerby II, 1867). Reported as: Tellina (Moerella) semitorta Sowerby, 1867;
and Tellina (Moerella) tenuilirata Sowerby, 1867. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata
(2009).
Serratina capsoides (Lamarck, 1818). Also reported as: Tellina capsoides Lamarck, 1818; Tellina (Serratina)
capsoides (Lamarck, 1818); Pristis capsoides (Lamarck, 1818); Tellina (Pistris) capsoides Lamarck, 1818; and
Tellina (Acropagia) capsoides Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Robba et al. (2002), Gemert (2003),
Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF
(2019b), J. D. Taylor et al. (unpublished data). The Andaman records of Serratina capsoides may actually be
Serratina pristis (Ujino & Matsukuma, 2013).
Serratina jonasi (Philippi, 1847). Reported as: Tellina serricostata Tokunaga, 1906. Gulf of Thailand. Reported
by: Sanpanich (2011).
Serratina perplexa (Hanley, 1844). Reported as: Tellina (Tellinella) hungerfordi Sowerby, 1894. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009), Huber et al. (2015).
Serratina pristis (Lamarck, 1818). Andaman Sea. Reported by: Ujino & Matsukuma (2013).
Serratina siamensis (Martens, 1860). Reported as: Tellina siamensis Martens, 1860. Gulf of Thailand. Reported
by: Suvatti (1938), Nabhitabhata (2009), Huber et al. (2015).
Sylvanus assimilis (Hanley, 1844). Reported as: Exotica assimilis (Hanley, 1844). Gulf of Thailand. Reported by:
Sanpanich (2011).
Sylvanus lilium (Hanley, 1844). Reported as: Exotica donaciformis (Deshayes, 1854); Exotica (Exotica)
donaciformis (Deshayes, 1854); Macoma awaijiensis (Sowerby III, 1914); Macoma awaijiensis (Sowerby III,
1914); Semelangulus lilium (Hanley, 1844); and Tellina lilium Hanley, 1844. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Swennen et al. (2001), Gemert (2003), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a).
Tellina candida Gmelin, 1791. Reported as: Tellina (Macoma) candida Lamarck, 1818. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Tellina cruciata (Spengler, 1798). Reported as: Tellinella cruciata (Spengler, 1798); and Tellinella stuarella
(Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Duangdee (2007),
BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a).
Tellina irus Hanley, 1845. Reported as: Tellina yantaiensis. Gulf of Thailand. Reported by: Suvatti (1937, 1938),
Nabhitabhata (2009).
Tellina plicata Valenciennes, 1827. Gulf of Thailand. Reported by: Sanpanich (2011).
Tellina rhodon Hanley, 1844. Reported as: Moerella cf. rhodon Hanley, 1844. Andaman Sea. Reported by:
Nabhitabhata (2009).
Tellina vernalis Hanley, 1844. Reported as: Tellina (Tellinides) vernalis Hanley, 1844. Gulf of Thailand. Reported
by: Nabhitabhata (2009), BEDO (2017a).
Tellina yemenensis (Melvill, 1898). Reported as: Arcopagia yemenensis (Melvill, 1898). Gulf of Thailand. Reported
by: Negri et al. (2014).
Tellinangulus aethiopicus (Thiele, 1931). Reported as: Tellina (Tellinangulus) aethiopica Thiele & Jaeckel, 1931.
Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009).
Tellinella cruciata (Spengler, 1798). Reported as: Tellina staurella Lamarck, 1818; and Tellina (Tellinella)
staurella Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999),
Subba Rao & Dey (2000), Duangdee et al. (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a),
Sanpanich & Duangdee (2018), GBIF (2019a).
Tellinella crucigera (Lamarck, 1818). Also reported as: Tellina exculta Gould, 1850. Gulf of Thailand, Andaman
Sea. Reported by: Wells (2005), Sanpanich (2011), Duangdee et al. (2013b).
Tellinella philippii (Anton, 1844). Also reported as: Tellina (Pharaonella) rastella Hanley, 1844; and Tellina
rastellum Hanley, 1844. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bernard et
al. (1993), Nabhitabhata (2009).
Tellinella virgata (Linnaeus, 1758). Reported as: Tellina virgata (Linnaeus, 1758); and Tellina (Tellinella)
virgata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Bussarawit
(1995), Carpenter & Niem (1998), Subba Rao & Dey (2000), Gemert (2003), Wells (2005), Duangdee et al.
(2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Tellinides coccineus (Gmelin, 1791). Gulf of Thailand. Reported by: Kurozumi et al. (1989).
Tellinides conspicuus (Hanley, 1846). Gulf of Thailand. Reported by: Huber et al. (2015).
Tellinides margaritinus (Lamarck, 1818). Reported as: Tellina umbonella Lamarck, 1818. Gulf of Thailand,
Andaman Sea. Reported by: Sanpanich (2011), GBIF (2019a, 2019b).
30
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Tellinides roseotinctus M. Huber, Langleit & Kreipl, 2015. Andaman Sea. Reported by: Huber et al. (2015).
Tellinides sinuatus (Spengler, 1798). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998),
Nabhitabhata (2009).
Tellinides striatus (Gmelin, 1791). Reported as: Tellina ovalis G. B. Sowerby I, 1825; and Tellina (Tellinides) ovalis
Sowerby, 1825. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Swennen et al. (2001), Gemert
(2003), Duangdee (2004), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Tellinides timorensis Lamarck, 1818. Also reported as: Tellina (Tellinides) timorensis (Lamarck, 1818); and
Tellina timorensis (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Swennen et al. (2001), Robba et al. (2002), Duangdee (2004), Kasetsart University (2006), Nabhitabhata
(2009), Sanpanich (2011), Sanpanich & Duangdee (2018), GBIF (2019b).
Tellinimactra edentula (Spengler, 1798). Reported as: Lutraria tellinoides Lamarck, 1818; Psammotreta
edentula (Spengler, 1797); Macoma edentula (Broderip & Sowerby); Psammotreta (Tellinimactra) edentula
(Spengler, 1798); and Metis angulata Chemnitz. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1938),
Robba et al. (2002), Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), GBIF (2019a, 2019b).
Tellinota imbellis (Hanley, 1844). Reported as: Tellina imbellis Hanley, 1844. Gulf of Thailand. Reported by:
Sanpanich (2011).
Tonganaella perna (Spengler, 1798). Reported as: Pharaonella perna (Spengler, 1798). Gulf of Thailand.
Reported by: Kurozumi et al. (1989).
Order Galeommatida
Superfamily Galeommatoidea
Family Galeommatidae
Aenictomya mirabilis (Lynge, 1909). Reported as: Corbula mirabilis Lynge, 1909. Gulf of Thailand. Reported by:
Lynge (1909), Bernard et al. (1993), Nabhitabhata (2009).
Aenictomya quadrangularis (Lynge, 1909). Reported as: Sphenia quadrangularis. Gulf of Thailand. Reported by:
Lynge (1909), Nabhitabhata (2009).
Amphilepida aurantia (Deshayes, 1835). Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999),
Gemert (2003), Nabhitabhata (2009).
Amphilepida callipareia (Melvill, 1899). Gulf of Thailand. Reported by: Gemert (2003).
Afrophysema tuticorina (Taylor & Glover, 2005). Andaman Sea. Reported by: Huber (2015).
Cymatioa agilis (Lützen & Nielsen, 2005). Andaman Sea. Reported by: Huber (2015), BEDO (2017a).
Ephippodonta gigas Kubo, 1996. Reported as: Ephippodonta (Ephippodontina) gigas Kubo, 1996. Andaman Sea.
Reported by: Lützen & Nielsen (2005), BEDO (2017a).
Ephippodontina murakamii (Kuroda, 1945). Reported as: Ephippodonta murakamii (Kuroda, 1945). Gulf of
Thailand. Reported by: Kurozumi et al. (1989).
Ephippodontomorpha orientalis (Lützen & Nielsen, 2005). Reported as: Aclistothyra orientalis Lützen & Nielsen
(2005). Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013).
Galeomma ambigua Deshayes, 1856. Reported as: Galeomma (Galeomma) ambigua Deshayes, 1856. Andaman
Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013), BEDO (2017a).
Galeomma augusta Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Galeomma macrochisma Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Lepirodes ambiguus (Deshayes, 1856). Reported as: Scintilla faba Deshayes, 1856. Andaman Sea. Reported by:
Subba Rao & Dey (2000), BEDO (2017a).
Lepirodes aurantius (Lamarck, 1818). Also reported as: Galeomma (Amphilepida) obockensis (Jousseaume,
1888). Andaman Sea. Reported by: Nateewathana et al. (1981), Lützen & Nielsen (2005), BEDO (2017a).
Lepirodes formosus (Deshayes, 1856). Reported as: Galeomma formosa Deshayes, 1856. Gulf of Thailand.
Reported by: Gemert (2003).
Lepirodes layardi (Deshayes, 1856). Reported as: Galeomma layardi Deshayes, 1856; and Galeomma (Galeomma)
layardi Deshayes, 1856. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013),
BEDO (2017a).
Lepirodes phuketi (Lützen & Nielsen, 2005). Reported as: Galeomma phuketi Lützen & Nielsen (2005). Andaman
Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013).
Nudiscintilla glabra Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Pseudogaleomma japonica (A. Adams, 1862). Andaman Sea. Reported by: Lützen & Nielsen (2005), Taylor et al.
(2007), Hylleberg & Aungtonya (2013), BEDO (2017a).
Scintilla agilis Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013).
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The Marine and Estuarine Molluscs of Thailand
Scintilla candida Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009), Huber
(2015).
Scintilla cuvieri Deshayes, 1856. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya
(2013), BEDO (2017a).
Scintilla hyalina Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Scintilla hydatina Deshayes, 1855. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Scintilla imperatoris Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla longitentaculata Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg
& Aungtonya (2013), BEDO (2017a).
Scintilla macrodactylus Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg
& Aungtonya (2013), Huber (2015), BEDO (2017a).
Scintilla minor Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla mortoni Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla nitidella Habe, 1962. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg & Aungtonya
(2013), BEDO (2017a).
Scintilla ovalis Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), WoRMS Editorial Board (2019) lists this as a temporary name as it is a junior homonym of
Scintilla ovalis Turton, 1932.
Scintilla papillosa Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla philippinensis Deshayes, 1855. Andaman Sea. Reported by: Lynge (1909), Lützen & Nielsen (2005),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017a).
Scintilla sannio Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla siamense Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla unicornia Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Scintilla verrucosa Lützen & Nielsen, 2005. Andaman Sea. Reported by: Lützen & Nielsen (2005), Hylleberg
& Aungtonya (2013), BEDO (2017a).
Scintilla violescens Kuroda & Habe, 1961. Andaman Sea. Reported by: Lützen & Nielsen (2005), Huber (2015),
BEDO (2017a).
Scintillula elongata (Lynge, 1909). Reported as: Erycina elongata; and Marikella elongata (Lynge, 1909). Gulf of
Thailand. Reported by: Lynge (1909), Robba et al. (2002), Gemert (2003), Nabhitabhata (2009).
Scintillula ovulina Deshayes, 1856. Reported as: Scintilla ovulina Deshayes, 1856. Andaman Sea. Reported by:
Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013), Huber (2015), BEDO (2017a).
Varotoga anomala (Deshayes, 1856). Reported as: Scintilla anomala Deshayes, 1856. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013), BEDO (2017a).
Family Lasaeidae
Borniola venusta (Lynge, 1909). Reported as: Montacuta venusta Lynge, 1909; and Pseudopythina venusta (Lynge,
1909). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009).
Borniopsis ariakensis Habe, 1959. Reported as: Pseudopythina ariakensis Habe, 1959. Gulf of Thailand. Reported
by: Robba et al. (2002), Nabhitabhata (2009).
Callomysia costata (Lynge, 1909). Reported as: Montacuta costata Lynge, 1909; and Mysella costata (Lynge,
1909). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009).
Callomysia matsuii Habe, 1951. Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009).
Curvemysella paula (A. Adams, 1856). Also reported as: Montacuta (Tellimya) paula A. Adams, 1856. Andaman
Sea. Reported by: Lynge (1909), Scott (1995), Robba et al. (2002), Nabhitabhata (2009), GBIF (2019a).
Kellia bullula (Deshayes, 1856). Gulf of Thailand. Reported by: Huber (2015).
Kellia cycladiformis (Deshayes, 1839). Reported as: Erycina cycladiformis Deshayes, 1839. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Kellia laperousii Deshayes, 1839. Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009).
Kellia porculus Pilsbry, 1904. Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009).
Kellia rosea (Lynge, 1909). Reported as: Erycina rosea. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata
(2009).
Kellia rotunda (Deshayes, 1856). Reported as: Erycina rotunda Deshayes, 1839. Gulf of Thailand. Reported by:
Lynge (1909), Nabhitabhata (2009).
32
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Kellia vitrina (Lynge, 1909). Reported as: Erycina vitrine. Gulf of Thailand. Reported by: Lynge (1909),
Nabhitabhata (2009). Reported as a taxon inquirendum by WoRMS Editorial Board (2019).
Kellya lilium Lynge, 1909. Reported as: Erycina lilium; and Mysella lilium (Lynge, 1909). Gulf of Thailand.
Reported by: Lynge (1909), Scott (1995), Nabhitabhata (2009). Reported as a taxon inquirendum by WoRMS
Editorial Board (2019).
Kellya lineata (Lynge, 1909). Reported as: Erycina lineata (Lynge, 1909); and Erycina lineate (Lynge, 1909). Gulf
of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009). Reported as a taxon
inquirendum by WoRMS Editorial Board (2019).
Koreamya arcuata (A. Adams, 1856). Reported as: Curvemysella arcuata (Adams, 1856). Gulf of Thailand.
Reported by: Robba et al. (2002).
Koreamya sibogai (Prashad, 1932). Andaman Sea. Reported by: BEDO (2017a).
Kurtiella triangularis (R. B. Watson, 1897). Reported as: Montacuta (Tellimya) triangularis A. Adams, 1856. Gulf
of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Lasaea undulata (A. A. Gould, 1861). Gulf of Thailand. Reported by: Kurozumi et al. (1989).
Montacutona japonica (Yokoyama, 1922). Reported as: Montacuta (Tellimya) japonica A. Adams, 1862. Gulf of
Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Montacutona mutsuwaensis (Yamamoto & Habe, 1959). Reported as: Mysella mutsuwaensis (Yamamoto &
Habe, 1959). Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009).
Montacutona rudis (Lynge, 1909). Reported as: Mysella rudis (Lynge, 1909). Gulf of Thailand. Reported by:
Robba et al. (2002), Nabhitabhata (2009).
Montacutona variabilis (Lynge, 1909). Reported as: Montacuta (Tellimya) variabilis Lynge, 1909; and Mysella
variabilis (Lynge, 1909). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009),
Huber (2015).
Nipponomysella subtruncata (Yokoyama, 1922). Gulf of Thailand. Reported by: Negri et al. (2014).
? Radobornia bryani (Pilsbry, 1921). Gulf of Thailand. Reported by: Gemert (2003).
Order Gastrochaenida
Superfamily Gastrochaenoidea
Family Gastrochaenidae
Cucurbitula cymbium (Spengler, 1783). Also reported as: Gastrochaena cymbium (Spengler, 1783). Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Swennen et al. (2001), ONEP (2007), Valentich-Scott &
Tongkerd (2008), Nabhitabhata (2009), Sanpanich (2011).
Dufoichaena dentifera (Dufo, 1840). Reported as: Gastrochaena dentifera Dufo, 1840. Andaman Sea. Reported
by: Scott (1995), Aungtonya et al. (1999), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), BEDO
(2017a).
Dufoichaena pexiphora (Sturany, 1899). Reported as: Gastrochaena cf. pexiphora Sturany, 1899. Gulf of Thailand,
Andaman Sea. Reported by: Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), Sanpanich (2011).
Gastrochaena carteri Nielsen, 1906. Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999),
Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Printrakoon et al.
(2016), BEDO (2017a).
Gastrochaena cuneiformis Spengler, 1783. Reported as: Gastrochaena gigantea Deshayes, 1830. Gulf of
Thailand, Andaman Sea. Reported by: Nielsen (1976a), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Aungtonya et al. (1999), Robba et al.
(2002), Taylor et al. (2007), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), Sanpanich & Duangdee
(2009), Sanpanich (2011), Hylleberg & Aungtonya (2013), Printrakoon et al. (2016), BEDO (2017a).
Lamychaena weinkauffi (Sturany, 1899). Reported as: Gastrochaena inaequistriata Jousseaume in Lamy, 1923.
Andaman Sea. Reported by: Aungtonya et al. (1999), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009),
BEDO (2017a).
Spengleria mytiloides (Lamarck, 1818). Reported as: Gastrochaena mytiloides Lamarck, 1818; and Spengleria
plicatilis (Deshayes, 1855). Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Aungtonya
et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO (2017a).
Spenglerichaena apertissima (Deshayes, 1855). Reported as: Spengleria apertissima (Deshayes, 1855).
Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001), Valentich-Scott & Tongkerd (2008),
Nabhitabhata (2009), Huber (2010), BEDO (2017a).
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The Marine and Estuarine Molluscs of Thailand
Order Lucinida
Superfamily Lucinoidea
Family Lucinidae
Austriella corrugata (Deshayes, 1843). Also reported as: Eamesiella corrugata (Deshayes, 1843); Pseudomiltha
corrugata (Deshayes, 1843); and Eamisiella philippinarum (Hanley). Gulf of Thailand, Andaman Sea. Reported
by: Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Subba Rao & Dey (2000),
Robba et al. (2002), Glover & Taylor (2016), Nabhitabhata (2009), BEDO (2017a).
Cardiolucina eucosmia (Dall, 1901). Reported as: Lucina (Phacoides) pisum Reeve, 1850. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Cardiolucina maccasari (Prashad, 1932). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001),
Meyer et al. (2008), Nabhitabhata (2009), Glover et al. (2016), Glover & Taylor (2016), BEDO (2017a).
Cardiolucina rugosa (Hedley, 1909). Reported as: Lucina (Phacoides) seminula Gould, 1861. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Cardiolucina semperiana (Issel, 1869). Also reported as: Parvilucina (Bellucina) semperiana (Issel, 1869). Gulf
of Thailand. Reported by: Robba et al. (2002), Nabhitabhata (2009), GBIF (2019b).
Cavatidens bullula (Reeve, 1850). Reported as: Anodontia bullula (Reeve, 1850); and Lucina (Thyasira) bullula
Reeve, 1850. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Robba et al. (2004), Taylor et al. (2007),
Meyer et al. (2008), Nabhitabhata (2009).
Chavania striata (Tokunaga, 1906). Gulf of Thailand. Reported by: Taylor & Glover (2001), Gemert (2003), GBIF
(2019b).
Codakia interrupta (Lamarck, 1816). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), ONEP
(2007), Nabhitabhata (2009), Duangdee et al. (2013a, 2013b), Sanpanich & Duangdee (2018), GBIF (2019a).
Codakia paytenorum (Iredale, 1937). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Kasetsart
University (2006), ONR & EPP (2006), Duangdee (2007), Nabhitabhata (2009), Duangdee et al. (2013b).
Codakia punctata (Linnaeus, 1758). Also reported as: Lucina punctata Linnaeus. Gulf of Thailand, Andaman Sea.
Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Nateewathana et al. (1981), Carpenter & Niem
(1998), Aungtonya et al. (1999), Duangdee et al. (2009), Nabhitabhata (2009), Galli (2015), BEDO (2017a).
Codakia tigerina (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b),
Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya
et al. (1999), Kasetsart University (2006), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Duangdee
et al. (2011, 2013a), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Cryptophysema vesicula (Gould, 1850). Also reported as: Loripes vesicula Gould; Anodontia vesicula (Reeve,
1850); Anodontia (Cryptophysema) vesicula (Gould, 1850); and Lucina (Thyasira) vesicula Gould, 1850. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937), Taylor & Glover (2005), Nabhitabhata
(2009), Sanpanich & Duangdee (2009), Sanpanich (2011). Reported as Anodontia pila (Reeve, 1850) by
Nabhitabhata (2009) and BEDO (2017a), but Anodontia pila was synonymised with the Atlantic Anodontia alba
Link, 1807, by Taylor & Glover (2005), and the species is actually Cryptophysema vesicula (Gould, 1850).
Ctena bella (Conrad, 1837)/Ctena divergens (Philippi, 1850). Also reported as: Epicodakia bella (Conrad,
1837); Epicodakia divergens (Philippi, 1850); and Codakia divergens Philippi, 1850. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Gemert (2003), Kasetsart University
(2006), Duangdee (2007), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2018), Sanpanich (2011), BEDO (2017a), GBIF (2019a). Note: The name Ctena bella is usually used for
Pacific shells and Ctena divergens for the Indian Ocean, but this may be a species complex (J. Taylor, pers.
comm.).
Ctena delicatula (Pilsbry, 1904). Also reported as: Epicodakia delicatula (Pilsbry, 1904). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Taylor et al. (2007), Meyer et al. (2008), Duangdee et al. (2011),
Huber (2015), Glover & Taylor (2016), Glover et al. (2016).
Discolucina virginea (Deshayes, 1832). Reported as: Lucina argentea Reeve, 1850. Gulf of Thailand. Reported
by: Nabhitabhata (2009).
Divalinga sechellensis (d’Orbigny, 1845). Also reported as: Divaricella seychellensis (d’Orbigny, 1846). Andaman
Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Divaricella ornatissima (d’Orbigny, 1846). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Easmithia desiderata (E. A. Smith, 1885). Reported as: Loripes (Loripes) desideratus (Smith, 1885). Gulf of
Thailand. Reported by: Lynge (1909), Robba et al. (2002, 2004), Nabhitabhata (2009).
Epicodakia sweeti (Hedley, 1899). Andaman Sea. Reported by: Duangdee et al. (2013b).
Euanodontia ovum (Reeve, 1850). Also reported as: Lucina (Thyasira) ovum Reeve, 1850; Anodontia ovum Reeve,
1850; and Anodontia (Euanodontia) ovum (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Lynge
34
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
(1909), Taylor & Glover (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Duangdee et al. (2011,
2013b), Sanpanich (2011).
Fimbria fimbriata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Wells (2005), Nabhitabhata (2009),
BEDO (2017a).
Fimbria soverbii (Reeve, 1842). Reported as: Fimbria sowerbyi (Reeve, 1841). Gulf of Thailand, Andaman Sea.
Reported by: Meyer et al. (2008), Nabhitabhata (2009), Sanpanich (2011).
Indoaustriella dalli (Lynge, 1909). Also reported as: Lucina (Phacoides) dalli. Gulf of Thailand. Reported by:
Lynge (1909), Nabhitabhata (2009), Duangdee et al. (2013a), Glover & Taylor (2016), Glover et al. (2016).
Indoaustriella lamprelli Glover, Taylor & Williams, 2008. Andaman Sea. Reported by: Glover et al. (2008).
Indoaustriella plicifera (A. Adams, 1855). Gulf of Thailand. Reported by: Glover et al. (2008).
Jallenia inanis (Prashad, 1932). Reported as: “Cavatidens” imajimai (Habe, 1981). Gulf of Thailand. Reported by:
Scott (1995).
Lamellolucina dentifera (Jonas, 1846). Reported as: Lucina dentifera (Jonas, 1846). Andaman Sea. Reported by:
Scott (1995), Nabhitabhata (2009), BEDO (2017a).
Lepidolucina odontotis (Salisbury, 1934). Gulf of Thailand, Andaman Sea. Reported by: Glover & Taylor (2007,
2016), Sanpanich (2011).
Lepidolucina venusta (Philippi, 1847). Reported as: Lucina venusta (Philippi, 1847); Lucina (Lucinisca) venusta
Philippi; and Miltha cf. venustra (Philippi, 1847). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889),
Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Glover & Taylor (2007), Nabhitabhata (2009), GBIF
(2019a).
Liralucina lyngei M. Huber, 2015. Reported as: Parvilucina (Parvilucina) pulchella (Lynge, 1909); and Codakia
pulchella (Lynge, 1909). Gulf of Thailand. Reported by: Lynge (1909), Scott (1995), Robba et al. (2002),
Nabhitabhata (2009), Huber (2015), Glover et al. (2016).
Liralucina sperabilis (Hedley, 1909). Reported as: Linga (Bellucina) cf. Liralucina sperabilis (Hedley, 1909). Gulf
of Thailand. Reported by: Gemert (2019).
Pegophysema bialata (Pilsbry, 1895). Reported as: Anodontia (? Pegophysema) bialata (Pilsbry, 1895). Gulf of
Thailand. Reported by: Taylor & Glover (2005).
Pegophysema philippiana (Reeve, 1850). Reported as: Anodontia philippiana (Reeve, 1850); and Anodontia
(Pegophysema) philippiana (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Taylor & Glover
(2005), Meyer et al. (2008). Reported as Anodontia edentula (Linnaeus, 1758); and Lucina (Thyasira) edentula
Linnaeus, 1758, by Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995), Scott (1995), Carpenter &
Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee
(2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Glover & Taylor (2016), and BEDO (2017a). Taylor
& Glover (2005) recommended abandoning the use of Anodontia edentula as there is no Linnean type and the
description is brief and could be any species. Most references are to Pegophysema philippiana.
Pillucina pisidium (Dunker, 1860). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Taylor & Glover (2001), Robba et al. (2002), Nabhitabhata (2009), Negri et al. (2014),
Glover et al. (2016), BEDO (2017a). Robba et al. (2004) stated that their specimens were actually Pillucina
australis Glover & Taylor, 2001, but Glover et al. (2016) refer to Pillucina pisidium.
Pillucina vietnamica Zorina, 1978. Andaman Sea. Reported by: Taylor & Glover (2001), Taylor et al. (2007),
Meyer et al. (2008), Glover & Taylor (2016), Glover et al. (2016), J. D. Taylor et al. (unpublished data).
Scabrilucina vitrea (Deshayes, 1844). Andaman Sea. Reported by: Taylor & Glover (2013), Glover et al. (2016),
GBIF (2019a).
Wallucina fijiensis (E. A. Smith, 1885). Reported as: Codakia fijiensis Smith, 1885. Gulf of Thailand. Reported by:
Lynge (1909), Taylor & Glover (2001), Nabhitabhata (2009).
Order Myida
Superfamily Dreissenoidea
Family Dreissenidae
Mytilopsis sallei [Récluz, 1849]. Also reported as: Mytilopsis adamsi Morrison, 1946. Gulf of Thailand. Reported
by: Wangkulangkul & Lheknim (2008), Wangkulangkul (2018), Sanpanich & Wells (2019). This is an invasive
species that originated either in the eastern Pacific Ocean (Mytilopsis adamsi) or Caribbean Sea (Mytilopsis
sallei) and is widespread in ports in the Indo-Pacific region. Most papers use Mytilopsis sallei for the species,
but some of the Thai literature uses Mytilopsis adamsi. The systematics of the species have not been formally
established. See Willan et al. (2000) for a discussion of the issue.
35
The Marine and Estuarine Molluscs of Thailand
Superfamily Myoidea
Family Corbulidae
Corbula archaeformis Lynge, 1909. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b),
Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO
(2017a).
Corbula erythrodon Lamarck, 1818. Also reported as: Solidocorbula erythrodon (Lamarck, 1818). Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Scott (1995), Kilburn &
Hylleberg (1998), Aungtonya et al. (1999), Kasetsart University (2000), Swennen et al. (2001), Nabhitabhata
(2009), Sanpanich & Duangdee (2009), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a).
Corbula fortisulcata (Smith, 1879). Reported as: Corbula (Corbula) fortisulcata (Smith, 1878). Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Robba et al. (2002), Taylor (2009), GBIF (2019a).
Corbula lineata Lynge, 1909. Reported as: Corbula (Caryocorbula) lineata Lynge, 1909. Gulf of Thailand. Reported
by: Lynge (1909), Bernard et al. (1993), Robba et al. (2002, 2003), Nabhitabhata (2009).
Corbula macgillivrayi Smith, 1885. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich (2011).
Corbula monilis Hinds, 1843. Also reported as: Corbula (Notocorbula) monilis Hinds, 1843. Gulf of Thailand.
Reported by: Lynge (1909), Swennen et al. (2001), Robba et al. (2002, 2003), Gemert (2003), Nabhitabhata
(2009).
Corbula ovalina Lamarck, 1818. Reported as: Corbula crassa Reeve, 1843; and Corbula (Serracorbula) crassa
Reeve, 1843. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a), Tantanasiriwong
(1979), Morris & Purchon (1981), Nateewathana et al. (1981), Aungtonya et al. (1999), Swennen et al. (2001),
Robba et al. (2002), Gemert (2003), Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a).
Corbula pallida Reeve, 1843. Reported as: Corbula (Anisocorbula) pallida L. A. Reeve, 1843. Andaman Sea.
Reported by: Huber (2010).
Corbula rotalis Hinds, 1843. Gulf of Thailand. Reported by: Suvatti (1937, 1938), Nabhitabhata (2009).
Corbula rugifera E. A. Smith, 1903. Reported as: Corbula rugifera A. Adams, 1903. Gulf of Thailand. Reported by:
Lynge (1909), Nabhitabhata (2009).
Corbula scaphoides Hinds, 1843. Also reported as: Anisocorbula scaphoides (Hinds, 1843). Gulf of Thailand,
Andaman Sea. Reported by: Robba et al. (2002), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich
(2011), BEDO (2017a).
Corbula sinensis F. R. Bernard, Cai & B. Morton, 1993. Gulf of Thailand. Reported by: Taylor et al. (2007).
Corbula smithiana Brazier, 1879. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
(2011).
Corbula solidula Hinds, 1843. Also reported as: Corbula (Anisocorbula) solidula Hinds, 1843. Gulf of Thailand.
Reported by: Nabhitabhata (2009), Negri et al. (2014).
Corbula subquadrata Melvill & Standen, 1907. Reported as: Corbula subquadrata Melvill, 1906. Gulf of Thailand.
Reported by: Lynge (1909).
Corbula sulcata Lamarck, 1801. Gulf of Thailand. Reported by: Lynge (1909).
Corbula taitensis Lamarck, 1818. Also reported as: Corbula modesta Hinds, 1843; and Corbula taheitensis
Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nielsen
(1976a), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Aungtonya et al.
(1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), J. D. Taylor et al. (unpublished data).
Corbula tunicata Hinds, 1843. Reported as: Corbula (Aloidis) tunicata Hinds, 1843. Gulf of Thailand. Reported
by: Lynge (1909).
Corbula cf. venusta (Gould, 1861). Reported as: Anisocorbula cf. venusta (Gould, 1861). Gulf of Thailand.
Reported by: Scott (1995).
Potamocorbula fasciata (Reeve, 1843). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata
(2009), Huber (2010), GBIF (2019b).
Potamocorbula laevis (Hinds, 1843). Gulf of Thailand. Reported by: Negri et al. (2014).
Family Myidae
Cryptomya busoensis Yokoyama, 1922. Gulf of Thailand. Reported by: Gemert (2003).
Cryptomya elliptica (A. Adams, 1851). Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Sphenia perversa Blandford, 1867. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Taylor et
al. (2007), Nabhitabhata (2009), Negri et al. (2014), Oliver et al. (2018).
Tugonia decurtata (A. Adams, 1851). Andaman Sea. Reported by: Nabhitabhata (2009).
36
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Superfamily Pholadoidea
Family Pholadidae
Aspidopholas ovum (W. Wood, 1828). Reported as: Aspidopholas ovata (G. B. Sowerby II, 1849). Andaman Sea.
Reported by: Valentich-Scott & Tongkerd (2008).
Aspidopholas tubigera (Valenciennes, 1846). Reported as: Aspidopholas obtecta (G. B. Sowerby II, 1849).
Andaman Sea. Reported by: Valentich-Scott & Tongkerd (2008).
Barnea dilatata (Souleyet, 1843). Reported as: Barnea (Anchomasa) dilatata (Souleyet, 1843). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999),
Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009), Haga (2011), GBIF (2019a).
Barnea manilensis (Philippi, 1847). Also reported as: Barnea erythraea (Gary, 1851) and Barnea (Anchomasa)
manilensis (Philippi, 1847). Gulf of Thailand. Reported by: Swennen et al. (2001), Robba et al. (2002),
Nabhitabhata (2009).
Diplothyra curta (G. B. Sowerby I, 1834). Reported as: Pholas (Martesia) grayana Leach. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Jouannetia cumingii (G. B. Sowerby II, 1849). Reported as: Jouanettia cumingii (Sowerby, 1849). Gulf of
Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001), Nabhitabhata (2009),
Duangdee et al. (2011), Hylleberg & Aungtonya (2013), BEDO (2017a).
Jouannetia globulosa (Quoy & Gaimard, 1835). Also reported as: Jouannetia globosa (Quoy & Gaimard).
Andaman Sea. Reported by: Nielsen (1976a), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana
et al. (1981), Aungtonya et al. (1999), ONEP (2007), Nabhitabhata (2009), BEDO (2017a).
Lignopholas rivicola (G. B. Sowerby II, 1849). Reported as: Martesia rivicola Sowerby; Pholas (Martesia) rivicola
Sowerby, 1848; Pholas rivicola Sowerby, 1848; and Mertasia rivicola Sowerby. Gulf of Thailand, Andaman Sea.
Reported by: Morlet (1889), Suvatti (1938), Nabhitabhata (2009).
Martesia multistriata (G. B. Sowerby II, 1849). Reported as: Aspidopholas multistriata (G. B. Sowerby II, 1849);
and Pholas (Martesia) multistriata Sowerby, 1849. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Suvatti (1938), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), Duangdee et al. (2011).
Martesia striata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909),
Suvatti (1938), Yoosukh & Jitkaew (1997), Carpenter & Niem (1998), Swennen et al. (2001), Robba et al.
(2002), Nabhitabhata (2009), GBIF (2019a, 2019b).
Parapholas quadrizonata (Spengler, 1797). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009),
Sanpanich (2011), BEDO (2017a).
Pholadidea fauroti Jousseaume, 1888. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich (2011).
Pholas orientalis Gmelin, 1791. Also reported as: Phragmopholas orientalis Gmelin, 1791; Barnea (Monothyra)
orientalis Gmelin, 1791; and Pholas siamensis (Spengler, 1788). Gulf of Thailand, Andaman Sea. Reported by:
Morlet (1889), Lynge (1909), Suvatti (1937, 1938), Chaitiamwong et al. (1971), Amornjaruchit (1988), Bernard
et al. (1993), Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Knudsen & Jensen
(2001), Swennen et al. (2001), Robba et al. (2002), Kasetsart University (2006), ONEP (2007), Nabhitabhata
(2009), BEDO (2017a), GBIF (2019b).
Family Teredinidae
Bactronophorus thoracites (Gould, 1856). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1995),
Yoosukh & Jitkaew (1997), Carpenter & Niem (1998), Aungtonya et al. (1999), Kasetsart University (2000),
Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Bankia gracilis Moll, 1935. Gulf of Thailand. Reported by: Yoosukh & Jitkaew (1997), Nabhitabhata (2009),
Huber (2015).
Dicyathifer mannii (Wright, 1866). Also reported as: Teredo manni Wright, 1866. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Yoosukh & Jitkaew (1997), Kasetsart University (2000), Nabhitabhata (2009).
Lyrodus pedicellatus (Quatrefages, 1849). Also reported as: Teredo (Teredo) siamensis Bartsch. Gulf of Thailand,
Andaman Sea. Reported by: Suvatti (1938), Carpenter & Niem (1998), Nabhitabhata (2009), GBIF (2019b).
Nausitora dunlopei Wright, 1864. Reported as: Bankia (Bankia) smithi Bartsch. Gulf of Thailand. Reported by:
Suvatti (1938), Nabhitabhata (2009).
Nausitora hedleyi Schepman, 1919. Gulf of Thailand. Reported by: Yoosukh & Jitkaew (1997), Nabhitabhata
(2009), Huber (2015).
Spathoteredo obtusa (Sivickis, 1928). Gulf of Thailand. Reported by: Yoosukh & Jitkaew (1997), Nabhitabhata
(2009).
Teredo furcifera von Martens, 1894. Gulf of Thailand. Reported by: Yoosukh & Jitkaew (1997), Swennen et al.
(2001), Nabhitabhata (2009), GBIF (2019b).
37
The Marine and Estuarine Molluscs of Thailand
Teredo johnsoni Clapp, 1924. Gulf of Thailand. Reported by: GBIF (2019b).
Teredora princesae (Sivickis, 1928). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Order Venerida
Superfamily Articoidea
Family Trapezidae
Coralliophaga coralliophaga (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009),
Sanpanich (2011), Printrakoon et al. (2016), BEDO (2017a).
Fluviolanatus subtortus (Dunker, 1857). Gulf of Thailand. Reported by: Gemert (2003).
Glossocardia stoliczkana Prashad, 1932. Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Neotrapezium sublaevigatum (Lamarck, 1819). Reported as: Trapezium sublaevigatum (Lamarck, 1819). Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Robba et al. (2004),
Taylor et al. (2007), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished
data).
Trapezium bicarinatum (Schumacher, 1817). Gulf of Thailand, Andaman Sea. Reported by: Morris & Purchon
(1981), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO
(2017a), GBIF (2019a).
Trapezium gilvum (Martens, 1872). Reported as: Trapezium angulatum (Lamarck, 1819); and Trapezium sowerbyi
(Hidalgo, 1903). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich
(2011).
Trapezium oblongum (Linnaeus, 1758). Gulf of Thailand. Reported by: Aungtonya et al. (1999), Nabhitabhata
(2009).
Superfamily Chamoidea
Family Chamidae
Amphichama scutulina (Poutiers, 1981). Reported as: Pseudochama scutulina Poutiers, 1981. Gulf of Thailand.
Reported by: Robba et al. (2002), Nabhitabhata (2009).
Chama asperella Lamarck, 1819. Also reported as: Chama jukesii Reeve, 1847; and Chama fibula Reeve, 1846.
Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Nielsen (1976b), Tantanasiriwong (1979),
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Robba et al. (2002), Gemert (2003),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a), GBIF (2019a,
2019b).
Chama aspersa Reeve, 1846. Also reported as: Chama aspera Reeve. Andaman Sea. Reported by: Nielsen (1976a),
Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Swennen et
al. (2001), Taylor et al. (2007), Nabhitabhata (2009), J. D. Taylor et al. (unpublished data). Reported as a nomen
dubium by WoRMS Editorial Board (2019).
Chama brassica Reeve, 1847. Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Swennen et al. (2001), Gemert (2003), Duangdee
(2004), Duangdee et al. (2007, 2013a), Nabhitabhata (2009), Sanpanich (2011), Printrakoon et al. (2016), BEDO
(2017a).
Chama croceata Lamarck, 1819. Also reported as: Chama plinthota Col, 1927; and Chama savigni Lamy, 1921.
Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Aungtonya et al. (1999), Kas etsart
University (2000), Gemert (2003), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Duangdee et al. (2011,
2013b), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Chama dunkeri Lischke, 1870. Gulf of Thailand. Reported by: Kurozumi et al. (1989).
Chama fragum Reeve, 1847. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009). Reported as a
nomen dubium by WoRMS Editorial Board (2019).
Chama japonica Lamarck, 1819. Gulf of Thailand. Reported by: Kurozumi et al. (1989), Kasetsart University (2006),
Duangdee (2007), Nabhitabhata (2009). Reported as a nomen dubium by WoRMS Editorial Board (2019).
Chama lazarus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Duangdee (2004, 2007), Kasetsart
University (2006), Duangdee et al. (2009, 2011), Nabhitabhata (2009), GBIF (2019a).
Chama limbula Lamarck, 1819. Reported as: Chama iostoma Conarad, 1837. Gulf of Thailand, Andaman Sea.
Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a).
38
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Chama macerophylla Gmelin, 1791. Reported as: Chama imbricata Lamarck, 1801. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a).
Chama pacifica Broderip, 1834. Also reported as: Chama reflexa Reeve, 1846; Chama multisquamosa Reeve,
1846; and Chama divaricata Reeve, 1846. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti
(1937, 1938), Nielsen (1976a), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981),
Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al.
(1999), Kasetsart University (2000, 2006), Gemert (2003), Duangdee (2004, 2007), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011, 2013a), ONEP (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO
(2017a), Sanpanich & Duangdee (2018), GBIF (2019a).
Chama pellucida Broderip, 1835. Andaman Sea. Reported by: Bussarawit (1995), Nabhitabhata (2009).
Chama pulchella Reeve, 1846. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Aungtonya et al.
(1999), Nabhitabhata (2009), BEDO (2017a).
Chama ruderalis Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
(2011).
Pseudochama cristella (Lamarck, 1819). Reported as: Chama cristella Lamarck, 1819. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Superfamily Cyrenoidea
Family Cyrenidae
Batissa insignis Deshayes, 1854. No Thai locality provided. Reported by: Huber (2015).
Batissa violacea (Lamarck, 1806). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Aungtonya et al. (1999), Nabhitabhata (2009).
Geloina bengalensis (Lamarck, 1818). Reported as: Cyrena sumatrensis G. B. Sowerby I, 1822; Polymesoda
(Geloina) bengalensis Lamarck, 1818; and Polymesoda (Geloina) galatheae (Mörch, 1850). Gulf of Thailand,
Andaman Sea. Reported by: Morlet (1889), Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al.
(1999), Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Geloina coaxans (Gmelin, 1791). Reported as: Polymesoda proxima (Gmelin, 1791); and Geloina zeylanica
Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999), Printrakoon et al. (2008),
Nabhitabhata (2009).
Geloina expansa (Mousson, 1849). Reported as: Geloina erosa (Solander, 1786); and Geloina (Polymesoda) erosa
(Solander, 1786). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter & Niem
(1998), Swennen et al. (2001), Kasetsart University (2006), Duangdee (2007), Printrakoon et al. (2008),
Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Geloina triangularis (Metcalfe, 1852). Gulf of Thailand. Reported by: Huber (2015).
Family Glauconomidae
Glauconome chinensis Gray, 1828. Gulf of Thailand. Reported by: Robba et al. (2002), Gemert (2003), Nabhitabhata
(2009), Negri et al. (2014).
Glauconome radiata Reeve, 1844. Reported as: Glauconome plankta (Iredale, 1936). Gulf of Thailand, Andaman
Sea. Reported by: Printrakoon et al. (2008), Nabhitabhata (2009), Sanpanich (2011).
Glauconome straminea Reeve, 1844. Gulf of Thailand. Reported by: GBIF (2019b).
Glauconome virens (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Huber
(2010), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Superfamily Glossoidea
Family Glossidae
Meiocardia cumingi (A. Adams, 1864). Reported as: Meiocardia globosa Matsukuma & Habe, 1995. Andaman
Sea. Reported by: Matsukuma & Habe (1995), Nabhitabhata (2009), Huber (2010), BEDO (2017a).
Meiocardia hawaiana Dall, Bartsch & Rehder, 1938. Andaman Sea. Reported by: Matsukuma & Habe (1995).
Meiocardia moltkiana (Gmelin, 1791). Reported as: Meiocardia moltkiana (Spengler, 1783). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Matsukuma & Habe (1995),
Aungtonya & Hylleberg (1998), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Meiocardia samarangiae Bernard, Cai & Morton, 1993. Also reported as: Meiocardia tetragona (Adams &
Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995),
Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
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The Marine and Estuarine Molluscs of Thailand
Meiocardia sangiuneomaculata (Dunker, 1882). Gulf of Thailand, Andaman Sea. Reported by: Kilburn &
Hylleberg (1998), Nabhitabhata (2009).
Meiocardia vulgaris (Reeve, 1845). Andaman Sea. Reported by: Matsukuma & Habe (1995), Nabhitabhata (2009),
BEDO (2017a), GBIF (2019a).
Family Kelliellidae
Alveinus ojianus (Yokoyama, 1927). Gulf of Thailand. Reported by: Robba et al. (2002, 2003), Nabhitabhata
(2009), Negri et al. (2014).
Superfamily Hemidonacoidea
Family Hemidonacidae
Hemidonax donaciformis (Bruguière, 1789). Andaman Sea. Reported by: GBIF (2019a).
Superfamily Mactroidea
Family Anatinellidae
Raeta pellicula (Reeve, 1854). Reported as: Raeta anatinoides Reeve, 1854. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Nabhitabhata (2009), Sanpanich (2011).
Raeta pulchella (Adams & Reeve, 1850). Also reported as: Raeta (Raetellops) pulchella (Adams & Reeve, 1850).
Gulf of Thailand. Reported by: Lynge (1909), Swennen et al. (2001), Robba et al. (2002), Kasetsart University
(2006), Nabhitabhata (2009), Negri et al. (2014).
Family Cardiliidae
Cardilia semisulcata (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017a).
Family Mactridae
Heterocardia gibbosula Deshayes, 1855. Reported as: Heterocardia gibbosula Philippi, 1871. Gulf of Thailand.
Reported by: Kilburn & Hylleberg (1998), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009),
Premcharoen et al. (2016).
Lutraria complanata (Gmelin, 1791). Reported as: Lutraria (Lutropbora) complanata (Gmelin, 1791). Gulf of
Thailand. Reported by: Suvatti (1937, 1938), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009).
Lutraria curta Reeve, 1854. Reported as: Lutraria (Psammophila) curta L. A. Reeve, 1854. No Thai locality
provided. Reported by: Huber (2010).
Lutraria impar Deshayes, 1854. Also reported as: Lutraria impar Deshayes, 1845. Gulf of Thailand, Andaman
Sea. Reported by: Bernard et al. (1993), Vongpanich (2000), Swennen et al. (2001), Nabhitabhata (2009).
Lutraria maxima Jonas, 1844. Gulf of Thailand. Reported by: Nateewathana et al. (1981), Vongpanich (2000),
Nabhitabhata (2009), Duangdee et al. (2011), BEDO (2017a).
Lutraria rhynchaena Jonas, 1844. Reported as: Lutraria australis (Reeve, 1854); Lutraria philippinarum Reeve,
1854; Lutraria arcuata Deshayes; and Lutraria curta Deshayes, 1854. Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Kurozumi et al. (1989), Sanpanich (1998, 2011), Vongpanich (2000), Gemert (2003), Wells
(2005), Kasetsart University (2006), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2018), BEDO (2017a).
Lutraria sieboldii Reeve, 1854. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Mactra abbreviata Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
(2011).
Mactra achatina Holten, 1802. Also reported as: Mactra ornata Gray, 1837; and Mactra achatina Dillwyn, 1817.
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995),
Carpenter & Niem (1998), Aungtonya et al. (1999), Vongpanich (2000), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a).
Mactra cf. alta Deshayes, 1855. Gulf of Thailand. Reported by: Gemert (2003).
Mactra antiquata Spengler, 1802. Also reported as: ? Coelomactra antiquata (Spengler, 1802). Gulf of Thailand,
Andaman Sea. Reported by: Suvatti (1937, 1938), Aungtonya et al. (1999), Gemert (2003), Nabhitabhata (2009),
BEDO (2017a).
Mactra artensis Montrouzier in Fischer, 1850. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003),
Nabhitabhata (2009), Sanpanich (2011).
40
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Mactra chinensis Philippi, 1846. Reported as: Mactra sulcataria Reeve, 1854. Gulf of Thailand, Andaman Sea.
Reported by: Vongpanich (2000), Duangdee et al. (2007), GBIF (2019a).
Mactra crossei (Dunker, 1877). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Mactra cumingii (Reeve, 1854). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), ONEP
(2007), Printrakoon et al. (2008), Nabhitabhata (2009).
Mactra cuneata Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bussarawit (1995),
Carpenter & Niem (1998), Vongpanich (2000), Gemert (2003), Kasetsart University (2006), ONR & EPP (2006),
Duangdee (2007), Duangdee et al. (2007), Nabhitabhata (2009), Sanpanich (2011), GBIF (2019a, 2019b).
Mactra cygnus Gmelin, 1791. Also reported as: Mactra incongrua Deshayes, 1854; and Mactra purpurea Spengler,
1802. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999),
Vongpanich (2000), Nabhitabhata (2009), Galli (2015), BEDO (2017a).
Mactra dissimilis Reeve, 1854. Reported as: Mactra (Mactra) dissimilis Reeve, 1854. Gulf of Thailand, Andaman
Sea. Reported by: Suvatti (1937, 1938), Nabhitabhata (2009), Sanpanich (2011), GBIF (2019a).
Mactra grandis Gmelin, 1791. Reported as: Mactra mera Reeve, 1854. Gulf of Thailand, Andaman Sea. Reported
by: Kurozumi et al. (1989), Carpenter & Niem (1998), Sanpanich (1998), Vongpanich (2000), Duangdee (2004),
Kasetsart University (2006), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Mactra incarnata Reeve, 1854. Also reported as: Mactra (Mactra) incarnata L. A. Reeve, 1854. Gulf of Thailand,
Andaman Sea. Reported by: Kasetsart University (2006), Nabhitabhata (2009), Huber (2010), GBIF (2019a).
Mactra iridescens Kuroda & Habe in Habe, 1958. Andaman Sea. Reported by: García & Oliver (2008).
Mactra lilacea Lamarck, 1818. Also reported as: Mactra (Mactra) lilacea Lamarck, 1818. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), Taylor
(2009), Huber (2010), BEDO (2017a), GBIF (2019a).
Mactra luzonica Reeve, 1854. Reported as: Mactra luzonica Deshayes, 1854; and Mactra (Mactra) luzonica. Gulf
of Thailand, Andaman Sea. Reported by: Lynge (1909), Vongpanich (2000), Swennen et al. (2001), Robba et al.
(2002, 2003), Nabhitabhata (2009), Negri et al. (2014), BEDO (2017a), GBIF (2019a).
Mactra maculata Gmelin, 1791. Also reported as: Mactra reevei Deshayes, 1853. Gulf of Thailand, Andaman Sea.
Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Carpenter & Niem (1998), Kasetsart University
(2006), ONR & EPP (2006), Nabhitabhata (2009).
Mactra opposita Reeve, 1854. Reported as: Mactra (Mactra) opposita L. A. Reeve, 1854. No Thai locality provided.
Reported by: Huber (2010).
Mactra pusilla (A. Adams, 1855). Also reported as: Mactra (Nanomactra) pusilla (A. Adams, 1855). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Mactra sericea Reeve, 1854. Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Sanpanich (2011).
Mactra violacea Gmelin, 1791. Also reported as: Mactra (Coelomactra) violacea “Chemnitz” Gmelin, 1791. Gulf
of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Kilburn & Hylleberg (1998), Aungtonya
et al. (1999), Vongpanich (2000), Robba et al. (2002), Duangdee (2007), Duangdee et al. (2009), Nabhitabhata
(2009), Huber (2010), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Mactrella striatula (Linnaeus, 1767). Reported as: Mactrinella striatula (Linnaeus, 1767). Andaman Sea. Reported
by: BEDO (2017a).
Mactrinula dolabrata Reeve, 1854. Reported as: Mactra dolabrata Reeve, 1854. Gulf of Thailand, Andaman Sea.
Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Vongpanich (2000), Swennen et al. (2001),
Robba et al. (2004), Duangdee et al. (2007), Nabhitabhata (2009).
Mactrinula plicataria (Linnaeus, 1767). Also reported as: Meropesta plicataria; and Harvella plicataria (Linnaeus,
1767). Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999), Vongpanich (2000), Nabhitabhata
(2009), BEDO (2017a), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Mactrinula reevesii (Gray, 1837). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Vongpanich (2000),
Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017a).
Mactrotoma angulifera (Reeve, 1854). Also reported as: Mactra angulifera Deshayes, 1854. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al.
(1981), Aungtonya et al. (1999), Swennen et al. (2001), Kasetsart University (2006), BEDO (2017a), Sanpanich
& Duangdee (2018).
Mactrotoma antecedens Iredale, 1930. Reported as: Mactra antecedens Iredale, 1930. Gulf of Thailand, Andaman
Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011).
Mactrotoma ovalina (Lamarck, 1818). Reported as: Mactra ovalina (Lamarck, 1818). Andaman Sea. Reported
by: Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Meropesta capillacea (Reeve, 1854). Reported as: Standella capillacea (Reeve, 1854). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009).
Meropesta nicobarica (Gmelin, 1791). Also reported as: Eastonia (Merope) nicobarica Gmelin, 1782; and Standella
(Eastonia) nicobarica Gmelin, 1790. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1938),
Kurozumi et al. (1989), Sanpanich (1998, 2011), Aungtonya et al. (1999), Vongpanich (2000), Swennen et al.
(2001), Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009), BEDO (2017a), GBIF (2019a).
Meropesta sinojaponica Zhuang, 1983. Andaman Sea. Reported by: Aungtonya et al. (1999).
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The Marine and Estuarine Molluscs of Thailand
Oxyperas coppingeri (E. A. Smith, 1884). Also reported as: Spisula coppingeri (Smith, 1884). Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata (2009), Sanpanich (2011).
Oxyperas lentiginosum (Gould, 1852). Andaman Sea. Reported by: Vongpanich (2000), Galli (2015), GBIF (2019a).
Standella pellucida (Gmelin, 1791). Reported as: Mactra pellucida Gmelin, 1791; and Meropesta pellucida (Gmelin,
1791). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Carpenter & Niem (1998), Vongpanich
(2000), Swennen et al. (2001), Robba et al. (2002), Meyer et al. (2008), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), GBIF (2019a, 2019b).
Tresus capax (Gould, 1850). Reported as: Lutraria maxima Jonas, 1844. Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Aungtonya et al. (1999).
Family Mesodesmatidae
Atactodea striata (Gmelin, 1791). Also reported as: Atactodea glabrata (Gmelin, 1791); Atactodea striata form
glabrata (Gmelin, 1791); Mesodesma striata (Gmelin, 1791); Mesodesma (Atactodea) glabrata Lamarck, 1805;
and Paphies striata (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937,
1938), Nielsen(1976a), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981),
Kurozumi et al. (1989), Bernard et al. (1993), Bussarawit (1995), Carpenter & Niem (1998), Kilburn &
Hylleberg (1998), Aungtonya et al. (1999), Gemert (2003), Duangdee (2004, 2007), Kasetsart University
(2006), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Sanpanich
(2011), BEDO (2017a), GBIF (2019a, 2019b).
Coecella horsfieldii Gray, 1853. Also reported as: Caecella horsfieldii Gray, 1853; Coecella cumingiana Deshayes,
1854; and Coecella transversalis Deshayes, 1854. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Suvatti (1937, 1938), Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya et al.
(1999), Gemert (2003), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Davila plana (Hanley, 1843). Reported as: Davilla crassula Deshayes, 1854. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Gemert (2003), Nabhitabhata (2009), GBIF (2019a).
Donacilla picta Dunker, 1877. Gulf of Thailand. Reported by: Gemert (2003).
Superfamily Ungulinoidea
Family Ungulinidae
Cycladicama luciniformis Valenciennes in Rousseau, 1854. Gulf of Thailand. Reported by: Swennen et al. (2001),
Nabhitabhata (2009). Reported by WoRMS Editorial Board (2019) as a nomen dubium.
Diplodonta amboinensis Smith, 1885. Also reported as: Phlyctiderma amboinensis (Smith, 1885). Gulf of Thailand.
Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009).
Diplodonta subrotunda Issel, 1869. Reported as: Diplodonta subrotundata Issel, 1869. Gulf of Thailand, Andaman
Sea. Reported by: Sanpanich & Duangdee (2009), Sanpanich (2011).
Diplodonta subrugosa Dunker, 1849. Andaman Sea. Reported by: Huber (2015).
Felaniella cuneata (Spengler, 1798). Andaman Sea. Reported by: Huber (2015).
Joannisiella cumingii (Hanley, 1846). Reported as: Cyclidacama cumingii (Hanley, 1846). Gulf of Thailand.
Reported by: Meyer et al. (2008).
Joannisiella oblonga (Hanley, 1846). Reported as: Cycladicama oblonga (Hanley, 1856); and Diplodonta
(Joannisilella) oblonga Hanley, 1856. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002) ,
Nabhitabhata (2009), GBIF (2019a).
Joannisiella philippinarum (Deshayes, 1855). Also reported as: Cycladicama subquadrata (Hedley, 1915).
Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Neodiplodonta conspicua (E. A. Smith, 1885). Reported as: Felaniella (Zemysia) conspicua (Smith, 1885); and
Diplodonta conspicua Smith, 1885. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata
(2009), Negri et al. (2014).
Neodiplodonta genethlia (Melvill, 1898). Reported as: Diplodonta genethlia (Melvill, 1898). Andaman Sea.
Reported by: Gemert (2003), Nabhitabhata (2009).
Phlyctiderma japonicum (Pilsbry, 1895). Gulf of Thailand. Reported by: Robba et al. (2002), Nabhitabhata
(2009).
Transkeia ethima (Melvill & Standen, 1899). Reported as: Diplodonta ethima Melvill & Standen, 1899. Gulf of
Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Transkeia globosa (Forsskål in Niebuhr, 1775). Reported as: Diplodonta globosa (Forskål, 1775). Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata
(2009), BEDO (2017a).
Zemysina semiasperoides (Nomura, 1932). Reported as: Diplodonta semiasperoides Nomura, 1932. Gulf of
Thailand. Reported by: Duangdee et al. (2011).
42
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Superfamily Veneroidea
Family Veneridae
Anomalodiscus squamosus (Linnaeus, 1758). Reported as: Anomalocardia squamosa (Linnaeus, 1758);
Anomalocardia (Anomalodiscus) squamosa (Linnaeus, 1758); Venus squamosa Linnaeus, 1758; and
Cryptogramma squamosa Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge
(1909), Suvatti (1938), Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya
et al. (1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee et al. (2007), ONEP (2007),
Meyer et al. (2008), Printrakoon et al. (2008), Sartori et al. (2008), Nabhitabhata (2009), BEDO (2017a), GBIF
(2019b).
Antigona chemnitzii (Hanley, 1845). Also reported as: Periglypta chemnitzii (Hanley, 1845); and Proxichione
chemnitzii (Hanley, 1845). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Aungtonya
et al. (1999), Swennen et al. (2001), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Antigona lacerata (Hanley, 1845). Gulf of Thailand. Reported by: Huber (2010).
Antigona lamellaris Schumacher, 1817. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Aungtonya & Hylleberg (1998), Sanpanich (1998, 2011), Swennen et al. (2001),
Gemert (2003), Duangdee (2004), ONEP (2007), Sartori et al. (2008), Nabhitabhata (2009), Duangdee et al.
(2011), BEDO (2017a), GBIF (2019a, 2019b).
Antigona magnifica (Hanley, 1845). Reported as: Periglypta magnifica (Hanley). Gulf of Thailand. Reported by:
Kurozumi et al. (1989).
Antigona persimilis (Iredale, 1930). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009,
2018), Duangdee et al. (2013a).
Antigona resticulata (G. B. Sowerby, 1853). Andaman Sea. Reported by: Wells (2005).
Aphrodora kurodai (Matsubara, 2007). Reported as: Pitar japonicus Kuroda & Kawamoto, 1956. Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Sanpanich (2011).
Aphrodora (s.l.) sulcata (Zorina, 1978). No Thai locality provided. Reported by: Huber (2010).
Aphrodora yerburyi (E. A. Smith, 1891). Reported as: Pitar yerburyi (E. A. Smith, 1891); and Cytherea (Caryatis)
yerburyi Smith, 1891. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Sartori et al. (2008),
Nabhitabhata (2009).
Callista chinensis (Holten, 1802). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Wells (2005), Kasetsart University (2006), Nabhitabhata (2009), BEDO (2017a).
Callista chione (Linnaeus, 1758). Andaman Sea. Reported by: Nateewathana et al. (1981).
Callista erycina (Linnaeus, 1758). Also reported as: Callista lilacina (Lamarck, 1818); and Costacallista erycina
(Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (1998, 2011), Sartori et al. (2008),
Nabhitabhata (2009), BEDO (2017a).
Callista hagenowi (Dunker, 1849). Reported as: Amiantis hagenowi (Dunker, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009),
Huber (2010).
Callista impar (Lamarck, 1818). Gulf of Thailand. Reported by: Sanpanich (2011).
Callista kingii (Gray in King, 1827). Gulf of Thailand. Reported by: Sanpanich (2011).
Callista pilsbryi Habe, 1960. Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009).
Callocardia guttata A. Adams, 1864. Gulf of Thailand, Andaman Sea. Reported by: Sartori et al. (2008),
Nabhitabhata (2009), BEDO (2017a).
Circe intermedia Reeve, 1863. Gulf of Thailand. Reported by: GBIF (2019b).
Circe lirata (Römer, 1860). Gulf of Thailand, Andaman Sea. Also reported as: Gafrarium liratum (Römer, 1869).
Reported by: Lynge (1909), Bernard et al. (1993), Swennen et al. (2001), Sartori et al. (2008), Nabhitabhata
(2009).
Circe rivularis (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Sartori et al.
(2008).
Circe scripta (Linnaeus, 1758). Reported as: Circe (Circe) scripta (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Nateewathana et al. (1981), Aungtonya & Hylleberg
(1998), Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al. (1999), Gemert (2003), Kasetsart
University (2006), Duangdee et al. (2007, 2009), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Clementia papyracea (Gmelin, 1791). Also reported as: Clementia vatheleti Mabille, 1910. Gulf of Thailand,
Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001), Nabhitabhata (2009), Sanpanich
(2011), BEDO (2017a).
Costellipitar manillae (G. B. Sowerby II, 1851). Also reported as: Pitar (Costellipitar) manillae (Sowerby, 1851);
and Pitar manillae (Sowerby, 1851). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002, 2003),
Sartori et al. (2008), Nabhitabhata (2009), Negri et al. (2014).
43
The Marine and Estuarine Molluscs of Thailand
Cyclina sinensis (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Cytherea isocardia Sowerby, 1888. Reported as: Cytherea (Caryatis) isocardia Sowerby, 1888. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Dorisca amica (E. A. Smith, 1885). Reported as: Dorisca nana (Melvill, 1898); Dorisca cf. nana (Melvill, 1898);
and Circe nana Melvill, 1898. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993),
Scott (1995), Sartori et al. (2008), Nabhitabhata (2009).
Dorisca melvilli (Lynge, 1909). Reported as: Gafrarium melvilli (Lynge, 1909); Circe melvilli; and Circe melvil. Gulf
of Thailand, Andaman Sea. Reported by: Lynge (1909), Robba et al. (2002), Sartori et al. (2008), Nabhitabhata
(2009), Negri et al. (2014).
Dosinia alta (Dunker, 1849). Gulf of Thailand. Reported by: GBIF (2019b).
Dosinia amphidesmoides (Reeve, 1850). Gulf of Thailand. Reported by: Duangdee et al. (2011).
Dosinia angulosa (Philippi, 1847). Also reported as: Dosinella angulosa (Philippi, 1847); Phacosoma angulosa
(Philippi, 1847); and Dosinia hanleyana H. & A. Adams, 1855. Gulf of Thailand, Andaman Sea. Reported by:
Lynge (1909), Suvatti (1937, 1938), Nateewathana et al. (1981), Bernard et al. (1993), Aungtonya et al. (1999),
Sartori et al. (2008), Nabhitabhata (2009), BEDO (2017a).
Dosinia areolata Römer, 1870. Reported as: Dosinia bruguieri (Gray, 1838). Gulf of Thailand. Reported by:
Duangdee et al. (2011), Sanpanich (2011).
Dosinia biscocta (Reeve, 1850). Andaman Sea. Reported by: Amornjaruchit (1988).
Dosinia cf. caelata (Reeve, 1850). Andaman Sea. Reported by: Nabhitabhata (2009).
Dosinia cingulifera Römer, 1862. Reported as: Dosinia (Bonartemis) cingulifera E. Römer, 1862. No Thai locality
provided. Reported by: Huber (2010).
Dosinia circularis Römer, 1862. Reported as: Dosinia (Asa) circularis E. Römer, 1862. Andaman Sea. Reported
by: Huber (2010).
Dosinia contusa Reeve, 1850. Gulf of Thailand. Reported by: Morlet (1889), Lynge (1909), Suvatti (1938),
Nabhitabhata (2009).
Dosinia cretacea (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909),
Suvatti (1938), Swennen et al. (2001), Robba et al. (2002), ONEP (2007), Sartori et al. (2008), Nabhitabhata
(2009), BEDO (2017a), GBIF (2019b).
Dosinia crocea Deshayes, 1853. Also reported as: Dosinia circinaria Deshayes, 1853. Gulf of Thailand, Andaman
Sea. Reported by: Kasetsart University (2006), Duangdee (2007), Meyer et al. (2008), Sartori et al. (2008),
Nabhitabhata (2009), Sanpanich (2011).
Dosinia dautzenbergi Fischer-Piette & Delmas, 1967. Also reported as: Asa dautzenbergi Fischer-Piette &
Delmas, 1967. Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002), Sartori et al. (2008),
Nabhitabhata (2009), Sanpanich (2011).
Dosinia dilecta A. Adams, 1855. Also reported as: Dosinia (Dosinella) dilecta A. Adams, 1856. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Swennen et al. (2001), Robba et al. (2002), Sartori et al. (2008),
Nabhitabhata (2009), Huber (2010), GBIF (2019b).
Dosinia erythraea Römer, 1860. Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999),
Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), Duangdee et al. (2013b).
Dosinia exasperata (Philippi, 1847). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Nateewathana (1995), Sartori et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
(2011), BEDO (2017a), GBIF (2019b).
Dosinia fibula (Reeve, 1850). Andaman Sea. Reported by: J. D. Taylor et al. (unpublished data).
Dosinia funiculata Römer, 1862. Reported as: Dosinia (Dosinella) funiculata E. Römer, 1862. No Thai locality
provided. Reported by: Huber (2010).
Dosinia glauca (Reeve, 1850). Also reported as: Dosinia (Asa) glauca (L. A. Reeve, 1850); and Dosinia brevilunata
Fischer-Piette & Delmas, 1967. Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Sartori et al.
(2008), Nabhitabhata (2009), Huber (2010), GBIF (2019a).
Dosinia histrio (Gmelin, 1790). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Dosinia juvenilis (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Kasetsart
University (2006), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich (2011), GBIF (2019b).
Dosinia labiosa Römer, 1862. Reported as: Dosinia pubescens Philippi, 1847. Gulf of Thailand. Reported by: Lynge
(1909).
Dosinia laminata Reeve, 1850. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Dosinia penicillata (Reeve, 1850). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Dosinia pubescens (Philippi, 1847). Reported as: Dosina cf. eunice A. Adams, 1855. Gulf of Thailand. Reported
by: Gemert (2003), ONEP (2007), Nabhitabhata (2009).
Dosinia sanata Fischer-Piette & Delmas, 1967. Andaman Sea. Reported by: Sartori et al. (2008).
Dosinia scabriuscula (Philippi, 1847). Reported as: Dosinia (Asa) scabriuscula (R. A. Philippi, 1847). Reported
by: Huber (2010).
44
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Dosinia sericea Reeve, 1850. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (1998), Aungtonya et al.
(1999), Nabhitabhata (2009), BEDO (2017a).
Dosinia subrosea (Gray, 1835). Andaman Sea. Reported by: BEDO (2017a).
Dosinia tenella Römer, 1860. Gulf of Thailand. Reported by: Kasetsart University (2006), Duangdee (2007),
Nabhitabhata (2009).
Dosinia trailli A. Adams, 1855. Also reported as: Dosinia (Dosinella) trailli A. Adams, 1856. Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Robba et al. (2002), ONEP (2007), Sartori et al. (2008),
Nabhitabhata (2009), Huber (2010), BEDO (2017a), GBIF (2019b).
Doisinia troscheli (Lischke, 1873). Andaman Sea. Reported by: Kasetsart University (2006).
Dosinia tumida (Gray, 1838). Also reported as: Phacosoma tumida (J. E. Gray, 1838). Gulf of Thailand, Andaman
Sea. Reported by: Swennen et al. (2001), Robba et al. (2002), Gemert (2003), ONEP (2007), Nabhitabhata
(2009), Sanpanich (2011), Negri et al. (2014).
Gafrarium aeguivocum (Holten, 1802). Andaman Sea. Reported by: BEDO (2017a).
Gafrarium catillum Hedley, 1909. Gulf of Thailand. Reported by: Sanpanich (2011).
Gafrarium dispar (Holten, 1802). Also reported as: Circe (Crista) dispar Chemnitz, 1795. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya
et al. (1999), Gemert (2003), Duangdee (2004), Duangdee et al. (2007, 2009, 2011, 2013b), Sartori et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Gafrarium divaricatum (Gmelin, 1791). Also reported as: Circe divaricata Chemnitz, 1795; and Gafrarium
(Crista) divaricatum (“Chemnitz” Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889),
Lynge (1909), Suvatti (1937, 1938), Nielsen(1976b), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Amornjaruchit (1988), Kurozumi et al. (1989), Nateewathana (1995), Bussarawit
(1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University
(2000, 2006), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007, 2011), Duangdee et al. (2007),
ONEP (2007), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Huber (2010), BEDO
(2017a), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Gafrarium pectinatum (Linnaeus, 1758). Also reported as: Gafrarium tumidum (Röding, 1798); Circe (Crista)
gibbia Lamarck, 1818; Circe gibba Lamarck; Circe pectinata (Linnaeus, 1758); and Circe (Crista) pectinata
(Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937,
1938), Nielsen(1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981),
Kurozumi et al. (1989), Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998,
2011), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Gemert (2003),
Duangdee (2004, 2007), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Meyer et al.
(2008), Sartori et al. (2008), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019a,
2019b), J. D. Taylor et al. (unpublished data).
Globivenus embrithes (Melvill & Standen, 1899). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich &
Duangdee (2009, 2018), Sanpanich (2011).
Globivenus mindorensis (E. A. Smith, 1916). Andaman Sea. Reported by: Aungtonya & Hylleberg (1998),
Nabhitabhata (2009), BEDO (2017a).
Globivenus toreuma (Gould, 1850). Also reported as: Chione (Ventricola) toreuma Gould, 1850; and Venus toreuma
Gould, 1850. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Aungtonya & Hylleberg (1998),
Sanpanich (1998), Sartori et al. (2008), Nabhitabhata (2009), Duangdee et al. (2013a, 2013b), BEDO (2017a).
Gomphina radiata (G. B. Sowerby II, 1835). Andaman Sea. Reported by: Sartori et al. (2008). This species cannot
be confirmed through WoRMS Editorial Board (2019), but is recorded by Sartori et al. (2008).
Gomphina cf. veneriformis Lamarck, 1818. Andaman Sea. Reported by: Nabhitabhata (2009). This species cannot
be confirmed through WoRMS Editorial Board (2019), but is listed in Bernard et al. (1993).
Gouldia minima (Montagu, 1803). Reported as: Cytherea (Caryatis) minuta Koch, 1845. Gulf of Thailand. Reported
by: Lynge (1909).
Hyphantosoma caperi (Lamprell & Healy, 1997). Reported as: Pitar caperi Lamprell & Healy, 1997. Gulf of
Thailand. Reported by: Kasetsart University (2006).
Hyphantosoma nancyae (Lamprell & Whitehead, 1990). Also reported as: Pitar nancyae Lamprell & Whitehead,
1990. Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), ONEP (2007), Sartori et al. (2008),
Nabhitabhata (2009), Sanpanich (2011).
Hyphantosoma spoori (Lamprell & Whitehead, 1990). Reported as: Pitar spoori Lamprell & Whitehead, 1990.
Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2013a, 2013b).
Irus irus (Linnaeus, 1758). Reported as: Venerupis irus Linnaeus, 1767; and Irus macrophylla (Deshayes,
1853). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979),
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Gemert (2003), Sartori et al. (2008),
Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
45
The Marine and Estuarine Molluscs of Thailand
Lajonkairia elegans (H. Adams, 1870). Reported as: Cyclina elegans H. Adams, 1870. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009). According to WoRMS Editorial Board (2019), the genus Cyclina is valid,
but there is no Cyclina elegans H. Adams, 1870, nor any elegans in the genus. Ruhoff (1980) lists one venerid
described by H. Adams in 1870: Lucinopsis elegans, which is now considered to be Lajonkairia elegans (H.
Adams, 1870).
Lamelliconcha paytensis (d’Orbigny, 1845). Reported as: Cytherea (Caryatis) affinis Sowerby, 1855. Gulf of
Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Lioconcha annettae Lamprell & Whitehead, 1990. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich &
Duangdee (2009, 2018), Duangdee et al. (2011), Sanpanich (2011).
Lioconcha castrensis (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009), Duangdee et al. (2011).
Lioconcha fastigiata G. B. Sowerby II, 1851. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989),
Sanpanich (1998), Aungtonya et al. (1999), Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a).
Lioconcha ornata (Dillwyn, 1817). Also reported as: Circe (Lioconcha) picta Lamarck, 1818. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem
(1998), Aungtonya et al. (1999), Gemert (2003), Duangdee (2004, 2007), Duangdee et al. (2007, 2009),
Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Lioconcha philippinarum (Hanley, 1844). Also reported as: Gouldia philippinarum (Hanley, 1844). Gulf of
Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981),
Bernard et al. (1993), Bussarawit (1995), Aungtonya et al. (1999), Robba et al. (2002), Nabhitabhata (2009),
Sanpanich (2011). Robba et al. (2004) reported that their specimens (Robba et al. [2002]) are actually Callista
erycina Linnaeus, 1758, but it is not clear whether this also applies to the other references.
Lioconcha polita (Röding, 1798). Reported as: Lioconcha sulcatina Lamarck, 1819. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Swennen et al. (2001), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Sanpanich (2011), BEDO (2017a).
Lioconcha rumphii Meij, Moolenbeek & Dekker (2010). Andaman Sea. Reported by: Meij et al. (2010).
Lioconcha trimaculata (Lamarck, 1818). Also reported as: Sulcilioconcha trimaculata (Lamarck, 1818). Gulf of
Thailand. Reported by: Lynge (1909), Gemert (2003), Sartori et al. (2008), Nabhitabhata (2009).
Marcia cordata (Forsskål in Niebuhr, 1775). Also reported as: Marcia flammea (Gmelin, 1791). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), BEDO (2017a).
Marcia hiantina (Lamarck, 1818). Reported as: Marcia (Hemitapes) hiantina (Lamarck, 1818); Marcia rimularis
Lamarck; Katelysia hiantina (Lamarck, 1818); and Tapes (Hemitapes) hiantinus Lamarck, 1818. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Amornjaruchit (1988), Nateewathana (1995),
Carpenter & Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Meyer et al.
(2008), Printrakoon et al. (2008), Sartori et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
(2011), GBIF (2019b).
Marcia japonica (Gmelin, 1791). Also reported as: Katelysia japonica (Gmelin, 1791); and Katelysia striata (Gmelin).
Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya
et al. (1999), Swennen et al. (2001), Gemert (2003), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Marcia opima (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Nabhitabhata
(2009), Huber (2010), BEDO (2017a).
Marcia recens (Holten, 1802). Also reported as: Marcia marmorata (Lamarck, 1818); and Katelysia marmorata
(Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Swennen et al. (2001),
Robba et al. (2002), Gemert (2003), Sartori et al. (2008), Nabhitabhata (2009), Huber (2010), GBIF (2019a).
Meretrix casta (Gmelin, 1791). Also reported as: Meretrix impudica (Lamarck, 1818); and Meretrix ovum (Hanley,
1845). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1938), Kilburn & Hylleberg (1998), Yoosukh &
Matsukuma (2001), Gemert (2003), Duangdee (2007), Nabhitabhata (2009), Sanpanich (2011), J. D. Taylor et
al. (unpublished data).
Meretrix lamarckii Deshayes, 1853. Andaman Sea. Reported by: Amornjaruchit (1988), BEDO (2017a).
Meretrix lusoria (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Nateewathana
et al. (1981), Amornjaruchit (1988), Nateewathana (1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO
(2017a).
Meretrix lyrata (G. B. Sowerby I, 1851). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge
(1909), Carpenter & Niem (1998), Yoosukh & Matsukuma (2001), Nabhitabhata (2009), Huber (2010).
Meretrix meretrix (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Amornjaruchit (1988), Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011),
Aungtonya et al. (1999), Kasetsart University (2000), Swennen et al. (2001), Yoosukh & Matsukuma (2001),
Robba et al. (2002), Gemert (2003), Duangdee (2004), ONEP (2007), Sartori et al. (2008), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Huber (2010), BEDO (2017a), GBIF (2019b).
Meretrix petechialis (Lamarck, 1818). Also reported as: Cytherea petechialis Lamarck, 1818. Gulf of Thailand.
Reported by: Morlet (1889), Suvatti (1938), Nabhitabhata (2009), Huber (2010), GBIF (2019b).
46
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Meretrix planisulcata (G. B Sowerby II, 1854). Gulf of Thailand, Andaman Sea. Reported by: Yoosukh &
Matsukuma (2001), Nabhitabhata (2009).
Paphia amabilis (Philippi, 1847). Andaman Sea. Reported by: Nabhitabhata (2009).
Paphia crassisulca (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (1998), Aungtonya
et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Paphia declivis (G. B. Sowerby II, 1852). Reported as: Paphia (Paratapes) declivis Sowerby, 1852. Gulf of
Thailand. Reported by: Suvatti (1937, 1938), Nabhitabhata (2009).
Paphia inflata (Deshayes, 1854). Also reported as: Paphia (Pitaria) inflata Sowerby. Gulf of Thailand. Reported
by: Suvatti (1938), Nabhitabhata (2009).
Paphia lirata (Philippi, 1848). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Paphia philippiana M. Huber, 2010. Also reported as: Paphia exarata (Philippi, 1846). Gulf of Thailand, Andaman
Sea. Reported by: Swennen et al. (2001), Duangdee (2007), Sartori et al. (2008), Nabhitabhata (2009).
Paphia rotundata (Linnaeus, 1758). Also reported as: Tapes (Paratapes) papilionaceus Lamarck, 1818; Tapes
(Parembola) radiatus Chemnitz, 1784; Tapes radiatus Chemnitz; and Paphia alapapilionis (Röding, 1798). Gulf
of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1938) , Nateewathana et al.
(1981), Aungtonya & Hylleberg (1998), Aungtonya et al. (1999), Kasetsart University (2006), Sartori et al.
(2008), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Paphia semirugata (Philippi, 1847). Also reported as: Tapes (Paratapes) semirugatus Philippi, 1847. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995), Carpenter
& Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Kasetsart University (2006), Sartori et al.
(2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Paphia vernicosa (A. A. Gould, 1861). Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Sartori
et al. (2008).
Paratapes textilis (Gmelin, 1791). Reported as: Paphia textile (Gmelin 1791). Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Gemert (2003), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Paratapes undulatus (Born, 1778). Also reported as: Neotapes undulata (Born, 1778); Tapes (Paratapes)
undulatus Born, 1778; Paphia (Paphia) undulata and Paphia undulata (Born, 1778). Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Chaitiamwong et al. (1971), Nateewathana et al. (1981),
Amornjaruchit (1988), Nateewathana (1995), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998),
Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University (2000), Swennen et al. (2001), Robba et
al. (2002), Gemert (2003), Duangdee (2007), ONEP (2007), Sartori et al. (2008), Duangdee et al. (2009, 2011),
Nabhitabhata (2009), Negri et al. (2014), BEDO (2017a), GBIF (2019a, 2019b).
Pelecyora ceylonica (Dunker, 1865). Andaman Sea. Reported by: BEDO (2017a).
Pelecyora eudeli (Fischer-Piette & Delmas, 1967). Gulf of Thailand. Reported by: GBIF (2019b).
Pelecyora excisa (Röding, 1798). Reported as: Pelecyora sinuata Gmelin, 1791; Pelecyora exilium (G. B. Sowerby
III, 1909); and Dosinia exilium G. B. Sowerby III, 1909. Gulf of Thailand, Andaman Sea. Reported by: Swennen
et al. (2001), Sartori et al. (2008), Nabhitabhata (2009), Huber (2010).
Pelecyora nana (Reeve, 1850). Reported as: Dosinia derupta Romer, 1860; and Phacosoma derupta (Römer,
1860). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002), Sartori et al. (2008), Nabhitabhata
(2009), Negri et al. (2014).
Pelecyora trigona (Reeve, 1850). Reported as: Dosinia trigona (Reeve, 1850); and Sinodia trigona (Reeve, 1850).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Carpenter & Niem (1998),
Robba et al. (2002), Gemert (2003), Sartori et al. (2008), Nabhitabhata (2009), Huber (2010), Sanpanich
(2011), GBIF (2019b).
Pelecyora tripla (Römer, 1860). Reported as: Pelecyora (Pelecyora) tripla (E. Römer, 1860). No Thai locality
provided. Reported by: Huber (2010).
Periglypta albocancellata (M. Huber, 2010). Reported as: Periglypta clathrata (Deshayes, 1854); and Antigona
clathrata (Deshayes, 1854). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Kilburn &
Hylleberg (1998), Aungtonya et al. (1999), Kasetsart University (2006), Duang dee (2007), ONEP (2007),
Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009), GBIF (2019b).
Periglypta corbis (Lamarck, 1818). Reported as: Venus corbis (Lamarck, 1818). Gulf of Thailand. Reported by:
Morlet (1889), Suvatti (1938), Nabhitabhata (2009).
Periglypta crispata (Deshayes, 1854). Also reported as: Paphia crispata (Deshayes, 1854). Gulf of Thailand,
Andaman Sea. Reported by: Bussarawit (1995), Carpenter & Niem (1998), Gemert (2003), Duangdee (2004,
2007), Kasetsart University (2006), ONR & EPP (2006), Nabhitabhata (2009), Huber (2010), GBIF (2019b).
Periglypta puerpera (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976a),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Kurozumi et al. (1989),
Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Robba et al. (2002), Duangdee
(2004), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a), ONEP (2007), Hylleberg
& Aungtonya (2013), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Periglypta reticulata (Linnaeus, 1758). Also reported as: Antigona reticulata (Linnaeus, 1758); Paphia reticulata
(Linnaeus, 1758); and Chione reticulata Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Lynge
47
The Marine and Estuarine Molluscs of Thailand
(1909), Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Kasetsart University
(2000), Duangdee (2007, 2009), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017a),
Sanpanich & Duangdee (2018).
Petricola aequistriata G. B. Sowerby II, 1874. Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Petricola lapicida (Gmelin, 1791). Also reported as: Petricola divergens (Gmelin, 1791). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981),
Aungtonya et al. (1999), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018),
Duangdee et al. (2011), Printrakoon et al. (2016), BEDO (2017a).
Petricola monstrosa (Gmelin, 1791). Also reported as: Claudiconcha monstrosa (Gmelin 1791). Andaman Sea.
Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009),
BEDO (2017a), J. D. Taylor et al. (unpublished data).
Pitar affinis (Gmelin, 1791). Also reported as: Pitarina affinis (Gmelin, 1791); and Pitar leata. Gulf of Thailand,
Andaman Sea. Reported by: Sartori et al. (2008), Sanpanich (2011).
Pitar alabastrum (Reeve, 1863). Andaman Sea. Reported by: Huber (2010), GBIF (2019a).
Pitar citrinus (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Duangdee
(2007), Duangdee et al. (2007), BEDO (2017a), GBIF (2019b).
Pitar deshayesi (Pfeiffer, 1869). Also reported as: Cyatherea (Caryatis) deshayesi (Pfeiffer, 1869). Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Sartori et al. (2008), Nabhitabhata (2009). This species could not be
confirmed through WoRMS Editorial Board (2019) but is listed by Ruhoff (1980) and Sartori et al. (2008).
Pitar gouldii (Reeve, 1864). Also reported as: Costellipitar gouldii (Reeve, 1864). Gulf of Thailand, Andaman Sea.
Reported by: Gemert (2003), Sartori et al. (2008), Nabhitabhata (2009). This species could not be confirmed
through WoRMS Editorial Board (2019) but is listed by Ruhoff (1980) and Sartori et al. (2008).
Pitar hebraeus (Lamarck, 1818). Reported as: Pitarina hebraea (Lamarck, 1818); Circe hebraea (Lamarck,
1818); and Cytherea (Caryatis) hebraea Lamarck, 1818. Gulf of Thailand, Andaman Sea. Reported by: Morlet
(1889), Lynge (1909), Suvatti (1938), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999),
Nabhitabhata (2009), BEDO (2017a).
Pitar laetus (Linnaeus, 1758). Reported as: Pitar marrowae Healy & Lamprell, 1992. Gulf of Thailand. Reported
by: Gemert (2003).
Pitar lineolatus (G. B. Sowerby II, 1854). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Kasetsart
University (2006), Duangdee (2007), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009), Sanpanich (2011).
Pitar pellucidus (Lamarck, 1818). Reported as: Pitar (Pitarina) pellucidum (Lamarck, 1818). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Sanpanich (1998), Swennen et al. (2001), Duangdee
(2004), Nabhitabhata (2009), Duangdee et al. (2013a), GBIF (2019a).
Pitar striatus (J. E. Gray, 1838). Also reported as: Pitarina striata (J. E. Gray, 1838); Pitar (Pitarina) striatus (Gray,
1838); and Cytherea (Caryatis) striata Gray, 1838. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Robba et al. (2002), Sartori et al. (2008), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Pitarina minuta (Koch in Philippi, 1845). Andaman Sea. Reported by: Sartori et al. (2008). This species could
not be confirmed through WoRMS Editorial Board (2019) but is listed by Sartori et al. (2008).
Placamen chloroticum (Philippi, 1849). Also reported as: Clausinella chlorotica Philippi, 1849. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana (1995), Sanpanich (1998), Aungtonya et al. (1999), Robba et al.
(2002), Sartori et al. (2008), Nabhitabhata (2009), BEDO (2017a), GBIF (2019b).
Placamen isabellina (Philippi, 1849). Also reported as: Venus isabellina (Philippi); and Chione (Circomphalus)
isabellina Philippi, 1848. Gulf of Thailand. Reported by: Lynge (1909), Suvatti (1938), Aungtonya et al. (1999),
Swennen et al. (2001), Nabhitabhata (2009).
Placamen lamellatum (Röding, 1798). Reported as: Callista planatella (Lamarck, 1818); Chione (Circomphalus)
calophylla Philippi, 1836; Cytherea (Circomphyllus) calophylla Hanley; Chione (Circomphalus) calophylla
Philippi, 1836; Placamen calophyllum (Philippi, 1836); Placamen tiara (Dillwyn, 1817); and Bassina callophylla
(Philippi, 1836). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1938), Nateewathana
et al. (1981), Carpenter & Niem (1998), Kilburn & Hylleberg (1998), Aungtonya et al. (1999), Swennen et al.
(2001), Robba et al. (2002), Gemert (2003), Duangdee (2004, 2007), Kasetsart University (2006), ONEP (2007),
Meyer et al. (2008), Sartori et al. (2008), Duangdee et al. (2009), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), GBIF (2019a).
Placamen lamellosum (G. B. Sowerby I, 1825). Andaman Sea. Reported by: GBIF (2019a).
Protapes gallus (Gmelin, 1791). Also reported as: Paphia malabarica (Chemnitz, 1782); Paphia gallus (Gemlin,
1791); Paphia (Protapes) gallus form lentiginosa (Reeve, 1864); and Tapes (Protapes) malabaricus Chemnitz,
1782. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nateewathana et al.
(1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Kasetsart
University (2006), ONEP (2007), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich (2011), GBIF (2019b),
J. D. Taylor et al. (unpublished data).
48
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Protapes swenneni Huber (2010). Gulf of Thailand. Reported by: Huber (2010).
Redicirce sulcata (Gray, 1838). Also reported as: Circe sulcata Gray, 1838; and Circe (Redicirce) sulcata Gray,
1838. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Swennen et al. (2001), Gemert (2003), Sartori
et al. (2008), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Ruditapes bruguieri (Hanley, 1845). No Thai locality provided. Reported by: Silina (2014).
Ruditapes decussatus (Linnaeus, 1758). Reported as: Tapes decussatus (Linnaeus, 1758). Gulf of Thailand.
Reported by: Sanpanich (2011).
Ruditapes philippinarum (Adams & Reeve, 1850). Reported as: Tapes philippinarum (A. Adams & Reeve, 1850).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979), Morris
& Purchon (1981), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Sartori et al.
(2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009).
Sunetta concinna Dunker, 1865. Reported as: Sunetta truncata (Deshayes, 1854). Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Kilburn & Hylleberg (1998), Aungtonya et al. (1999), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a).
Sunetta donacina (Gmelin, 1791). Reported as: Sunetta (Sunetta) donacina (J. F. Gmelin, 1791). No Thai locality
provided. Reported by: Huber (2010).
Sunetta effossa (Reeve, 1843). Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001),
Gemert (2003), ONEP (2007), Sartori et al. (2008), Nabhitabhata (2009), BEDO (2017a).
Sunetta menstrualis (Menke, 1843). Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009),
GBIF (2019a).
Sunetta meroe (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Sunetta scripta (Linnaeus, 1758). Andaman Sea. Reported by: Taylor (2009).
Sunetta solanderii (Gray, 1825). Reported as: Sunetta (Solanderina) solanderi Gray. Gulf of Thailand. Reported
by: Suvatti (1937, 1938), Nabhitabhata (2009).
Sunetta sunettina (Jousseaume, 1891). Reported as: Sunetta contempta E. A. Smith, 1891; Sunetta (Sunemeroe)
sunettina (F. P Jousseaume, 1891); and Sunetta (Cyclosunetta) comtempta Smith, 1891. Gulf of Thailand,
Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), ONEP (2007),
Sartori et al. (2008), Nabhitabhata (2009), Huber (2010), Sanpanich (2011), GBIF (2019a).
Tapes belcheri G. B. Sowerby ll, 1852. Also reported as: Tapes obscuratus Deshayes, 1853; and Paphia obscurata
(Deshayes, 1853). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Carpenter & Niem (1998), Sartori
et al. (2008), Nabhitabhata (2009), BEDO (2017a).
Tapes conspersus (Gmelin, 1791). Reported as: Tapes dorsatus (Lamarck, 1818). Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al. (1999), Duangdee (2004), Duangdee
et al. (2007), Nabhitabhata (2009), BEDO (2017a).
Tapes literatus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Morris & Purchon (1981), Nateewathana et al. (1981), Carpenter & Niem (1998), Sanpanich (1998,
2011), Aungtonya et al. (1999), Kasetsart et al. (2004), Wells (2005), Kasetsart University (2006), ONR & EPP
(2006), Duangdee (2007), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2018), BEDO (2017a), GBIF (2019a).
Tapes platyptycha Pilsbry, 1901. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009, 2013b).
Tapes sericeus Matsukuma, 1986. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Sartori
et al. (2008), Duangdee et al. (2009), GBIF (2019a).
Tapes sulcarius (Lamarck, 1818). Gulf of Thailand. Reported by: Sanpanich (2011).
Tawera laticostata (Ohdner, 1917). Gulf of Thailand. Reported by: Sanpanich (2011).
Timoclea arakana (G. Nevill & H. Nevill, 1871). Also reported as: Timoclea cf. farsiana Biggs, 1973. Andaman
Sea. Reported by: Scott (1995), Sartori et al. (2008).
Timoclea costellifera (Adams & Reeve, 1850). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Timoclea habei Fischer-Piette & Vukadinovic, 1977. Reported as: Veremolpa habei (Fischer-Piette & Vukadinovic,
1977). No Thai locality stated. Reported by: Bernard et al. (1993).
Timoclea imbricata (G. B. Sowerby II, 1853). Reported as: Timoclea (Glycodonta) imbricata (Sowerby, 1853); and
Glycydonta imbricata (Sowerby, 1845). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Bernard et al. (1993), Bussarawit (1995), Aungtonya et al. (1999), Duangdee (2004), Sartori et al.
(2008), Nabhitabhata (2009), BEDO (2017a).
Timoclea lionota (E. A. Smith, 1885). Reported as: Timoclea (Timoclea) lionota (Smith, 1885). Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Robba et al. (2002), Sartori et al. (2008).
Timoclea marica (Linnaeus, 1758). Also reported as: Glycydonta marica (Linnaeus, 1758) and Chione
(Omphaloclathrum) marica Sowerby. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Duangdee et al. (2007), Sartori et al.
(2008), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), Sanpanich & Duangdee (2018).
Timoclea cf. micra (Pilsbry, 1904). Reported as: Veremolpa cf. micra (Pilsbry, 1904). Gulf of Thailand. Reported
by: Gemert (2003).
49
The Marine and Estuarine Molluscs of Thailand
Timoclea scabra (Hanley, 1845). Also reported as: Chione (Timoclea) scabra (Hanley, 1844); and Timoclea
(Chioneryx) scabra. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1938), Scott (1995),
Swennen et al. (2001), Robba et al. (2002, 2003), Sartori et al. (2008), Nabhitabhata (2009), Huber (2010),
Negri et al. (2014), GBIF (2019b).
Timoclea scandularis (Hedley, 1909). Gulf of Thailand. Reported by: Sanpanich (2011).
Timoclea siamensis (Lynge, 1909). Reported as: Timoclea (Timoclea) siamensis (Lynge, 1909). Gulf of Thailand.
Reported by: Lynge (1909), Robba et al. (2002), Sartori et al. (2008), Nabhitabhata (2009), Huber (2010).
Timoclea subnodulosa (Hanley, 1845). Also reported as: Chione (Omphaloclathrum) subnodulosa Hanley, 1844;
Timoclea recognita (Smith, 1885); Timoclea (Glycydonta) recognita (Smith, 1885); and Chione recognita E. A.
Smith, 1885. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Swennen et al. (2001), Robba et al.
(2004), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), GBIF
(2019b).
Tivela tripla (Linnaeus, 1771). Reported as: Tivela bicolor (Gray, 1838). Gulf of Thailand. Reported by: Morlet
(1889), Suvatti (1938).
Venerupis aspera (Quoy & Gaimard, 1835). Reported as: Tapes variegatus Sowerby, 1852; Tapes (Ruditapes)
variegatus (Sowerby, 1852); Timoclea variegata (Sowerby, 1853); and Ruditapes variegatus (Sowerby, 1852).
Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Kurozumi et al.
(1989), Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Gemert (2003), Duangdee
(2004, 2007), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), ONEP
(2007), Sartori et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Huber (2010),
Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Venus sinuosa Lamarck, 1818. Reported as: Tapes (Paratapes) sinuosus Lamarck, 1818; and Paphia sinuosa
(Lamarck, 1819). Gulf of Thailand. Reported by: Lynge (1909), Gemert (2003), Nabhitabhata (2009), BEDO
(2017a). WoRMS Editorial Board (2019) lists this species as a nomen dubium as it is an invalid junior homonym
of Venus sinuosa Pennant, 1777.
Subclass Protobranchia
Order Nuculanida
Superfamily Nuculanoidea
Family Malletiidae
Malletia conspicua E. A. Smith, 1895. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017a).
Family Nuculanidae
Jupiteria puellata (Hinds, 1843). Reported as: Nuculana (Jupiteria) puellata Hinds, 1843. Gulf of Thailand.
Reported by: Lynge (1909), Robba et al. (2002, 2003), Nabhitabhata (2009), Negri et al. (2014).
Lembulus belcheri (Hinds, 1843). Reported as: Nuculana belcheri Hinds, 1843; and Yoldia (Yoldia) belcheri
(Hinds, 1843). Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009).
Saccella bellula (A. Adams, 1856). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2002), Nabhitabhata
(2009), BEDO (2017a).
Saccella mauritiana (G. B. Sowerby I, 1833). Also reported as: Nuculana mauritina (G. B. Sowerby I, 1833);
Nuculana (Scaeoleda) mauritiana (Sowerby, 1833); and Nuculana (Scaeoleda) cuspidata (Gould, 1861). Gulf of
Thailand. Reported by: Lynge (1909), Swennen et al. (2001), Robba et al. (2002, 2004), Nabhitabhata (2009),
Negri et al. (2014).
Family Yoldiidae
Megayoldia japonica (Adams & Reeve, 1850). Reported as: Portlandia japonica (Adams & Reeve, 1850). Gulf of
Thailand. Reported by: Scott (1995), Robba et al. (2002), Nabhitabhata (2009).
Orthoyoldia lepidula (A. Adams, 1856). Andaman Sea. Reported by: BEDO (2017a).
Orthoyoldia serotina (Hinds, 1843). Reported as: Nucula serotina Hinds, 1843; and Yoldia cf. serotina (Hinds,
1843). Gulf of Thailand. Reported by: Lynge (1909), Scott (1995), Nabhitabhata (2009).
Orthoyoldia tenella (Hinds, 1843). Reported as: Nucula tenella Hinds, 1843. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009).
Yoldiella retusa (Hinds, 1843). Reported as: Nuculana retusa Hinds, 1843. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009).
50
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Order Nuculida
Superfamily Nuculoidea
Family Nuculidae
Ennucula cumingii Hinds, 1843. Reported as: Nucula cumingii R. B. Hinds, 1843. Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Nabhitabhata (2009), BEDO (2017a).
Ennucula layardii (A. Adams, 1856). Reported as: Nuculoma layardi (A. Adams, 1856). Gulf of Thailand. Reported
by: Scott (1995), Robba et al. (2002), Nabhitabhata (2009).
Ennucula niponica (Smith, 1885). Gulf of Thailand. Reported by: Scott (1995).
Nucula mitralis (Hinds, 1843). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata
(2009), BEDO (2017a).
Nucula tokyoensis (Yokoyama, 1920). Gulf of Thailand. Reported by: Scott (1995).
Sinonucula cyrenoides (Kuroda, 1929). Reported as: Ennucula cyrenoides (Kuroda, 1929). Gulf of Thailand.
Reported by: Scott (1995).
Order Solemyida
Superfamily Solemyoidea
Family Solemyidae
Acharax grandis (Verrill & Bush, 1898). Andaman Sea. Reported as: Solemya cf. grandis Verrill & Bush, 1871.
Reported by: Nabhitabhata (2009).
Solemya japonica Dunker, 1882. Reported as: Acharax japonicus (Dunker, 1833). Andaman Sea. Reported by:
Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata
(2009).
Subclass Pteriomorpha
Order Arcida
Superfamily Arcoidea
Family Arcidae
Acar congenita E. A. Smith, 1885. Reported as: Arca congenita E. A. Smith, 1885; and Barbatia (Acar) iota (Iredale,
1939). Gulf of Thailand. Reported by: Gemert (2003), Nabhitabhata (2009), Sanpanich & Duangdee (2009).
Acar plicata (Dillwyn, 1817). Reported as: Barbatia plicata (Dillwyn, 1817); and Arca (Fossularca) plicata
Chemnitz, 1795. Gulf of Thailand, Andaman Sea. Reported by: Scott (1995), Gemert (2003), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Anadara antiquata (Linnaeus, 1758). Also reported as: Anadara scapha (Gmelin, 1791). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Amornjaruchit (1988), Nateewathana (1995), Vongpanich
(1996), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al. (1999),
Gemert (2003), Duangdee (2004, 2007), Kasetsart University (2006), Duangdee et al. (2007, 2011, 2013b),
ONEP (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Anadara auriculata (Lamarck, 1819). Reported as: Arca auriculata Lamarck, 1819. Gulf of Thailand. Reported
by: Morlet (1889), Suvatti (1938), Nabhitabhata (2009).
Anadara broughtonii (Schrenck, 1867). Reported as: Arca (Scapharca) inflata (Reeve, 1844). Gulf of Thailand.
Reported by: Suvatti (1937, 1938), Carpenter & Niem (1998), Nabhitabhata (2009).
Anadara consociata (E. A. Smith, 1885). Reported as: Scapharca consociata E. A. Smith, 1885. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Anadara cornea (Reeve, 1844). Also reported as: Scapharca cornea (Reeve, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Bernard et al. (1993), Carpenter & Niem (1998), Duangdee et al. (2009), BEDO (2017a).
Anadara craticulata (Nyst, 1848). Also reported as: Anadara clathrata (Reeve, 1843); and Scapharca clathrata
Reeve, 1844. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981),
Bussarawit (1995), Aungtonya & Hylleberg (1998), Aungtonya et al. (1999), Subba Rao & Dey (2000), Robba
et al. (2004), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Anadara crebricostata (Reeve, 1844). Also reported as: Scapharca crebricostata (Reeve, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Robba et al. (2002), Nabhitabhata (2009), Sanpanich (2011).
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The Marine and Estuarine Molluscs of Thailand
Anadara ferruginea (Reeve, 1844). Also reported as: Diluvarca ferruginea (Reeve, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Bernard et al. (1993), Scott (1995), Vongpanich (1996),
Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Aungtonya et al. (1999), Robba et al. (2 002),
Kasetsart University (2006), Duangdee (2007), Duangdee et al. (2009), Nabhitabhata (2009), Huber (2010),
Sanpanich (2011), Negri et al. (2014), BEDO (2017a), GBIF (2019a).
Anadara globosa (Reeve, 1844). Reported as: Scapharca globosa (Tanaka, 1959); and Anadara binakayanensis
(Faustino, 1932). Gulf of Thailand. Reported by: Lynge (1909), Bernard et al. (1993), Carpenter & Niem (1998),
Nabhitabhata (2009), BEDO (2017a).
Anadara gubernaculum (Reeve, 1844). Reported as: Arca (Scapharca) gubernaculum Reeve, 1844. Gulf of
Thailand. Reported by: Suvatti (1937, 1938), Carpenter & Niem (1998), Nabhitabhata (2009), J. D. Taylor et al.
(unpublished data).
Anadara inaequivalvis (Bruguière, 1789). Also reported as: Scapharca inaequivalvis (Bruguière, 1789). Gulf of
Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Nateewathana et al. (1981), Amornjaruchit (1988),
Vongpanich (1996), Carpenter & Niem (1998), Aungtonya et al. (1999), Robba et al. (2002), Duangdee (2007),
ONEP (2007), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich (2011), Negri et al. (2014), J. D.
Taylor et al. (unpublished data).
Anadara indica (Gmelin, 1791). Also reported as: Arca indica Gmelin, 1791; and Scapharca indica (Gmelin, 1791).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Vongpanich (1996), Carpenter & Niem (1998),
Aungtonya et al. (1999), Kasetsart University (2000, 2006), Robba et al. (2002, 2003), Gemert (2003),
Nabhitabhata (2009), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Anadara jousseaumei (Lamy, 1907). Also reported as: Anadara (Anadara) jousseaumei (E. Lamy, 1907). Gulf of
Thailand, Andaman Sea. Reported by: Huber (2010), BEDO (2017a).
Anadara jurata Iredale, 1939. Gulf of Thailand. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Anadara kagoshimensis (Tokunaga, 1906). Reported as: Scapharca subcrenata (Lischke, 1869). Gulf of
Thailand, Andaman Sea. Reported by: Amornjaruchit (1988).
Anadara oceanica (Lesson, 1831). Reported as: Arca maculosa (Reeve, 1844). Gulf of Thailand. Reported by:
Suvatti (1937, 1938), Carpenter & Niem (1998).
Anadara paucigranosa (Dunker, 1889). Reported as: Tegillarca paucigranosa (Dunker, 1889). Gulf of Thailand.
Reported by: Kurozumi et al. (1989).
Anadara pilula (Reeve, 1843). Also reported as: Scapharca pilula (Reeve, 1843). Gulf of Thailand, Andaman Sea.
Reported by: Suvatti (1937, 1938), Vongpanich (1996), Carpenter & Niem (1998), Sanpanich (1998, 2011),
Aungtonya et al. (1999), ONEP (2007), Nabhitabhata (2009), Galli (2015), BEDO (2017a), GBIF (2019b), J. D.
Taylor et al. (unpublished data).
Anadara rhomboidalis (Schumacher, 1817). Reported as: Scapharca rhomboidalis Chemnitz, 1784; Anadara
(Scapharca) rhomboidalis (H. C. F. Schumacher, 1817); and Anadara penangana (Jousseaume, 1893). Gulf of
Thailand. Reported by: Lynge (1909), Nabhitabhata (2009), Huber (2010), GBIF (2019b).
Anadara rotundicostata (Reeve, 1843). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich (2011).
Anadara rufescens (Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
(2011).
Anadara sabinae (Morlet, 1889). Reported as: Scapharca sabinae Morlet, 1889; Tegillarca sabinae (Morlet, 1889);
and Arca sabinae Morlet, 1889. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1938),
Bernard et al. (1993), Nabhitabhata (2009).
Anadara satowi (Dunker, 1882). Gulf of Thailand, Andaman Sea. Reported by: Chaitiamwong et al. (1971), BEDO
(2017a).
Anadara setigericosta (Nyst, 1848). Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999),
Nabhitabhata (2009).
Anadara subrubra (Dunker, 1866). Gulf of Thailand. Reported by: Nabhitabhata (2009).
Anadara tricenicosta (Nyst, 1848). Also reported as: Anadara (Diluvarca) tricenicosta (H. J. P. Nyst, 1848). Gulf
of Thailand, Andaman Sea. Reported by: Vongpanich (1996), Aungtonya et al. (1999), Nabhitabhata (2009),
Huber (2010), Sanpanich (2011), BEDO (2017a).
Anadara troscheli (Dunker, 1852). Also reported as: Anadara trocheli (Dunker). Gulf of Thailand, Andaman Sea.
Reported by: Chaitiamwong et al. (1971), Nateewathana et al. (1981), Amornjaruchit (1988), Nateewathana
(1995), Vongpanich (1996), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Anadara uropigimelana Bory Saint-Vincent, 1824. Also reported as: Arca haloserica Reeve, 1843. Gulf of Thailand,
Andaman Sea. Reported by: Morlet (1889), Suvatti (1938), Aungtonya et al. (1999), Nabhitabhata (2009).
Anadara vellicata (Reeve, 1844). Also reported as: Arca (Barbatia) velata Reeve, 1844. Gulf of Thailand, Andaman
Sea. Reported by: Suvatti (1938), Nateewathana et al. (1981), Vongpanich (1996), Aungtonya & Hylleberg
(1998), Aungtonya et al. (1999), Nabhitabhata (2009).
Arca avellana Lamarck, 1819. Also reported as: Barbatia maculata (Sowerby, 1833), Arca patriarchalis Röding,
1798; and Arca kraussi Philippi, 1848. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen
(1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Bussarawit (1995),
52
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Vongpanich (1996), Carpenter & Niem (1998), Aungtonya et al. (1999), Gemert (2003), Meyer et al. (2008),
Nabhitabhata (2009), BEDO (2017a).
Arca boucardi Jousseaume, 1894. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Arca navicularis (Bruguière, 1789). Also reported as: Arca linter Jonas, 1845; Arca subnavicularis (Iredale); Arca
(Arca) navicularis Bruguière, 1789; and Arca (Arca) fuscomarginata Dunker, 1868. Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937, 1938), Nateewathana et al. (1981), Kurozumi et al.
(1989), Vongpanich (1996), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Swennen
et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee (2004), Wells (2005), Duangdee et al. (2007, 2009,
2011), ONEP (2007), Nabhitabhata (2009), Negri et al. (2014), BEDO (2017a), GBIF (2019a, 2019b).
Arca ventricosa (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Kurozumi et al. (1989), Vongpanich (1996), Carpenter & Niem (1998),
Sanpanich (1998, 2011), Aungtonya et al. (1999), Gemert (2003), Duangdee (2004, 2007), Wells (2005),
Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP
(2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), BEDO (2017a), GBIF (2019a).
Barbatia amygdalumtostum (Röding, 1798). Also reported as: Barbatia (Barbatia) amygdalumtostum (Röding,
1798); Barbatia fusca (Bruguière, 1789); and Barbatia (Ustalarca) bicolorata (Dillwyn, 1817). Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nielsen(1976b), Morris & Purchon (1981),
Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Kilburn & Hylleberg (1998),
Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001),
Gemert (2003), Duangdee (2004, 2007, 2009), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009,
2011, 2013a, 2013b), ONEP (2007), García & Oliver (2008), Nabhitabhata (2009), Hylleberg & Aungtonya
(2013), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Barbatia candida (Helbling, 1779). Reported as: Barbatia helbingi Bruguière, 1789. Gulf of Thailand, Andaman
Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al.
(1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009).
Barbatia cometa (Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO
(2017a).
Barbatia complanata (Bruguière, 1789). Reported as: Barbatia complanata Chemnitz, 1784. Gulf of Thailand.
Reported by: Morlet (1889), Lynge (1909), Nabhitabhata (2009), BEDO (2017a).
Barbatia foliata (Forsskål in Niebuhr, 1775). Also reported as: Barbatia (Barbatia) foliata (Forsskål in Niebuhr,
1775); Barbatia velata (Sowerby, 1843); Barbatia (Abarbatia) velata (Sowerby); Barbatia lima (Reeve, 1844);
and Barbatia (Abarbatia) lima (Reeve). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Kurozumi
et al. (1989), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University
(2000, 2006), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee (2004, 2007), Wells (2005),
ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2018), Negri et al. (2014), BEDO (2017a), GBIF (2019a, 2019b).
Barbatia grayana (Dunker, 1867). Gulf of Thailand. Reported by: García & Oliver (2008), Sanpanich (2011).
Barbatia hachijoensis Hatai, Niino & Kotaka in Niino, 1952. Gulf of Thailand, Andaman Sea. Reported by:
Aungtonya & Hylleberg (1998), Nabhitabhata (2009), BEDO (2017a).
Barbatia lacerata (Bruguière, 1789). Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit
(1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Barbatia multivillosa (Iredale, 1939). Reported as: Barbatia (Arbatia) multivillosa (Iredale, 1939). Gulf of
Thailand. Reported by: Kurozumi et al. (1989).
Barbatia parva G. B. Sowerby I, 1833. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009),
Printrakoon et al. (2016).
Barbatia pistachia (Lamarck, 1819). Gulf of Thailand. Reported by: Gemert (2003).
Barbatia trapezina (Lamarck, 1819). Also reported as: Barbatia (Barbatia) trapezina (Lamarck, 1819); and Arca
(Barbatia) decussata (Sowerby, 1823). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938),
Carpenter & Niem (1998), Gemert (2003), García & Oliver (2008), Nabhitabhata (2009), Duangdee et al. (2011),
GBIF (2019a, 2019b).
Barbatia virescens (Reeve, 1844). Reported as: Barbatia (Savignyarca) virescens (Reeve, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Bussarawit (1995), García & Oliver (2008), Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich (2011), GBIF (2019b).
Bathyarca pectunculoides (Scacchi, 1835). Reported as: Arca (Fossularca) pectunculiformis Dunker, 1866. Gulf
of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Calloarca tenella (Reeve, 1844). Also reported as: Barbatia tenella (Reeve, 1844); and Arca tenella Reeve, 1844.
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Aungtonya et al.
(1999), Nabhitabhata (2009), Duangdee et al. (2013a), BEDO (2017a).
Hawaiarca yamamotoi (Sakurai & Habe in Habe, 1961). Reported as: Barbatia yamamotoi Sakurai & Habe,
1961. Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), Hylleberg & Aungtonya (2013).
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The Marine and Estuarine Molluscs of Thailand
Mabellarca dautzenbergi (Lamy, 1907). Also reported as: Anadara dautzenbergi (Lamy, 1907). Andaman Sea.
Reported by: Bernard et al. (1993), Nabhitabhata (2009), BEDO (2017a).
Mesocibota bistrigata (Dunker, 1866). Reported as: Barbatia bistrigata (Dunker, 1866); Barbatia (Nipponarca)
bistrigata (Dunker, 1866); and Hawaiarca bistrigata (Dunker, 1853). Gulf of Thailand, Andaman Sea. Reported
by: Swennen et al. (2001), Robba et al. (2002), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007).
Mesocibota signata (Dunker, 1868). Reported as: Arca signata Dunker, 1858. Gulf of Thailand. Reported by:
Lynge (1909), Robba et al. (2002), Gemert (2003), Nabhitabhata (2009), Galli (2015).
Miratacar wendti (Lamy, 1907). Also reported as: Barbatia (Barbatia) wendti (Lamy, 1907). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Nabhitabhata (2009), BEDO (2017a).
Tegillarca aequilatera (W. R. Dunker, 1868). No Thai locality provided. Reported by: Huber (2010).
Tegillarca granosa (Linnaeus, 1758). Reported as: Anadara granosa (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Chaitiamwong et al. (1971), Nateewathana et al. (1981), Amornjaruchit (1988),
Gemert (2003), Sanpanich (1998, 2011), Duangdee et al. (2009, 2011), Hylleberg & Aungtonya (2013), Negri
et al. (2014), Premcharoen et al. (2016), BEDO (2017a), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished
data).
Tegillarca nodifera Martens, 1860. Reported as: Arca nodifera von Martens, 1860; Anadara nodifera (von Martens,
1860); Anadara (Tegillarca) nodifera (von Martens, 1860); and Arca oblonga Philippi, 1849. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Suvatti (1938), Chaitiamwong et al. (1971), Nateewathana et al.
(1981), Amornjaruchit (1988), Bernard et al. (1993), Nateewathana (1995), Bussarawit (1995), Vongpanich
(1996), Carpenter & Niem (1998), Aungtonya et al. (1999), Robba et al. (2002, 2003), ONEP (2007),
Nabhitabhata (2009), Huber (2010), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019a).
Tegillarca rhombea (Born, 1778). Andaman Sea. Reported by: Galli (2015), BEDO (2017a).
Tegillarca zanzibarensis (Nyst, 1848). Reported as: Anadara cuneata (Reeve, 1844). Andaman Sea. Reported by:
Vongpanich (1996), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Trisidos semitorta (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Bussarawit (1995), Vongpanich (1996), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Sanpanich
(1998), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Robba et al. (2002),
Gemert (2003), Duangdee (2004, 2007), Wells (2005), Duangdee et al. (2007, 2011), ONEP (2007), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Trisidos torta (Mörch, 1850). Gulf of Thailand. Reported by: Nabhitabhata (2009).
Trisidos tortuosa (Linnaeus, 1758). Also reported as: Parallelipedum tortuosa Linnaeus, 1758; Arca (Barbatia)
tortosa Linnaeus, 1758; Arca (Barbatia) tortuosum; and Trisidos yongei Iredale, 1939. Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Suvatti (1937, 1938), Bernard et al. (1993), Vongpanich (1996), Carpenter &
Niem (1998), Kilburn & Hylleberg (1998), Sanpanich (1998), Aungtonya et al. (1999), Swennen et al. (2001),
Robba et al. (2002), Kasetsart University (2006), ONEP (2007), Nabhitabhata (2009), BEDO (2017a), GBIF
(2019b).
Vitracar sulcata (Lamarck, 1819). Reported as: Scapharca dichotoma Deshayes, 1863. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
Family Cucullaeidae
Cucullaea granulosa Jonas, 1846. Andaman Sea. Reported by: GBIF (2019a).
Cucullaea labiata (Lightfoot, 1786). Reported as: Cucullaea labiata (Solander, 1786); and Arca (Cucullaea)
concamerata Martini, 1777. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al.
(1981), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University (2000,
2006), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee et al. (2007), Nabhitabhata (2009),
BEDO (2017a), GBIF (2019a, 2019b).
Family Glycymerididae
Glycymeris pertusa (Reeve, 1843). Andaman Sea. Reported by: BEDO (2017a).
Glycymeris pilsbryi (Yokoyama, 1920). Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit
(1995), Nabhitabhata (2009), BEDO (2017a).
Glycymeris reevei (Mayer, 1868). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Carpenter & Niem (1998), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Glycymeris tenuicostata (Reeve, 1843). Reported as: Glycymeris amamiensis (Kuroda, 1930). Andaman Sea.
Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Tucetona auriflua (Reeve, 1843). Andaman Sea. Reported by: BEDO (2017a), GBIF (2019a).
Tucetona hanzawai (Nomura & Zinbo, 1934). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Tucetona hoylei (Melvill & Standen, 1899). Reported as: Melaxinaea hoylei (Melvill & Standen, 1899). Andaman
Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Tucetona mindoroensis (E. A. Smith, 1916). No Thai locality provided. Reported by: Huber (2010).
Tucetona montrouzieri (Angas, 1872). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Tucetona pectunculus (Linnaeus, 1758). Reported as: Pectunculus pectiniformis Lamarck, 1819; and Pectunculus
amboinensis Gmelin, 1790. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Carpenter
& Niem (1998), Kilburn & Hylleberg (1998), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Tucetona odhneri Iredale, 1939. Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata (2009).
Tucetona planata (G. Nevill & H. L. Nevill, 1874). Andaman Sea. Reported by: BEDO (2017a).
Tucetona tegulicia (Melvill, 1898). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Family Noetiidae
Arcopsis afra (Gmelin, 1791). Reported as: Striarca afra (Gmelin, 1791). Andaman Sea. Reported by: Nielsen
(1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya et al. (1999), Nabhitabhata (2009),
BEDO (2017a).
Arcopsis sculptilis (Reeve, 1844). Reported as: Arca (Fossularca) sculputilis Reeve, 1844; and Arcopsis minabensis
(Habe, 1981). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nabhitabhata (2009), BEDO (2017a).
Didimacar tenebrica (Reeve, 1844). Also reported as: Arca (Fossularca) tenebrica Reeve, 1844. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a), GBIF
(2019a, 2019b).
Estellacar galactodes (Benson, 1842). Gulf of Thailand. Reported by: Negri et al. (2014).
Estellacar olivacea (Reeve, 1844). Also reported as: Striarca olivacea (Reeve, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Swennen et al. (2001), Robba et al. (2002), ONEP (2007),
Printrakoon et al. (2008), Nabhitabhata (2009), Negri et al. (2014), BEDO (2017a).
Mulinarca aceraea (Melvill & Standen, 1899). Reported as: Striarca acerea (Melvill & Standen, 1899). Gulf of
Thailand. Reported by: Robba et al. (2004), BEDO (2017a).
Noetiella pectunculiformis (Dunker, 1866). Reported as: Scelidionarca pectunculiformis (Dunker, 1866); and
Striarca pectunculiformis (Dunker, 1866). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Aungtonya et al. (1999), Robba et al. (2002), Nabhitabhata (2009), BEDO (2017a).
Ribriarca polycymoides (Thiele & Jaeckel, 1931). Reported as: Arcopsis polycymoides (Thiele & Jaeckel, 1931).
Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Sheldonella lateralis (Reeve, 1844). Reported as: Sheldonella venustopsis (Iredale, 1939). Gulf of Thailand,
Andaman Sea. Reported by: Robba et al. (2002), Gemert (2003), Nabhitabhata (2009), Duangdee et al. (2011),
BEDO (2017a), GBIF (2019a).
Striarca erythraea (Issel, 1869). Gulf of Thailand. Reported by: Printrakoon et al. (2016).
Striarca navicella (Reeve, 1844). Gulf of Thailand. Reported by: Printrakoon et al. (2016).
Striarca symmetrica (Reeve, 1844). Also reported as: Arcopsis symmetrica (Reeve, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Robba et al. (2002), Gemert (2003), Nabhitabhata
(2009), Printrakoon et al. (2016), BEDO (2017a), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Verilarca bivia Iredale, 1939. Reported as: Verilarca sinensis (Thiele & Jaeckel, 1931). Gulf of Thailand, Andaman
Sea. Reported by: Kilburn & Hylleberg (1998), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009),
Hylleberg & Aungtonya (2013).
Verilarca interplicata (Grabau & S. G. King, 1928). Andaman Sea. Reported by: BEDO (2017a).
Verilarca mortenseni (Lynge, 1909). Also reported as: Anadara mortenseni (Lynge, 1909); Arca (Anadara)
mortenseni Lynge, 1909; and Verilarca (Spinearca) mortenseni (Lynge, 1909). Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Scott (1995), Robba et al. (2002), Nabhitabhata (2009), Huber (2010), BEDO
(2017a).
Verilarca thielei (Schenck & Reinhart, 1938). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Superfamily Limopsoidea
Family Limopsidae
Limopsis cf. crenata A. Adams, 1863. Reported as: Crenulilimopsis cf. oblonga (A. Adams, 1860). Andaman Sea.
Reported by: Nabhitabhata (2009).
Limopsis multistriata (Forsskål, 1775). Also reported as: Oblimopa multistriata (Forsskål in Niebuhr, 1775).
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999),
Nabhitabhata (2009), BEDO (2017a).
Limopsis tenisoni Tenison Woods, 1878. Reported as: Limopsis cancellata Reeve, 1843. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Limopsis tokaiensis Yokoyama, 1910. Also reported as: Limopsis tajimae (Sowerby, 1914). Andaman Sea.
Reported by: Nabhitabhata (2009), BEDO (2017a).
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The Marine and Estuarine Molluscs of Thailand
Order Limida
Superfamily Limoidea
Family Limidae
Ctenoides ales (Finley, 1927). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee (2007),
Nabhitabhata (2009), BEDO (2017a).
Ctenoides annulatus (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b),
Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Subba Rao
& Dey (2000), Nabhitabhata (2009), Duangdee et al. (2013a, 2013b), BEDO (2017a).
Ctenoides lischkei (Lamy, 1930). Andaman Sea. Reported by: Kasetsart University (2006), ONR & EPP (2006).
Ctenoides scaber (Born, 1778). Reported as: Ctenoides scabra. Andaman Sea. Reported by: Bussarawit (1995),
Nabhitabhata (2009).
Lima lima (Linnaeus, 1767). Andaman Sea. Reported by: Nielsen (1976a), Tantanasiriwong (1979), Morris &
Purchon (1981), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Wells (2005),
Nabhitabhata (2009), BEDO (2017a).
Lima nimbifer Iredale, 1924. Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Lima tomlini Prashad, 1932. Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Lima vulgaris Link, 1807. Also reported as: Lima lima vulgaris (Link, 1807). Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Aungtonya et al. (1999), Duangdee (2004), Kasetsart University (2006),
ONR & EPP (2006), Duangdee et al. (2009), Nabhitabhata (2009), BEDO (2017a), Sanpanich & Duangdee (2018).
Limaria aurilirata Stuardo, 1968. Reported as: Limaria (Limatulella) aurilirata J. R. Stuardo, 1968. No Thai
locality provided. Reported by: Huber (2010). This species could not be confirmed through WoRMS Editorial
Board (2019).
Limaria basilanica (A. Adams & Reeve, 1850). Gulf of Thailand. Reported by: Aungtonya et al. (1999), Robba
et al. (2002), Gemert (2003), Duangdee (2007), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a).
Limaria dentata (G. B. Sowerby II, 1843). Gulf of Thailand. Reported by: Lynge (1909), Kasetsart University
(2006), Duangdee (2007), Nabhitabhata (2009).
Limaria fragilis (Gmelin, 1791). Also reported as: Lima fragilis (Gmelin, 1791). Gulf of Thailand, Andaman Sea.
Reported by: Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Subba Rao
& Dey (2000), Swennen et al. (2001), Robba et al. (2002), Duangdee (2004, 2007), Wells (2005), Kasetsart
University (2006), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Duangdee et al. (2011),
Sanpanich (2011), Printrakoon et al. (2016), BEDO (2017a).
Limaria orbignyi (Lamy, 1930). Reported as: Lima (Mantellum) angulata Sowerby, 1843. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Limaria orientalis A. Adams & Reeve, 1850. Also reported as: Lima orientalis (Adams & Reeve). Gulf of Thailand,
Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit
(1995), Aungtonya et al. (1999), Swennen et al. (2001), Wells (2005), Kasetsart University (2006), Nabhitabhata
(2009), BEDO (2017a).
Limaria viali (Jousseaume in Lamy, 1919). Reported as: Limatuella viali (Jousseaume in Lamy, 1919). Gulf of
Thailand. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Limatula bullata (Born, 1778). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Order Mytilida
Superfamily Mytiloidea
Family Mytilidae
Adipicola crypta (Dall, Bartsch & Rehder, 1938). Gulf of Thailand. Reported by: Scott (1995).
Amygdalum soyoae Habe, 1958. Reported as: Amygdalum soyae Habe, 1958. Gulf of Thailand. Reported by: Scott
(1995).
Arcuatula arcuatula (Hanley, 1843). Also reported as: Modiola arcuatula Hanley, 1843; and Brachidontes
arcuatulus (Hanley, 1844). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Amornjaruchit (1988),
Carpenter & Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Gemert (2003),
Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009), Huber (2010), Negri et al. (2014), BEDO
(2017a), GBIF (2019b).
Arcuatula glaberrima (Dunker, 1857). Reported as: Modiola glaberrima Dunker, 1856. Gulf of Thailand. Reported
by: Lynge (1909), Nabhitabhata (2009).
56
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Arcuatula japonica (Dunker, 1857). Reported as: Musculista japonica (Dunker, 1857). Gulf of Thailand, Andaman
Sea. Reported by: Robba et al. (2002), Nabhitabhata (2009), Huber (2010), GBIF (2019a).
Arcuatula senhousia (Benson, 1842). Reported as: Musculista senhousia (Benson in Cantor, 1842); Modiolus
senhauseni (Reeve); and Modiola senhausi Reeve. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937,
1938), Chaitiamwong et al. (1971), Nateewathana et al. (1981), Nateewathana (1995), Carpenter & Niem (1998),
Aungtonya et al. (1999), Robba et al. (2002), Kasetsart University (2006), Duangdee (2007), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a).
Botula cinnamomea (Gmelin, 1791). Also reported as: Lithodomus (Botula) cinnamonea Chemnitz, 1758; and
Botula silicula (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a,
1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999),
Robba et al. (2002), Gemert (2003), Kleeman (2007), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009),
Printrakoon et al. (2016), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Brachidontes crebristriatus (Conrad, 1837). Reported as: Brachidontes maritimus (Pilsbry, 1921). Gulf of
Thailand. Reported by: Kasetsart University (2006), Nabhitabhata (2009).
Brachidontes evansi (E. A. Smith, 1903). No Thai locality provided. Reported by: Galli (2015).
Brachidontes mutabilis (Gould, 1861). Reported as: Hormomya mutabilis (Gould, 1861). Gulf of Thailand.
Reported by: Kurozumi et al. (1989).
Brachidontes setiger (W. R. Dunker, 1857). No Thai locality provided. Reported by: Huber (2010).
Brachidontes striatulus (Hanley, 1843). Reported as: Brachidontes emarginatus Bens. (Reeve); and Gregariella
striatula (Hanley, 1843). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1 976b),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Aungtonya et al. (1999),
Nabhitabhata (2009), BEDO (2017a).
Brachidontes subramosus Hanley, 1843. Also reported as: Brachidontes (Hormomys) subramosus Hanley, 1844.
Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Brachidontes variabilis (Krauss, 1848). Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit
(1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Also reported as Brachidontes pharaonis (Fischer, 1870), Gulf of Thailand, by Swennen et al. (2001) and
Nabhitabhata (2009), but these are misidentifications of Brachidontes variabilis (see Appendix 1).
Gregariella barbata (Reeve, 1858). Reported as: Trichomusculus barbatus (Reeve, 1858). Gulf of Thailand.
Reported by: Gemert (2003).
Gregariella coralliophaga (Gmelin, 1791). Reported as: Coralliophaga (Coralliophaga) coralliophaga (Gmelin,
1791); and Botulina coralliophaga (Gmelin, 1791). Andaman Sea. Reported by: Nielsen (1976a, 1976b),
Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Swennen
et al. (2001), Gemert (2003), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Printrakoon et al. (2016),
J. D. Taylor et al. (unpublished data).
Jolya arata (Dunker, 1857). Reported as: Modiolus aratus Dunker, 1857; and Modiolus atratus Dunker, 1856. Gulf
of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009), Sanpanich (2011),
BEDO (2017a), J. D. Taylor et al. (unpublished data).
Jolya elongata (Swainson, 1821). Reported as: Modiolus (Modiolus) elongastus (Swainson, 1821); and Modiola
elongata Swainson, 1821. Gulf of Thailand. Reported by: Lynge (1909), Robba et al. (2002), Nabhitabhata
(2009), Huber (2010), GBIF (2019b).
Jolya rhomboidea (Reeve, 1857). Reported as: Modiola rhomboidea Hanley. Gulf of Thailand. Reported by: Lynge
(1909), Nabhitabhata (2009).
Leiosolenus divaricalx (Iredale, 1939). Reported as: Lithophaga divaricalx Iredale, 1939. Gulf of Thailand,
Andaman Sea. Reported by: Bussarawit (1995).
Leiosolenus hanleyanus (Reeve, 1857). Reported as: Gulf of Thailand, Andaman Sea. Reported by: Bussarawit
(1995), Nabhitabhata (2009).
Leiosolenus kleemanni (Valentich-Scott, 2008). Reported as: Botula kleemanni Valentich-Scott & Tongkerd
(2008). Gulf of Thailand. Reported by: Valentich-Scott & Tongkerd (2008).
Leiosolenus kuehnelti (Kleeman, 1977). Reported as: Lithophaga (Leiosolenus) kuehnelti Kleeman, 1977. Gulf of
Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Leiosolenus lessepsianus (Vaillant, 1865). Reported as: Lithophaga (Leiosolenus) lessepsianus (Vaillant, 1865).
Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (1999), Swennen et al. (2001), Nabhitabhata
(2009), BEDO (2017a).
Leiosolenus lima (Jousseaume in Lamy, 1919). Reported as: Lithophaga lima Lamy, 1919; and Lithophaga
(Diberus) lima Lamy, 1919. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Bussarawit
(1995), Aungtonya et al. (1999), Nabhitabhata (2009), Printrakoon et al. (2016), BEDO (2017a).
Leiosolenus lithurus (Pilsbry, 1905). Reported as: Lithophaga (Diberus) lithurus (Pilsbry). Gulf of Thailand.
Reported by: Kurozumi et al. (1989).
Leiosolenus malaccanus (Reeve, 1857). Also reported as: Lithophaga malaccana (Reeve, 1858); and Lithophaga
(Diberus) malaccana (Reeve, 1857). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a,
1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Kleeman (1984),
57
The Marine and Estuarine Molluscs of Thailand
Bussarawit (1995), Aungtonya et al. (1999), Kasetsart University (2006), ONEP (2007), Valentich-Scott &
Tongkerd (2008), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Printrakoon et al. (2016), J. D. Taylor
et al. (unpublished data).
Leiosolenus nasutus (Philippi, 1846). Reported as: Lithophaga nasuta (Philippi, 1846); and Lithophaga
(Leiosolenus) nasuta (Philippi, 1846). Andaman Sea. Reported by: Nielsen (1976a, 1976b), Tantanasiriwong
(1979), Morris & Purchon (1981), Nateewathana et al. (1981), Kleeman (1984), Bussarawit (1995), Aungtonya
et al. (1999), Subba Rao & Dey (2000), Nabhitabhata (2009), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Leiosolenus obesus (Philippi, 1847). Also reported as: Lithophaga obesus (Philippi, 1847). Andaman Sea. Reported
by: Bussarawit (1995), Aungtonya et al. (1999), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009).
Leiosolenus tripartilus (Jousseaume, 1894). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata
(2009).
Lioberus ligneus (Reeve, 1858). Reported as: Modiola lignea Reeve, 1858; and Modiolus ligneus (Reeve, 1858).
Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009), BEDO (2017a).
Lithophaga caperata (Philippi, 1849). Also reported as: Lithophaga straminea (Reeve, 1857). Andaman Sea.
Reported by: Kleeman (1984), Duangdee et al. (2009), BEDO (2017a), GBIF (2019a).
Lithophaga curta (Lischke, 1874). Reported as: Lithophaga (Leiosolenus) curta (Lischke, 1874). Gulf of Thailand,
Andaman Sea. Reported by: Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009). This species
cannot be confirmed with WoRMS Editorial Board (2019), but is reported by Highsmith (1980) and Kleeman
(1983).
Lithophaga teres (Philippi, 1846). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976a, 1976b),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Kurozumi et al. (1989),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Subba Rao & Dey (2000), Swennen et al.
(2001), Duangdee (2004, 2007), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007), Valentich-Scott
& Tongkerd (2008), Nabhitabhata (2009), Sanpanich (2011), Hylleberg & Aungtonya (2013), Printrakoon et al.
(2016), BEDO (2017a).
Lithophaga zitteliana Dunker, 1882. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Valentich-Scott & Tongkerd (2008), Nabhitabhata (2009), Hylleberg & Aungtonya
(2013).
Modiolatus flavidus (Dunker, 1857). Reported as: Modiolus flavidus (Dunker, 1857). Andaman Sea. Reported by:
Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Aungtonya & Hylleberg (1998),
Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Modiolatus hanleyi (Dunker, 1882). Reported as: Modiola hanleyi Dunker, 1882; and Modiolus hanleyi (Dunker,
1882). Gulf of Thailand. Reported by: Lynge (1909), Sanpanich (1998), Nabhitabhata (2009).
Modiolus areolatus Gould, 1850. Gulf of Thailand. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Modiolus auriculatus (Krauss, 1848). Reported as: Modiolus agripetus (Iredale, 1939). Andaman Sea. Reported
by: Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Kilburn & Hylleberg (1998),
Aungtonya et al. (1999), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019a).
Modiolus modulaides (Röding, 1798). Reported as: Modiolus metcalfei (Hanley, 1843) and Modiolus microptesis
(Deshayes, 1836). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Suvatti (1938), Nateewathana
et al. (1981), Amornjaruchit (1988), Kurozumi et al. (1989), Nateewathana (1995), Carpenter & Niem (1998),
Sanpanich (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Duangdee et al. (2007,
2011), ONEP (2007), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Modiolus philippinarum (Hanley, 1843). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti
(1937, 1938), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998),
Sanpanich (1998), Aungtonya et al. (1999), Robba et al. (2002), Wells (2005), Duangdee et al. (2009), Nabhitabhata
(2009), Sanpanich (2011), Hylleberg & Aungtonya (2013), BEDO (2017a), Sanpanich & Duangdee (2018).
Modiolus rufus (Fischer von Waldheim, 1807). Reported as: Modiolus proclivis Iredale, 1939; and Modiolus rumphii
(R. A. Philippi, 1847). Gulf of Thailand. Reported by: Swennen et al. (2001), Gemert (2003), Huber (2010).
Modiolus tumescens (Clessin, 1888). Reported as: Modiola plumescens Dunker, 1868; and Modiolus plumescens
(Dunker, 1868). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Nabhitabhata (2009), Huber (2010), BEDO (2017a).
Musculus coenobitus (Vaillant, 1865). Also reported as: Modiolaria cuneata Gould, 1861; and Musculus chinensis
Bernard, Cai & Morton, 1993. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), Duangdee et al. (2011), Sanpanich (2011).
Musculus cumingianus (Reeve, 1857). Reported as: Modiolaria cumingiana Dunker (Reeve). Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009).
Musculus cupreus (A. A. Gould, 1861). Gulf of Thailand. Reported by: Kurozumi et al. (1989).
Musculus panhai Moolenbeek, 2009. Also reported as: Modiolaria nana Dunker, 1856. Gulf of Thailand. Reported
by: Lynge (1909), Moolenbeek (2009a), Nabhitabhata (2009).
Musculus viridulus (H. Adams, 1871). Reported as: Modiolaria miranda E. A. Smith, 1884; Musculus mirandus
(Smith, 1884); and Musculista miranda (E. A. Smith, 1884). Gulf of Thailand. Reported by: Lynge (1909),
Nateewathana et al. (1981), Kurozumi et al. (1989), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
58
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Mytella strigata (Hanley, 1843). Also reported as: Musculus (Modiolarca) strigatus (S. C. T. Hanley, 1843). Gulf of
Thailand. Reported by: Huber (2010), Sanpanich & Wells (2019). This is an invasive species from the Caribbean
Sea and eastern Pacific Ocean that was recently found in densities of up to 40,000 m−2 in the upper Gulf of
Thailand.
Mytilisepta virgata (Wiegmann, 1837). Reported as: Septifer virgatus (Wiegmann, 1837). Gulf of Thailand.
Reported by: Duangdee et al. (2007).
Perna viridis (Linnaeus, 1758). Also reported as: Mytilus smaragdinus (Chemnitz, 1785); and Mytilus viridis
Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Chaitiamwong
et al. (1971), Nateewathana et al. (1981), Amornjaruchit (1988), Kurozumi et al. (1989), Bussarawit (1995),
Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Subba Rao
& Dey (2000), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee (2004, 2007), Kasetsart
University (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata (2009), Ramakrishna &
Dey (2010), BEDO (2017a), GBIF (2019a, 2019b).
Rhomboidella malaccana Ockelmann, 1983. Andaman Sea. Reported by: Ockelmann (1983).
Rhomboidella perstriata (Hedley, 1906). Reported as: Musculus perstriata (Hedley, 1906). Gulf of Thailand.
Reported by: Gemert (2003).
Rhomboidella vaillanti (Issel, 1869). Reported as: Solamen vaillanti Issel, 1869. Andaman Sea. Reported by:
Ockelmann (1983), Aungtonya et al. (1999), Nabhitabhata (2009), Huber (2010).
Septifer bilocularis (Linnaeus, 1758). Also reported as: Septifer kraussii (Küster, 1841). Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nielsen (1976b), Tantanasiriwong (1979),
Morris & Purchon (1981), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter &
Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et
al. (2001), Robba et al. (2002), Duangdee (2004, 2007), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011,
2013a, 2013b), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), BEDO (2017a), GBIF
(2019a, 2019b), J. D. Taylor et al. (unpublished data).
Septifer cumingii Récluz, 1849. Reported as: Septifer australis Laseron, 1956. Gulf of Thailand, Andaman Sea.
Reported by: Gemert (2003).
Septifer excisus Weigmann, 1837. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana
et al. (1981), Aungtonya et al. (1999), Swennen et al. (2001), Gemert (2003), Nabhitabhata (2009), Duangdee
et al. (2011, 2013a), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Sinomytilus harmandi (A. T. de Rochebrune, 1881). Gulf of Thailand. Reported by: Huber (2010).
Solamen persicum (E. A. Smith, 1906). Andaman Sea. Reported by: Huber (2010).
Xenostrobus balani Ockelmann, 1983. Gulf of Thailand, Andaman Sea. Reported by: Ockelmann (1983),
Aungtonya et al. (1999), Nabhitabhata (2009), Galli (2015), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Xenostrobus mangle Ockelmann, 1983. Reported as: Limnoperna mangle (K. W. Ockelmann, 1983). Gulf of
Thailand, Andaman Sea. Reported by: Ockelmann (1983), Huber (2010).
Order Ostreida
Superfamily Ostreoidea
Bussarawit & Cedhagen (2010) reported that previous studies on oysters of the families Gryphaeidae and
Ostreidae in Thailand are scarce and there is considerable confusion regarding the scientific names. Inconsistent
taxonomic classification renders most of the published information uncertain. In some cases, unidentified or
misidentified oysters (Nielsen, 1976; Tantanasiriwong, 1979) were preserved in the PMBC collection so that it was
possible for Bussarawit & Cedhagen (2010) to revise them. The list presented here of the two families follows
Bussarawit & Cedhagen (2010) with the taxonomy updated according to WoRMS Editorial Board (2019).
Family Gryphaeidae
Hyotissa hyotis (Linnaeus, 1758). Reported as: Ostrea hyotis Linnaeus, 1758. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya
et al. (1999), Yoosukh & Duangdee (1999), Kasetsart University (2000, 2006), Robba et al. (2002), Duangdee
(2004, 2007), Kasetsart et al. (2004), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b),
ONEP (2007), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), Sanpanich (2011), Hylleberg & Aungtonya
(2013), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Hyotissa inermis (G. B. Sowerby II, 1871). Reported as: Parahyotissa inermis (G. B. Sowerby II, 1871);
Parahyotissa imbricata (Lamarck, 1819); and Hyotissa imbricata (Lamarck, 1819). Gulf of Thailand, Andaman
Sea. Reported by: Aungtonya et al. (1999), Yoosukh & Duangdee (1999), Swennen et al. (2001), Gemert (2003),
Kasetsart University (2006), Duangdee (2007), Nabhitabhata (2009), Sanpanich (2011). Not listed by
Bussarawit & Cedhagen (2010), BEDO (2017a), GBIF (2019b).
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The Marine and Estuarine Molluscs of Thailand
Hyotissa numisma (Lamarck, 1818). Reported as: Parahyotissa numisma (Lamarck, 1819) and Parahyotissa
imbricata (Lamarck, 1819). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya
et al. (1999), Nabhitabhata (2009), Sanpanich (2011), Hylleberg & Aungtonya (2013). Not listed by Bussarawit
& Cedhagen (2010).
Hyotissa sinensis (Gmelin, 1791). Reported as: Parahyotissa sinensis (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich (2011). Not listed by Bussarawit &
Cedhagen (2010), BEDO (2017a).
Family Ostreidae
Alectryonella plicatula (Gmelin, 1791). Reported as: Ostrea plicata var. plicatula (Gmelin, 1791); and
Alectryonella plicata (Gmelin). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Nielsen
(1976a), Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Carpenter & Niem
(1998), BEDO (2017a).
Anomiostrea coralliophila Habe, 1975. Also reported as: Ostrea pyxidata Adams & Reeve, 1850. Gulf of Thailand,
Andaman Sea. Reported by: Aungtonya et al. (1999), Gemert (2003), Nabhitabhata (2009), Hylleberg &
Aungtonya (2013).
Dendostrea cristata (Born, 1778). Reported as: Dendostrea gluacina (Lamarck, 1819). Gulf of Thailand. Reported
by: Swennen et al. (2001).
Dendostrea folium (Linnaeus, 1758). Also reported as: Alectryonella haliotidaea (Lamarck, 1819); Lopha folium
(Linnaeus, 1758); and Pterostrea sp. cf. Pterostrea affinis (Sowerby). Gulf of Thailand, Andaman Sea. Reported
by: Nateewathana et al. (1981), Kurozumi et al. (1989), Aungtonya et al. (1999), Yoosukh & Duangdee (1999),
Kasetsart University (2000), Robba et al. (2002), Duangdee (2004), Bussarawit et al. (2006), Duangdee et al.
(2007, 2009), ONEP (2007), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), Hylleberg & Aungtonya
(2013), BEDO (2017a).
Dendostrea rosacea (Deshayes, 1836). Reported as: Ostrea chemnitzii Hanley, 1846; Parahyotissa chemnitzii
(Hanley, 1846); and Ostrea plicata Chemnitz. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938),
Aungtonya et al. (1999), Robba et al. (2002, 2004), Bussarawit et al. (2006), Kasetsart University (2006),
Nabhitabhata (2009), Bussarawit & Cedhagen (2010), Negri et al. (2014), BEDO (2017a).
Dendostrea sandvichensis (G. B. Sowerby II, 1871). Reported as: Dendostrea crenulifera (Sowerby, 1871); and
Alectryonella crenulifera Sowerby, 1871. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989),
Wells (2005), Bussarawit et al. (2006), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), Hylleberg &
Aungtonya (2013), BEDO (2017a).
Lopha cristagalli (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong
(1979), Nateewathana et al. (1981), Carpenter & Niem (1998), Aungtonya et al. (1999), Duangdee (2004,
2007), Wells (2005), Bussarawit et al. (2006), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007),
Duangdee et al. (2009, 2013a, 2013b), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), BEDO (2017a),
Sanpanich & Duangdee (2018).
Magallana belcheri (G. B. Sowerby II, 1871). Reported as: Crassostrea belcheri (Sowerby, 1871). Gulf of Thailand,
Andaman Sea. Reported by: Chaitiamwong et al. (1971), Nateewathana et al. (1981), Nateewathana (1995),
Aungtonya et al. (1999), Kasetsart University (2000), Klinbunga et al. (2005), Bussarawit et al. (2006), ONEP
(2007), Duangdee et al. (2009), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), BEDO (2017a).
Magallana bilineata (Röding, 1798). Reported as: Crassostrea lugubris (Sowerby, 1871); Crassostrea bilineata
(Röding, 1798); and Crassostrea iredalei Faustino, 1932. Gulf of Thailand, Andaman Sea. Reported by:
Chaitiamwong et al. (1971), Nateewathana (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Kasetsart
University (2000, 2006), Lam & Morton (2004), Klinbunga et al. (2005), Bussarawit et al. (2006), ONEP (2007),
Nabhitabhata (2009), Bussarawit & Cedhagen (2010), J. D. Taylor et al. (unpublished data).
Magallana dactylena (Iredale, 1939). Saccostrea dactylena Iredale, 1939. Gulf of Thailand. Reported by: Gemert
(2003).
Magallana gigas (Thunberg, 1793). Reported as: Crassostrea gigas Thunberg, 1793; and Saccostrea gigas
(Thunberg, 1793). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter & Niem
(1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Printrakoon et al. (2008), Sanpanich
(2011), GBIF (2019b).
Magallana rivularis (Gould, 1861). Reported as: Ostrea rivularis Gould, 1861. Gulf of Thailand. Reported by:
Lynge (1909), GBIF (2019b). This species is shown as a taxon inquirendum by WoRMS Editorial Board (2019).
Nanostrea fluctigera (Jousseaume in Lamy, 1925). Reported as: Nanostrea exigua Harry, 1985. Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Bussarawit & Cedhagen (2010), BEDO (2017a).
Ostrea denselamellosa Lischke, 1869. Gulf of Thailand. Reported by: Robba et al. (2002).
Planostrea pestigris (Hanley, 1846). Also reported as: Ostrea pestigris Hanley, 1846; Dendostrea paulucciae
(Crosse, 1869); Crassostrea paulucciae (Crosse, 1869); and Ostrea paulucciae Crosse, 1869. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Bernard et al. (1993), Carpenter & Niem (1998), Robba et al. (2002),
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Gemert (2003), Lam & Morton (2004, 2009), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Tan & Morton
(2009), Bussarawit & Cedhagen (2010), Negri et al. (2014), BEDO (2017a), GBIF (2019b).
Pustulostrea australis (Lamarck, 1819). Reported as: Pustulostrea tuberculata (Lamarck, 1819); and Ostrea
tuberculata Lamarck, 1804. Andaman Sea. Reported by: Nabhitabhata (2009), Bussarawit & Cedhagen (2010).
Saccostrea cucullata (Born, 1778). Also reported as: Saccostrea forskalii Chemnitz, 1758. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a, 1976b), Tantanasiriwong (1979), Morris & Purchon
(1981), Nateewathana et al. (1981), Amornjaruchit (1988), Bussarawit (1995), Nateewathana (1995),
Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Yoosukh & Duangdee (1999),
Kasetsart University (2000, 2006), Yoosukh & Sukhsangchan (2000), Swennen et al. (2001), Robba et al.
(2002), Gemert (2003), Duangdee (2004, 2007), Klinbunga et al. (2005), Wells (2005), Bussarawit et al. (2006),
ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Printrakoon et al. (2008), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2018), Bussarawit & Cedhagen (2010), Samakraman et al. (2010),
Sørensen (2012), Negri et al. (2014), BEDO (2017a), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Saccostrea echinata (Quoy & Gaimard, 1835). Also reported as: Striostrea (Parastriostrea) mytiloides (Lamarck,
1819). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon
(1981), Nateewathana et al. (1981), Kurozumi et al. (1989), Aungtonya et al. (1999), Yoosukh & Duangdee
(1999), Kasetsart University (2000), Yoosukh & Sukhsangchan (2000), Klinbunga et al. (2005), Bussarawit
et al. (2006), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Bussarawit & Cedhagen (2010), Hylleberg &
Aungtonya (2013), BEDO (2017a), J. D. Taylor et al. (unpublished data). Note: Bussarawit & Cedhagen (2010)
consider the Nielsen (1976b) identification of Saccostrea cuccullata and a number of other identifications as a
misidentification of Saccostrea echinata.
Saccostrea glomerata (Gould, 1850). Reported as: Saccostrea commercialis (Ireland & Roughley, 1933) and
Crassostrea commercialis (Ireland & Roughley). Andaman Sea. Reported by: Chaitiamwong et al. (1971),
Bussarawit et al. (2006).
Saccostrea scyphophilla (Peron & Lesueur, 1807). Reported as: Saccostrea mordax (Gould, 1850). Gulf of
Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Bussarawit (1995).
Superfamily Pinnoidea
Family Pinnidae
Atrina chinensis (Deshayes, 1841). Reported as: Atrina lamellata (Habe, 1961); Pinna chemnitzii Hanley, 1858;
and Atrina (Servatrina) chinensis (Deshayes in Cuvier, 1841). Gulf of Thailand. Reported by: Morlet (1889),
Suvatti (1938), Swennen et al. (2001), Nabhitabhata (2009), Schultz & Huber (2013).
Atrina exusta (Gmelin, 1791). Reported as: Atrina (Servatrina) exusta (Gmelin, 1791). Andaman Sea. Reported
by: Schultz & Huber (2013).
Atrina hystrix (Hanley, 1858). Reported as: Atrina (Servatrina) hystrix (Hanley, 1858). Andaman Sea. Reported
by: Schultz & Huber (2013).
Atrina inflata (Dillwyn, 1817). Also reported as: Pinna inflata Dillwyn, 1787; and Atrina (Servatrina) inflata
(Dillwyn, 1817). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata (2009),
Schultz & Huber (2013).
Atrina pectinata (Linnaeus, 1767). Also reported as: Pinna hanleyi (Reeve, 1858). Gulf of Thailand, Andaman
Sea. Reported by: Suvatti (1937, 1938), Morris & Purchon (1981), Nateewathana et al. (1981), Amornjaruchit
(1988), Bussarawit (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Robba
et al. (2002), Kasetsart University (2006), Duangdee (2007), Duangdee et al. (2007, 2009, 2011), ONEP (2007),
Nabhitabhata (2009), Schultz & Huber (2013), BEDO (2017a), GBIF (2019a, 2019b).
Atrina penna (Reeve, 1858). Reported as: Atrina (Servatrina) penna (Reeve, 1858). Gulf of Thailand.
Atrina serra (Reeve, 1858). Reported as: Pinna serra Reeve, 1858. Gulf of Thailand. Reported by: Lynge (1909),
Nabhitabhata (2009).
Atrina vexillum (Born, 1778). Reported as: Atrina (Atrina) vexillum (Born, 1778). Gulf of Thailand, Andaman
Sea. Reported by: Rosewater (1961), Nielsen (1976a), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Amornjaruchit (1988), Bussarawit (1995), Carpenter & Niem (1998), Sanpanich
(1998, 2011), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Robba et al.
(2002), Duangdee (2004, 2007), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), ONEP
(2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Subba Rao & Dey (2000), Schultz & Huber
(2013), BEDO (2017a), GBIF (2019b), Printrakoon et al. (2019).
Pinna atropurpurea G. B. Sowerby I, 1825. Also reported as: Pinna mutica Reeve, 1858; and Pinna (Subitopinna)
atropurpurea Sowerby I, 1825. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Lynge
(1909), Aungtonya et al. (1999), Nabhitabhata (2009), Schultz & Huber (2013), BEDO (2017a), Printrakoon
et al. (2019).
Pinna attenuata Reeve, 1858. Also reported as: Pinna (Cyrtopinna) attenuata Reeve, 1858. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009), Schultz & Huber (2013).
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The Marine and Estuarine Molluscs of Thailand
Pinna bicolor Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: Rosewater (1961), Nielsen (1976a),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Amornjaruchit (1988),
Bussarawit (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart
University (2000), Subba Rao & Dey (2000), Swennen et al. (2001), Robba et al. (2002), Duangdee (2004, 2007),
Duangdee et al. (2007, 2009), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018),
Hylleberg & Aungtonya (2013), BEDO (2017a), GBIF (2019b).
Pinna deltodes Menke, 1843. Gulf of Thailand. Reported by: Printrakoon et al. (2019).
Pinna incurva Gmelin, 1791. Also reported as: Pinna (Cyrtopinna) incurva Gmelin, 1791. Gulf of Thailand.
Reported by: Swennen et al. (2001), Nabhitabhata (2009), Schultz & Huber (2013).
Pinna muricata Linnaeus, 1758. Also reported as: Pinna (Quantulopinna) muricata Linnaeus, 1758. Gulf of
Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Kasetsart University (2000, 2006), Gemert
(2003), Duangdee (2007), ONEP (2007), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Schultz & Huber
(2013), BEDO (2017a).
Streptopinna saccata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998),
Aungtonya et al. (1999), Wells (2005), Duangdee et al. (2009, 2013a, 2013b), Nabhitabhata (2009), Sanpanich
& Duangdee (2018).
Superfamily Pterioidea
Family Malleidae
Malleus albus Lamarck, 1819. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al.
(1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya
et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Robba et al. (2002), Duangdee (2004,
2007), Wells (2005), Duangdee et al. (2007, 2011), ONEP (2007), Tëmkin (2010), BEDO (2017a), Sanpanich &
Duangdee (2018), GBIF (2019b).
Malleus anatinus (Gmelin, 1791). Reported as: Malleus figuratus Chemnitz, 1785. Gulf of Thailand. Reported by:
Lynge (1909).
Malleus legumen Reeve, 1858. Reported as: Malleus (Malvifundus) irregularis Jousseaume, 1894. Gulf of Thailand.
Reported by: Kurozumi et al. (1989).
Malleus malleus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Duangdee et al. (2009), Sanpanich (2011), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019b).
Malleus regula (Forsskål in Niebuhr, 1775). Also reported as: Malleus decurtatus Lamarck, 1819; and Malvifundus
regulus (Forsskål, 1775). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Morris & Purchon (1981),
Nateewathana et al. (1981), Aungtonya et al. (1999), Gemert (2003), Sanpanich (2011), Duangdee et al. (2013b),
BEDO (2017a).
Vulsella minor Röding, 1798. Reported as: Vulsella spongiarum Lamarck, 1819. Gulf of Thailand, Andaman Sea.
Reported by: Sanpanich & Duangdee (2009), Sanpanich (2011).
Vulsella vulsella (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998),
Sanpanich (1998, 2011), Wells (2005), Duangdee et al. (2009, 2011), Tëmkin (2010).
Family Margaritidae
Pinctada albina (Lamarck, 1819). Also reported as: Pinctada fimbriata (Dunker, 1852). Gulf of Thailand,
Andaman Sea. Reported by: Suvatti (1937, 1938), Swennen et al. (2001), Wells (2005), Nabhitabhata (2009),
Duangdee et al. (2011).
Pinctada chemnitzii (Philippi, 1849). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b),
Tantanasiriwong (1979), Nateewathana et al. (1981), Amornjaruchit (1988), Kurozumi et al. (1989),
Aungtonya et al. (1999), Gemert (2003), Nabhitabhata (2009), BEDO (2017a).
Pinctada fucata (Gould, 1850). WoRMS Editorial Board (2019) lists Pinctada imbricata fucata (Gould, 1850) as
an alternate representation. Also reported as: Pinctada lurida (Gould, 1850). Gulf of Thailand, Andaman Sea.
Reported by: Suvatti (1937, 1938), Nabhitabhata (2009), Tëmkin (2010), Sanpanich (2011), BEDO (2017a).
GBIF (2019a).
Pinctada imbricata Röding, 1798. Also reported as: Pteria (Margaritifera) vulgaris Schumacher, 1817; and
Pinctada martensii (Dunker, 1872). Gulf of Thailand. Reported by: Lynge (1909), Chaitiamwong et al. (1971),
Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem
(1998), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Robba et al. (2002), Duangdee (2007,
2011), Nabhitabhata (2009), BEDO (2017a), GBIF (2019b).
Pinctada maculata (Gould, 1850). Also reported as: Pinctada panasesae (Jameson). Gulf of Thailand, Andaman
Sea. Reported by: Kurozumi et al. (1989), Carpenter & Niem (1998), Gemert (2003), Duangdee (2004, 2007),
Kasetsart University (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata (2009),
Sanpanich & Duangdee (2009), Sanpanich (2011), BEDO (2017a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Pinctada margaritifera (Linnaeus, 1758). Reported as: Gulf of Thailand, Andaman Sea. Reported by:
Chaitiamwong et al. (1971), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998),
Sanpanich (1998, 2011), Aungtonya et al. (1999), Duangdee (2004), Wells (2005), Kasetsart University (2006),
ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Printrakoon et al. (2016), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a), J.
D. Taylor et al. (unpublished data).
Pinctada maxima (Jameson, 1901). Gulf of Thailand, Andaman Sea. Reported by: Chaitiamwong et al. (1971),
Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Swennen
et al. (2001), ONEP (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a).
Pinctada nigra (Gould, 1850). Also reported as: Pteria (Margaritifera) nigra Gould, 1850. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2018), Sanpanich (2011), BEDO (2017a).
Pinctada radiata (Leach, 1814). Gulf of Thailand, Andaman Sea. Reported by: Morris & Purchon (1981),
Nateewathana et al. (1981), Duangdee et al. (2009), Nabhitabhata (2009), Sanpanich (2011). WoRMS Editorial
Board (2019) lists Pinctada imbricata radiata (Leach, 1814) as an alternate representation.
Family Pteriidae
Electroma alacorvi (Dillwyn, 1817). Also reported as: Pteria pulchella Reeve, 1857; and Electroma ovata (Quoy
& Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Kurozumi et al.
(1989), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009),
Sanpanich (2011), Hylleberg & Aungtonya (2013), BEDO (2017a).
Isognomon alatus (Gmelin, 1791). Reported as: Perna ephippium Lamarck. Gulf of Thailand, Andaman Sea.
Reported by: Suvatti (1937, 1938), BEDO (2017a).
Isognomon ephippium (Linnaeus, 1758). Also reported as: Perna cumingi Reeve, 1858. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a), Tantanasiriwong (1979), Morris & Purchon (1981),
Nateewathana et al. (1981), Amornjaruchit (1988), Kurozumi et al. (1989), Carpenter & Niem (1998),
Aungtonya et al. (1999), Swennen et al. (2001), Duangdee (2004, 2007), Kasetsart University (2006), ONR &
EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007), Printrakoon et al. (2008),
Nabhitabhata (2009), Tëmkin (2010), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b), J. D. Taylor
et al. (unpublished data).
Isognomon isognomum (Linnaeus, 1758). Also reported as: Perna isognomon Linnaeus, 1758; Perna rudis Reeve,
1858; Perna vespertilio Reeve, 1858; Perna spengleri Chemnitz, 1784; and Isognomon attenuata (Reeve, 1858).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Nielsen (1976a), Tantanasiriwong (1979), Morris
& Purchon (1981), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem
(1998), Aungtonya et al. (1999), Kasetsart University (2000, 2006), Swennen et al. (2001), Robba et al. (2002),
Duangdee (2004, 2007), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a), ONEP
(2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Tëmkin (2010), Sanpanich (2011), BEDO
(2017a), GBIF (2019a, 2019b).
Isognomon legumen (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Kurozumi et al. (1989), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Duangdee et al. (2011, 2013b),
Sanpanich (2011), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Isognomon nucleus (Lamarck, 1819). Also reported as: Isognomon acutirostris (Dunker, 1848); Isognomon
(Parvipinna) nucleus (Lamarck, 1819); Pervipinna nucleus Lamarck, 1819; and Perna nucleus Lamarck, 1819.
It has also been reported as Isognomon vitreum (Reeve, 1853), which is now recognised as Isognomon
californicum (Conrad, 1837), but this is a misidentification. Gulf of Thailand, Andaman Sea. Reported by: Lynge
(1909), Nielsen (1976b), Tantanasiriwong (1979), Kurozumi et al. (1989), Bussarawit (1995), Carpenter &
Niem (1998), Kilburn & Hylleberg (1998), Aungtonya et al. (1999), Swennen et al. (2001), Duangdee (2004,
2007), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2013a, 2013b), ONEP (2007),
Sanpanich & Duangdee (2009), Samakraman et al. (2010), Sanpanich (2011), GBIF (2019b), J. D. Taylor et al.
(unpublished data).
Isognomon perna (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: Morris & Purchon (1981),
Nateewathana et al. (1981), Gemert (2003), Duangdee et al. (2011), BEDO (2017a), GBIF (2019a).
Isognomon spathulatus (Reeve, 1858). Gulf of Thailand. Reported by: Tëmkin (2010), Tëmkin & Printrakoon
(2016).
Pterelectroma physoides (Lamarck, 1819). Reported as: Electroma physoides (Lamarck, 1819); and Electroma
zebra (Reeve, 1857). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Kasetsart University (2006),
ONR & EPP (2006), Sanpanich & Duangdee (2009), Sanpanich (2011).
Pteria admirabilis Z. Wang, 2001. No Thai locality provided. Reported by: Huber (2010).
Pteria avicular (Holten, 1802). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Aungtonya
et al. (1999), Nabhitabhata (2009), Tëmkin (2010), BEDO (2017a).
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The Marine and Estuarine Molluscs of Thailand
Pteria heteroptera (Lamarck, 1819). Reported as: Pteria brevialata (Dunker, 1872). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Sanpanich (1998,
2011), Aungtonya et al. (1999), Robba et al. (2002), Kasetsart University (2006), ONR & EPP (2006), Duangdee
(2007), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2018), BEDO (2017a).
Pteria lata (Gray in Eyre, 1845). Reported as: Pteria serrulata Dunker, 1848. Gulf of Thailand. Reported by: Lynge
(1909), Tëmkin (2010).
Pteria maura (Reeve, 1857). Reported as: Pteria coturnix (Dunker, 1879). Andaman Sea. Reported by:
Nateewathana et al. (1981), Bussarawit (1995), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Pteria peasei (Dunker, 1872). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University (2006), ONR &
EPP (2006), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2018), Sanpanich (2011), BEDO (2017a).
Pteria penguin (Röding, 1798). Also reported as: Pteria penguin form lotorium Lamarck, 1819; and Pteria
semisagita (Lamarck, 1818). Gulf of Thailand, Andaman Sea. Reported by: Chaitiamwong et al. (1971),
Nateewathana et al. (1981), Kurozumi et al. (1989), Smitasiri et al. (1994), Bussarawit (1995), Carpenter &
Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Swennen et al. (2001), Duangdee (2004,
2007), Kasetsart et al. (2004), Wells (2005), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al.
(2007, 2009, 2011, 2013a), ONEP (2007), Nabhitabhata (2009), Tëmkin (2010), BEDO (2017a), Sanpanich &
Duangdee (2018), GBIF (2019a).
Pteria spectrum Reeve, 1857. Gulf of Thailand. Reported by: Lynge (1909).
Pteria straminea (Dunker, 1852). Reported as: Pteria scabriuscula (Reeve, 1857). Gulf of Thailand. Reported by:
Suvatti (1938).
Pteria tortirostris (Dunker, 1849). Gulf of Thailand. Reported by: Aungtonya et al. (1999), Swennen et al. (2001).
Order Pectinida
Superfamily Anomioidea
Family Anomiidae
Anomia acheus Gray, 1849. Also reported as: Anomia sol Reeve, 1859. Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Nateewathana et al. (1981), Aungtonya et al. (1999), Robba et al. (2002), Nabhitabhata
(2009), BEDO (2017a).
Anomia chinensis Philippi, 1849. Also reported as: Anomia amabaeus Gray, 1849. Gulf of Thailand. Reported
by: Lynge (1909), Swennen et al. (2001), Robba et al. (2002), Kasetsart University (2006), ONEP (2007),
Nabhitabhata (2009), Negri et al. (2014).
Anomia simplex d’Orbigny, 1853. Also reported as: Anomis simplex Orbigny. Gulf of Thailand, Andaman Sea.
Reported by: Nabhitabhata (2009), Sanpanich (2011).
Enigmonia aenigmatica (Holten, 1802). Also reported as: Anomia aenigmatica Holten, 1802. Gulf of Thailand,
Andaman Sea. Reported by: Morlet (1889), Nateewathana et al. (1981), Subba Rao & Dey (2000), Printrakoon
et al. (2008), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), J. D. Taylor et al. (unpublished data).
Monia colon (Gray, 1850). Reported as: Monia (s.l.) colon (J. E. Gray, 1850). No Thai locality provided. Reported
by: Huber (2010).
Monia timida Iredale, 1939. Andaman Sea. Reported by: Duangdee et al. (2013b).
Pododesmus macrochisma (Deshayes, 1839). Reported as: Monia macroschisma Deshayes, 1839. Gulf of
Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Family Placunidae
Placuna ephippium (Philipsson, 1788). Also reported as: Placuna sella (Gmelin, 1791); and Placuna (Ephippium)
ephippium (Philipsson, 1788). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937), Kurozumi et al.
(1989), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart University
(2000, 2006), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), ONEP (2007), Nabhitabhata
(2009), BEDO (2017a), GBIF (2019b).
Placuna lincolnii (Gray, 1849). Gulf of Thailand. Reported by: Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich (2011), BEDO (2017a).
Placuna placenta (Linnaeus, 1758). Also reported as: Placuna orbicularis Retzius, 1758. Gulf of Thailand,
Andaman Sea. Reported by: Morlet (1889), Lynge (1909), Suvatti (1937), Nateewathana et al. (1981),
Bussarawit (1995), Carpenter & Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al.
(2002), Duangdee (2004, 2007), Kasetsart University (2006), Duangdee et al. (2007, 2011), ONEP (2007),
Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019b).
Placuna quadrangula (Philipsson, 1788). Andaman Sea. Reported by: Huber (2010), GBIF (2019a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Superfamily Pectinoidea
Family Pectinidae
Aequipecten opercularis (Linnaeus, 1758). Reported as: Pecten cretatus Reeve, 1853. Gulf of Thailand. Reported
by: Lynge (1909).
Amusium pleuronectes (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Chaitiamwong et al. (1971), Nateewathana et al. (1981), Amornjaruchit (1988), Roussy (1991b), Bussarawit
(1995), Nateewathana (1995), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999),
Kasetsart University (2000, 2006), Swennen et al. (2001), Duangdee (2004, 2007), Nabhitabhata (2009),
Mynhardt et al. (2014), BEDO (2017a), Dijkstra & Beu (2018), GBIF (2019a, 2019b).
Anguipecten picturatus Dijkstra, 1995. Andaman Sea. Reported by: Dijkstra & Beu (2018), GBIF (2019a).
Annachlamys striatula (Linnaeus, 1758). Reported as: Chlamys (Annachlamys) macassarensis (Chenu, 1845);
and Annachlamys macassarensis (Chenu, 1845). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991b),
Aungtonya et al. (1999), Swennen et al. (2001), Gemert (2003), Poppe & Groh (2006), Nabhitabhata (2009),
BEDO (2017a), Dijkstra & Beu (2018), GBIF (2019a, 2019b).
Bractechlamys adorabilis (Dijkstra & Roussy, 1994). Gulf of Thailand, Andaman Sea. Reported by: Aungtonya
et al. (1999), Poppe & Groh (2006), Nabhitabhata (2009), Galli (2015), BEDO (2017a), GBIF (2019a).
Bractechlamys nodulifera (G. B. Sowerby II, 1842). Reported as: Decatopecten noduliferus (Sowerby, 1842); and
Nodipecten noduliferus (Sowerby, 1842). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991 b),
Nabhitabhata (2009), Sanpanich (2011), Duangdee et al. (2013a), GBIF (2019a).
Bractechlamys vexillum (Reeve, 1853). Reported as: Comptopallium vexillum (Reeve, 1853); and Pecten
(Chlamys) vexillum (Reeve, 1853). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Carpenter
& Niem (1998), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Bractechlamys schmeltzii (Kobelt, 1892). Reported as: Bractechlamys schaltzii (Kobelt). Gulf of Thailand,
Andaman Sea. Reported by: Kurozumi et al. (1989). It cannot be verified through WoRMS Editorial Board
(2019), but is listed by Bernard et al. (1893).
Complicachlamys wardiana Iredale, 1939. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2009), Sanpanich (2011), Dijkstra & Beu (2018), GBIF (2019b).
Coralichlamys madreporarum (G. B. Sowerby II, 1842). Reported as: Pecten (Chlamys) madreporarum Sowerby;
and Chlamys madreporarum (G. B. Sowerby II, 1842). Gulf of Thailand, Andaman Sea. Reported by: Lynge
(1909), Kurozumi et al. (1989), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), GBIF
(2019b).
Cryptopecten nux (Reeve, 1853). Reported as: Pecten nux Reeve, 1853. Gulf of Thailand, Andaman Sea. Reported
by: Lynge (1909), Bernard et al. (1993), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Decatopecten amiculum (Phillipi, 1851). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991b),
Carpenter & Niem (1998), Kasetsart University (2000), Poppe & Groh (2006), Nabhitabhata (2009), Galli
(2015), GBIF (2019a, 2019b).
Decatopecten plica (Linnaeus, 1758). Also reported as: Pecten subplicatus Sowerby, 1842; and Pecten (Phalium)
plica Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Roussy (1991b), Kilburn &
Hylleberg (1998), Sanpanich (1998, 2011), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017a), GBIF
(2019a, 2019b).
Decatopecten radula (Linnaeus, 1758). Also reported as: Comptopallium radula (Linnaeus, 1758). Gulf of
Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Wells (2005), Nabhitabhata (2009).
Decatopecten strangei (Reeve, 1852). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004),
Nabhitabhata (2009), Sanpanich (2011).
Excellichlamys histrionica (Gmelin, 1791). Also reported as: Pecten (Vola) sannionis Chemnitz. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Roussy (1991b), Bernard et al. (1993), Sanpanich & Duangdee
(2009, 2018), Sanpanich (2011), BEDO (2017a), GBIF (2019a, 2019b).
Excellichlamys spectabilis (Reeve, 1853). Also reported as: Pecten spectabilis (Reeve, 1853). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Sanpanich (2011),
Duangdee et al. (2013b), Galli (2015), BEDO (2017a), Dijkstra & Beu (2018), GBIF (2019a).
Glorichlamys quadrilirata (Lischke, 1870). Reported as: Bractaechlamys quadrilirata (Lischke, 1868). Gulf of
Thailand, Andaman Sea. Reported by: Dijkstra & Knudsen (1998), Nabhitabhata (2009), Sanpanich & Duangdee
(2009), Sanpanich (2011), BEDO (2017a). GBIF (2019a).
Gloripallium pallium (Linnaeus, 1758). Also reported as: Crytopecten pallium Linnaeus, 1758; and Chlamys
pallium (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991b), Bussarawit (1995),
Carpenter & Niem (1998), Aungtonya et al. (1999), Kasetsart et al. (2004), Wells (2005), Nabhitabhata (2009),
Duangdee et al. (2013a, 2013b), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a).
Gloripallium speciosum (Reeve, 1853). Reported as: Crytopecten speciosus (Reeve, 1853). Gulf of Thailand,
Andaman Sea. Reported by: Roussy (1991b), Nabhitabhata (2009), BDO (2017a).
Glorichlamys elegantissima (Deshayes, 1863). Andaman Sea. Reported by: Dijkstra & Beu (2018), GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Haumea rehderi (Grau, 1960). Andaman Sea. Reported by: Dijkstra & Beu (2018), GBIF (2019a, 2019b).
Hemipecten forbesianus A. Adams & Reeve, 1849. Andaman Sea. Reported by: Dijkstra & Beu (2018), GBIF
(2019a).
Juxtamusium maldivense (E. A. Smith, 1903). Andaman Sea. Reported by: Dijkstra & Beu (2018), GBIF (2019a).
Laevichlamys cuneatus (Reeve, 1853). Also reported as: Chlamys irregularis (Sowerby, 1842). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), Duangdee
et al. (2013b), Dijkstra & Beu (2018), GBIF (2019a).
Laevichlamys deliciosa (Iredale, 1939). Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Laevichlamys gladysiae (Melvill, 1888). Gulf of Thailand. Reported by: Dijkstra (2013), GBIF (2019b).
Laevichlamys mollita (Reeve, 1853). Reported as: Chlamys mollita (Reeve, 1853). Gulf of Thailand, Andaman Sea.
Reported by: Nabhitabhata (2009), Sanpanich (2011), Duangdee et al. (2013b).
Laevichlamys squamosa (Gmelin, 1791). Reported as: Chlamys squamosa (Gmelin, 1791); and Chlamys larvata
(Reeve, 1853). Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Roussy (1991b), Bussarawit
(1995), Sanpanich (1998), Nabhitabhata (2009), Dijkstra & Beu (2018), Sanpanich & Duangdee (2018), GBIF
(2019a, 2019b).
Laevichlamys superficialis (Forsskål in Niebuhr, 1775). Reported as: Chlamys superficialis (Forsskål in Niebuhr,
1775). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Mimachlamys albolineata (G. B. Sowerby II, 1842). Also reported as: Chlamys albolineata (Sowerby, 1825). Gulf
of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981),
Roussy (1991b), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a), Dijkstra & Beu (2018), GBIF
(2019a).
Mimachlamys cloacata (Reeve, 1853). Also reported as: Chlamys (Argopecten) pelseneeri (Dautzenberg & Bavay,
1912); Chlamys curtisiana (Iredale, 1939); Chlamys cloacata (Reeve, 1853); Mimachlamys cloacta (Reeve,
1853); and Pecten rugosus G.B. Sowerby I, 1842. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Roussy (1991b), Swennen et al. (2001), Robba et al. (2002), Nabhitabhata (2009), Sanpanich (2011), Galli
(2015), Dijkstra & Beu (2018), GBIF (2019a, 2019b).
Mimachlamys crassicostata (Sowerby II, 1842). Reported as: Chlamys senatoria nobilis (Reeve, 1852). Gulf of
Thailand. Reported by: Sanpanich (2011).
Mimachlamys gloriosa (Reeve, 1853). Also reported as: Mimachlamys glovisa (Reeve, 1853). Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata (2009), Sanpanich (2011), Dijkstra (2013),
GBIF (2019b).
Mimachlamys sanguinea (Linnaeus, 1758). Also reported as: Chlamys (Mimachlamys) senatoria (Gmelin, 1791);
Mimachlamys senatoria (Gmelin, 1791); Pecten senatorius Gemlin, 1791; and Chlamys senatoria (Gmelin, 1791).
Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nateewathana et al. (1981),
Roussy (1991b), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya et al. (1999), Kasetsart
University (2000), Swennen et al. (2001), Robba et al. (2002), Gemert (2003), Duangdee (2004), Duangdee
et al. (2009), Nabhitabhata (2009), Sanpanich & Duangdee (2009), BEDO (2017a), Dijkstra & Beu (2018), GBIF
(2019a, 2019b).
Minnivola pyxidata (Born, 1778). Also reported as: Pecten pyxidata (Born, 1780); Pecten (Minnivola) pyxidatus
(Born, 1778); and Minnivola pyxidata (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Roussy (1991b), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Sanpanich (1998, 2011), Aungtonya
et al. (1999), Swennen et al. (2001), Robba et al. (2002), Poppe & Groh (2006), Nabhitabhata (2009), Sanpanich
& Duangdee (2009), Duangdee et al. (2011), BEDO (2017a), Dijkstra & Beu (2018), GBIF (2019a, 2019b).
Mirapecten mirificus (Reeve, 1853). Andaman Sea. Reported by: Dijkstra (2013), Dijkstra & Beu (2018), GBIF
(2019a).
Pecten excavatus Anton, 1838. Andaman Sea. Reported by: Galli (2015).
Pedum spondyloideum (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Nielsen (1976b),
Tantanasiriwong (1979), Morris & Purchon (1981), Nateewathana et al. (1981), Kurozumi et al. (1989),
Bussarawit (1995), Aungtonya et al. (1999), Duangdee (2004, 2007), Wells (2005), Duangdee et al. (2007,
2009, 2011, 2013a, 2013b), ONEP (2007), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), Dijkstra &
Beu (2018), Sanpanich & Duangdee (2018), GBIF (2019a).
Scaeochlamys squamata (Gmelin, 1791). Reported as: Chlamys squamata (Gmelin, 1791). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009).
Semipallium dringi (Reeve, 1853). Reported as: Complicachlamys dringi (Reeve, 1853). Gulf of Thailand.
Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), Dijkstra & Beu (2018),
GBIF (2019b).
Semipallium flavicans (Linnaeus, 1758). Reported as: Semipallium tigris (Lamarck, 1819). Andaman Sea. Reported
by: Roussy (1991b), Nabhitabhata (2009), GBIF (2019a).
Semipallium fulvicostatum (A. Adams & Reeve, 1850). Reported as: Pecten fulvicostatus A. Adams & Reeve,
1848. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Semipallium marybellae Raines, 1996. Andaman Sea. Reported by: Galli (2015).
Serratovola angusticostata Dijkstra, 2008. Andaman Sea. Reported by: Dijkstra (2008), GBIF (2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Serratovola gardineri (E. A. Smith, 1903). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991b),
Nabhitabhata (2009), GBIF (2019a).
Serratovola rubicunda (Récluz, 1843). Reported as: Serratovola passerina (Hinds, 1845); Serratovolva tricarinata
(Anton, 1839); and Pecten tricarinatus Anton, 1939. Gulf of Thailand, Andaman Sea. Reported by: Roussy
(1991b), Scott (1995), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Volachlamys singaporina (Sowerby II, 1842). Also reported as: Pecten singaporinus Sowerby, 1853. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Roussy (1991b), Carpenter & Niem (1998), Aungtonya et
al. (1999), Swennen et al. (2001), Gemert (2003), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a, 2019b).
Volachlamys tranquebarica (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991b),
Aungtonya et al. (1999), Nabhitabhata (2009), Galli (2015), BEDO (2017a), GBIF (2019a).
Ylistrum japonicum (Gmelin, 1791). Reported as: Amusium japonicum taiwanense Dijkstra, 1988. Andaman Sea.
Reported by: Roussy (1991b), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
Family Propeamussiidae
Propeamussium caducum (E. A. Smith, 1885). Andaman Sea. Reported by: BEDO (2017a).
Propeamussium investigatoris (E. A. Smith, 1906). Andaman Sea. Reported by: BEDO (2017a).
Parvamussium pauciliratum (E. A. Smith, 1903). Reported as: Amusium paucilirata Smith, 1903. Gulf of Thailand,
Andaman Sea. Reported by: Lynge (1909), Nabhitabhata (2009), GBIF (2019a).
Parvamussium scitulum (E. A. Smith, 1885). Andaman Sea. Reported by: GBIF (2019a).
Family Spondylidae
Spondylus anacanthus Mawe, 1823. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Sanpanich (2011), Duangdee et
al. (2013a), BEDO (2017a).
Spondylus asiaticus Chenu, 1844. No Thai locality provided. Reported by: Huber (2010).
Spondylus asperrimus G. B. Sowerby II, 1847. Andaman Sea. Reported by: GBIF (2019a).
Spondylus butleri Reeve, 1856. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Carpenter
& Niem (1998), Aungtonya et al. (1999), Nabhitabhata (2009), Sanpanich (2011), BEDO (2017a), GBIF (2019a).
Spondylus croceus Schreibers, 1793. Gulf of Thailand. Reported by: Swennen et al. (2001), Duangdee (2007),
Nabhitabhata (2009), GBIF (2019b).
Spondylus crassisquama Lamarck, 1819. Reported as: Spondylus princeps Broderip, 1833. Andaman Sea.
Reported by: Bussarawit (1995), Nabhitabhata (2009).
Spondylus echinatus Schreibers, 1793. Reported as: Spondylus albibarbatus Sowerby, 1847; Spondylus castus
Reeve, 1856; and Spondylus spectrum Reeve, 1856. Gulf of Thailand, Andaman Sea. Reported by: Lynge (1909),
Gemert (2003), Nabhitabhata (2009), Sanpanich (2011), Duangdee et al. (2013a).
Spondylus imperialis (Chenu, 1843). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Swennen et al. (2001), Nabhitabhata (2009), Sanpanich (2011).
Spondylus lamarcki Chenu, 1845. Gulf of Thailand. Reported by: Sanpanich (2011).
Spondylus layardi Reeve, 1856. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009).
Spondylus limbatus (Lamarck, 1819). Reported as: Spondylus coccineus Lamarck, 1819. Gulf of Thailand. Reported
by: Lynge (1909).
Spondylus multimuricatus Reeve, 1846. Gulf of Thailand. Reported by: Huber (2010).
Spondylus nicobaricus Schreibers, 1793. Also reported as: Spondylus nicobanicus Schreibers, 1793; Spondylus
hysterix Röding, 1798; Spondylus hystrix Roeding; and Spondylus percea Iredale, 1939. Gulf of Thailand, Andaman
Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Morris & Purchon (1981), Bussaraw it (1995),
Robba et al. (2002), Gemert (2003), Kasetsart University (2006), ONR & EPP (2006), Duangdee (2007), ONEP
(2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Sanpanich (2011), Duangdee et al. (2013a,
2013b), Hylleberg & Aungtonya (2013), BEDO (2017a).
Spondylus occidens G. B. Sowerby III, 1903. Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Spondylus sinensis Schreibers, 1793. Also reported as: Spondylus ducalis Röding, 1798. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Suvatti (1937, 1938), Nielsen(1976b), Tantanasiriwong (1979), Nateewathana
et al. (1981), Aungtonya et al. (1999), Duangdee (2004), Kasetsart University (2006), ONR & EPP (2006),
Duangdee (2007), Nabhitabhata (2009), Sanpanich (2011), Hylleberg & Aungtonya (2013), BEDO (2017a).
Spondylus squamosus Schreibers, 1793. Also reported as: Spondylus barbatus Reeve, 1856. Gulf of Thailand,
Andaman Sea. Reported by: Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Aungtonya
et al. (1999), Robba et al. (2002), Gemert (2003), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a, 2019b).
Spondylus varius G. B. Sowerby I, 1827. Also reported as: Spondylus varians. Gulf of Thailand, Andaman Sea.
Reported by: Kasetsart University (2006), ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009, 2013a,
2013b), Nabhitabhata (2009), BEDO (2017a), GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Spondylus versicolor Schreibers, 1793. Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995),
Carpenter & Niem (1998), Sanpanich (1998), Aungtonya et al. (1999), Duangdee (2004, 2007), Kasetsart
University (2006), ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata
(2009), Sanpanich (2011), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019a).
Spondylus victoriae G. B. Sowerby II, 1860. Reported as: Spondylus wrightianus Crosse, 1872. Andaman Sea.
Reported by: Nabhitabhata (2009), BEDO (2017a).
Spondylus virgineus Reeve, 1856. Gulf of Thailand. Reported by: Sanpanich (2011).
Superfamily Plicatuloidea
Family Plicatulidae
Plicatula australis Lamarck, 1819. Also reported as: Plicatulostrea onca Simone & Simão de Amaral (2008). Gulf
of Thailand, Andaman Sea. Reported by: Nielsen (1976b), Tantanasiriwong (1979), Nateewathana et al. (1981),
Kurozumi et al. (1989), Aungtonya et al. (1999), Gemert (2003), Kasetsart University (2006), Duangdee (2007),
Nabhitabhata (2009), Sanpanich & Duangdee (2009), Dornellas & Simone (2011), Duangdee et al. (2011,
2013b), Sanpanich (2011), BEDO (2017a).
Plicatula horrida Dunker, 1882. Gulf of Thailand. Reported by: Gemert (2003).
Plicatula muricata G. B. Sowerby II, 1873. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Aungtonya et al. (1999), Robba et al. (2002), Kasetsart University (2006), Simone & Simão do Amaral
(2008), Nabhitabhata (2009), Duangdee et al. (2011), BEDO (2017a), Sanpanich & Duangdee (2018), GBIF (2019b).
Plicatula plicata (Linnaeus, 1767). Also reported as: Plicatula pricata (Linnaeus); Plicatula chinensis Mörch,
1853; Plicatula philippinarum Hanley, 1856; and Plicatula imbricata Menke, 1843. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Kurozumi et al. (1989), Sanpanich (1998, 2011), Aungtonya et al.
(1999), Robba et al. (2002), Gemert (2003), Duangdee (2004, 2007), Kasetsart University (2006), ONR & EPP
(2006), Duangdee et al. (2007, 2009, 2013a), ONEP (2007), Simone & Simão do Amaral (2008), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2018), BEDO (2017a).
Plicatula regularis Philippi, 1849. Reported as: Plicatula simplex (Gould, 1861). Andaman Sea. Reported by:
Nateewathana et al. (1981), Aungtonya et al. (1999), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017a).
Class Cephalopoda
Subclass Nautiloida
Family Nautilidae
Nautilus pompilius Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a),
Nabhitabhata & Wongkamhaeng (2012), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a), Nabhitabhata
et al. (2019).
Subclass Coleoidea
Superorder Decapodiformes
Order Myopsida
Family Loliginidae
Loliolus (Loliolus) affinis Steenstrup, 1856. Reported as: Loliolus affinis Steenstrup, 1856. Gulf of Thailand,
Andaman Sea. Reported by: Chotiyaputta et al. (1992), Nateewathana (1992), Chaitiamwong (1993), Vecchione
et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb & Roper (2010), Aungtonya et al. (2014), BEDO (2017d),
GBIF (2019b).
Loliolus (Loliolus) hardwickei (Gray, 1849). Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al.
(2019).
Loliolus (Nipponololigo) beka (Sasaki, 1929). Reported as Loligo beka Sasaki, 1929. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana (1992), Vecchione et al. (2005), Nabhitabhata et al. (2009a), Jereb & Roper
(2010), Aungtonya et al. (2014), BEDO (2017d), Nabhitabhata et al. (2019).
Loliolus (Nipponololigo) japonica (Hoyle, 1885). Gulf of Thailand. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Loliolus (Nipponololigo) sumatrensis (d’Orbigny [in Férussac & d’Orbigny], 1835). Also reported as: Loligo
sumatrensis d’Orbigny, 1835. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Nateewathana (1992), Chaitiamwong (1993), Vecchione et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb
& Roper (2010), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a).
Loliolus (Nipponololigo) uyii (Wakiya & Ishikawa, 1921). Also reported as: Loligo cf. tagoi Sasaki, 1929. Gulf
of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Aungtonya et al. (2014), BEDO (2017d),
Nabhitabhata et al. (2019).
Sepioteuthis lessoniana Ferussac, 1825. Also reported as: Sepioteuthis lessoniana Lesson, 1830. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Nateewathana (1992),
Chaitiamwong (1993), Vecchione et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb & Roper (2010),
Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a, GBIF (2019b).
Sepioteuthis sepioidea (Blainville, 1823). Andaman Sea. Reported by: Nateewathana et al. (1981), GBIF (2019a).
Uroteuthis (Photololigo) chinensis (Gray, 1849). Reported as: Loligo chinensis Gray, 1849. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Nateewathana (1992),
Chaitiamwong (1993), Vecchione et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb & Roper (2010),
Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a).
Uroteuthis (Photololigo) duvaucelii (d’Orbigny [in Férussac & d’Orbigny], 1835). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Nateewathana (1992), Chaitiamwong
(1993), Vecchione et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb & Roper (2010), Aungtonya et al.
(2014), BEDO (2017d), GBIF (2019a, 2019b).
Uroteuthis (Photololigo) edulis (Hoyle, 1885). Reported as: Loligo edulis Hoyle, 1885. Gulf of Thailand, Andaman
Sea. Reported by: Chotiyaputta et al. (1992), Nateewathana (1992), Vecchione et al. (2005), Nabhitabhata et al.
(2009a, 2019), Jereb & Roper (2010), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019b).
Uroteuthis (Photololigo) sibogae (Adam, 1954). Reported as Loligo (Doryteuthis) sibogae (Adam, 1854). Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana (1992), Chaitiamwong (1993), Nabhitabhata et al. (2009a,
2019), Jereb & Roper (2010), Aungtonya et al. (2014), BEDO (2017d).
Uroteuthis (Photololigo) singhalensis (Ortmann, 1891). Gulf of Thailand, Andaman Sea. Reported by:
Chotiyaputta et al. (1992), Vecchione et al. (2005), Nabhitabhata et al. (2009a, 2019), Jereb & Roper (2010),
Aungtonya et al. (2014), BEDO (2017d).
Order Oegopsida
Family Ancistrocheiridae
Ancistrocheirus lesueurii (Orbigny, 1842, in Férussac & Orbigny, 1834–1848). Gulf of Thailand, Andaman Sea.
Reported by: Aungtonya et al. (2014), BEDO (2017d), Nabhitabhata et al. (2019).
Family Brachioteuthidae
Brachioteuthis picta Chun, 1910. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al.
(2019).
Brachioteuthis riisei (Steenstrup, 1882). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Family Chiroteuthidae
Chiroteuthis imperator Chun, 1910. Reported as: Chiroteuthis (Chirotauma) imperator Chun, 1908. Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana (1995, 1997b), Nabhitabhata et al. (2009a, 2019),
Aungtonya et al. (2014), BEDO (2017d).
Chiroteuthis picteti Joubin, 1894. Andaman Sea. Reported by: Aungtonya et al. (2014).
Chiroteuthis veranii (Férussac, 1834). Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Grimalditeuthis bonplandii (Vérany, 1839). Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al.
(2019).
Family Cranchiidae
Bathothauma lyromna Chun, 1906. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2019).
Cranchia scabra Leach, 1817. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019),
BEDO (2017d).
Egea inermis Joubin, 1933. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO
(2017d).
Leachia pacifica (Issel, 1908). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019),
BEDO (2017d).
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The Marine and Estuarine Molluscs of Thailand
Liguriella podophthalma Issel, 1908. Gulf of Thailand. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO
(2017d).
Liocranchia reinhardti (Steenstrup, 1856). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1995b),
Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Sandalops melancholicus Chun, 1906. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Family Cycloteuthidae
Cycloteuthis sirventi Joubin, 1919. Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Discoteuthis discus Young & Roper, 1969. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Family Enoploteuthidae
Abralia (Abralia) armata (Quoy & Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta
et al. (1992), Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Abralia (Abralia) multihamata Sasaki, 1929. Reported as: Abralia (Stenabralia) lucens Voss, 1962. Andaman
Sea. Reported by: Nateewathana (1995b), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO
(2017d).
Abralia (Abralia) spaercki Grimpe, 1931. Gulf of Thailand, Andaman Sea. Reported by: Aungtonya et al. (2014),
Nabhitabhata et al. (2019).
Abralia (Hererabralia) andamanica Goodrich, 1896. Also reported as: Abralia (Asteroteuthis) andamanica
Goodrich, 1986. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1995b), Nabhitabhata et al.
(2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Abralia steindachneri Weindl, 1912. Andaman Sea. Reported by: Nabhitabhata et al. (2019).
Abraliopsis (Abraliopsis) hoylei (Pfeffer, 1884). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Enoploteuthis reticulata Rancurel, 1970. Reported as: Enoploteuthis (Enoploteuthis) reticulata Rancurel, 1970.
Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Family Histioteuthidae
Histioteuthis pacifica (Voss, 1962). Reported as: Histioteuthis celetaria pacifica (Voss, 1962). Andaman Sea.
Reported by: Nateewathana (1995b), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Stigmatoteuthis hoylei (Goodrich, 1896). Also reported as: Histioteuthis hoylei (Goodrich, 1896). Andaman Sea.
Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Family Joubiniteuthidae
Joubiniteuthis portieri (Joubin, 1916). Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Family Lepidoteuthidae
Lepidoteuthis grimaldii Joubin, 1895. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Family Mastigoteuthidae
Idioteuthis cordiformis (Chun, 1908). Reported as: Mastigoteuthis cordiformis Chun, 1908. Andaman Sea.
Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Family Octopoteuthidae
Octopoteuthis megaptera (Verrill, 1885). Andaman Sea. Reported by: Aungtonya et al. (2014).
Octopoteuthis sicula Rüppell, 1844. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
Taningia danae Joubin, 1931. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al.
(2019).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Family Ommastrephidae
Hyaloteuthis pelagica (Bosc, 1802). Gulf of Thailand. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Nototodarus hawaiiensis (Berry, 1912). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1995b),
Aungtonya et al. (2014), BEDO (2017d), Nabhitabhata et al. (2019).
Ornithoteuthis volatilis (Sasaki, 1915). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
Sthenoteuthis oualaniensis (Lesson, 1830). Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al.
(1992), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), GBIF (2019a).
Todaropsis eblanae (Ball, 1841). Gulf of Thailand. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Family Onychoteuthidae
Onychoteuthis banksi (Leach, 1817). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
Onykia loennbergii (Ishikawa & Wakiya, 1914). Gulf of Thailand. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Walvisteuthis virilis Nesis & Nikitina, 1986. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Family Pyroteuthidae
Pterygioteuthis giardi Fischer, 1896. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
Pyroteuthis margaritifera (Rüppell, 1844). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Family Thysanoteuthidae
Thysanoteuthis rhombus Troschel, 1857. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana & Hylleberg
(1989), Chotiyaputta et al. (1992), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Order Sepiida
Family Sepiadariidae
Sepiadarium kochii Steenstrup, 1881. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
Chaitiamwong (1993), Nabhitabhata et al. (2009a, 2019), BEDO (2017d), GBIF (2019b).
Family Sepiidae
Metasepia tullbergi (Appellöf, 1886). Gulf of Thailand. Reported by: Jereb & Roper (2005), Nateewathana
(2008), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Sepia aculeata Van Hasselt [in Férussac & d’Orbigny], 1835. Reported as: Sepia (Acanthosepion) aculeata
Ferussac & d’Orbigny, 1835; and Sepia aculeata recurvirostra Steenstrup. Gulf of Thailand, Andaman Sea.
Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Chaitiamwong (1993), Nateewathana
(1996, 2008), Voss et al. (1998), Jereb & Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al.
(2014), BEDO (2017d), GBIF (2019a, 2019b).
Sepia arabica Massy, 1916. Reported as: Sepia (Doratosepion) arabica Massy, 1916. Andaman Sea. Reported by:
Nateewathana (1996), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Sepia brevimana Steenstrup, 1875. Reported as: Sepia (Acanthosepion) brevimana Steenstrup, 1875. Gulf of
Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993), Nateewathana (1996,
2008), Voss et al. (1998), Jereb & Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014),
BEDO (2017d).
Sepia esculenta Hoyle, 1885. Reported as: Sepia (Acanthosepion) esculenta Hoyle, 1885. Gulf of Thailand, Andaman
Sea. Reported by: Voss et al. (1998), Nabhitabhata et al. (2009a, 2019), BEDO (2017d), GBIF (2019a).
Sepia kobiensis Hoyle, 1885. Reported as: Sepia (Doratosepion) kobiensis Hoyle, 1885. Gulf of Thailand, Andaman
Sea. Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993), Jereb & Roper (2005), Nabhitabhata et al.
(2009a, 2019), BEDO (2017d).
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The Marine and Estuarine Molluscs of Thailand
Sepia latimanus Quoy & Gaimard, 1832. Reported as: Sepia (Sepia) latimanus (Quoy & Gaimard, 1832). Gulf of
Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d), GBIF (2019a).
Sepia lycidas Gray, 1849. Reported as: Sepia (Acanthosepion) lycidas Gray, 1849. Gulf of Thailand, Andaman Sea.
Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993), Voss et al. (1998), Jereb & Roper (2005),
Nateewathana (2008), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Sepia pharaonis Ehrenberg, 1831. Reported as: Sepia (Sepia) pharaonis Ehrenberg, 1831. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Chaitiamwong (1993),
Nateewathana (1996), Jereb & Roper (2005), Duangdee et al. (2009), Nabhitabhata et al. (2009a, 2019), Hylleberg
& Aungtonya (2013), Aungtonya et al. (2014), BEDO (2017d), Sanpanich & Duangdee (2018), GBIF (2019).
Sepia prashadi Winckworth, 1936. Reported as: Sepia (Acanthosepion) prashadi Winckworth, 1936. Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana (1996, 2008), Nabhitabhata et al. (2009a, 2019),
Aungtonya et al. (2014), BEDO (2017d).
Sepia recurvirostra Steenstrup, 1875. Reported as: Sepia (Acanthosepion) recurvisostra Steenstrup, 1875. Gulf
of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993), Nateewathana
(1996, 2008), Voss et al. (1998), Jereb & Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al.
(2014), BEDO (2017d), GBIF (2019a).
Sepia stellifera Homenko & Khromov, 1916. Reported as: Sepia (Acanthosepion) stellifera Homenko & Khromov,
1916. Gulf of Thailand, Andaman Sea. Reported by: Voss et al. (1998), Nabhitabhata et al. (2009a, 2019), BEDO
(2017d).
Sepiella inermis (Van Hasselt [in Férussac & d’Orbigny], 1835). Reported as: Sepia inermis Ferussac &
d’Orbigny, 1835. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993),
Nateewathana (1996, 2008), Jereb & Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014),
BEDO (2017d), GBIF (2019a).
Family Sepiolidae
Euprymna berryi Sasaki, 1929. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
Chaitiamwong (1993), Nabhitabhata et al. (2009a, 2019), BEDO (2017d), GBIF (2019a).
Euprymna hoylei Adam, 1986. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2019).
Euprymna hyllebergi Nateewathana, 1997. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana
(1997d), Jereb & Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2011, 2014), Aungtonya &
Sungjeen (2014), BEDO (2017d), GBIF (2019a).
Euprymna morsei (Verrill, 1881). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019),
Aungtonya et al. (2014), BEDO (2017d), GBIF (2019b).
Euprymna stenodactyla (Grant, 1833). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Chotiyaputta et al. (1992), Chaitiamwong (1993), Norman & Lu (1997), Nabhitabhata et al. (2009 a, 2019),
Aungtonya et al. (2014), BEDO (2017d).
Inioteuthis maculosa Goodrich, 1896. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
Chaitiamwong (1993), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Sepiola trirostrata Voss, 1962. Gulf of Thailand. Chotiyaputta et al. (1992), Chaitiamwong (1993), Nabhitabhata
et al. (2009a, 2019), BEDO (2017d).
Order Spirulida
Family Spirulidae
Spirula spirula (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019),
BEDO (2017d), GBIF (2019a).
Order [unassigned] Decapodiformes
Superfamily Bathyteuthoidea
Family Chtenopterygidae
Chtenopteryx sicula (Vérany, 1851). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
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Family Idiosepiidae
Idiosepius minimus (d’Orbigny [in Férussac & d’Orbigny], 1835). Reported as: Idiosepius biserialis Voss, 1962.
Gulf of Thailand, Andaman Sea. Reported by: Hylleberg & Nateewathana (1991b), Chaitiamwong (1993), Jereb
& Roper (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019b).
Idiosepius pygmaeus Steenstrup, 1881. Gulf of Thailand, Andaman Sea. Reported by: Hylleberg & Nateewathana
(1991a), Chotiyaputta et al. (1992), Suwanmala et al. (2006), Nabhitabhata et al. (2009a, 2019), Hylleberg &
Aungtonya (2013), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a, 2019b).
Idiosepius thailandicus Chotiyaputta, Okutani & Chaitiamvong, 1991. Gulf of Thailand, Andaman Sea.
Reported by: Chotiyaputta et al. (1992), Chaitiamwong (1993), Jereb & Roper (2005), Nabhitabhata et al.
(2009a, 2019), Aungtonya et al. (2014), Aungtonya & Sungjeen (2014), BEDO (2017d), GBIF (2019b).
Superorder Octopodiformes
Order Octopoda
Suborder Incirrata
Superfamily Argonautoidea
Family Alloposidae
Haliphron atlanticus Steenstrup, 1861. Reported as: Alloposus mollis Verrill, 1880. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana (1995b, 1997b), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019b),
Nabhitabhata et al. (2019).
Family Argonautidae
Argonauta argo Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009b, 2019),
Aungtonya et al. (2014), BEDO (2017d).
Argonauta hians Solander, 1786. Also reported as: Argonauta boettgeri Maltzan, 1881. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992), Nateewathana (1997b),
Sukhsangchan & Nabhitabhata (2007), Sukhsangchan et al. (2008, 2009), Aungtonya et al. (2014), BEDO (2017b),
Nabhitabhata et al. (2019).
Family Tremoctopodidae
Tremoctopus gracilis (Souleyet, 1852). Reported as: Tremoctopus violaceus cf. gracilis (Eydoux & Souleyet, 1852)
and Tremoctopus violaceus Delle Chiaie, 1830. Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2009b,
2019), Aungtonya et al. (2014), BEDO (2017d).
Superfamily Octopodoidea
Family Amphitretidae
Amphitretus pelagicus Hoyle, 1885. Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Bolitaena pygmaea (A. E. Verrill, 1884). Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Japetella diaphana Hoyle, 1885. Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
Vitreledonella richardi Joubin, 1918. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Family Octopodidae
Abdopus aculeatus (d’Orbigny [in Férussac & d’Orbigny], 1834). Gulf of Thailand. Reported by: BEDO (2017d),
Nabhitabhata et al. (2019).
Abdopus horridus (Orbigny, 1826). Gulf of Thailand. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO
(2017d).
Amphioctopus aegina (Gray, 1849). Also reported as: Amphioctopus dollfusi (Robson, 1928); and Octopus dolifusi
Robson, 1928. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981, 1997b), Chotiyaputta
et al. (1992), Norman & Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Hylleberg & Aungtonya (2013),
Aungtonya et al. (2014), Jereb et al. (2014), BEDO (2017d), GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Amphioctopus exannulatus (Norman, 1993). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1997b),
Norman & Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Amphioctopus marginatus (Taki, 1964). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1997b),
Norman & Hochberg (2005), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a), Nabhitabhata et al.
(2019).
Amphioctopus membranaceus (Quoy & Gaimard, 1832). Also reported as: Octopus membranaceus (Quoy &
Gaimard). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Chotiyaputta et al. (1992),
Chaitiamwong (1993), Norman & Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014),
BEDO (2017d).
Amphioctopus neglectus (Nateewathana & Norman, 1999). Gulf of Thailand, Andaman Sea. Reported by:
Nateewathana (1997b), Nateewathana & Norman (1999), Norman & Hochberg (2005), Nabhitabhata et al.
(2009a, 2019), Aungtonya & Sungjeen (2014), Jereb et al. (2014), BEDO (2017d).
Amphioctopus rex (Nateewathana & Norman, 1999). Gulf of Thailand, Andaman Sea. Reported by:
Nateewathana (1997b), Norman & Hochberg (1999, 2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al.
(2014), Aungtonya & Sungjeen (2014), Jereb et al. (2014), BEDO (2017d).
Amphioctopus siamensis (Nateewathana & Norman, 1999). Gulf of Thailand, Andaman Sea. Reported by:
Nateewathana (1997b), Norman & Hochberg (1999, 2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al.
(2014), Aungtonya & Sungjeen (2014), Jereb et al. (2014), BEDO (2017d).
Callistoctopus aspilosomatis (Norman, 1992). Gulf of Thailand. Reported by: Nabhitabhata et al. (2019).
Callistoctopus graptus Norman, 1993. Andaman Sea. Reported by: Norman & Hochberg (2005), Aungtonya
et al. (2014).
Callistoctopus luteus (Sasaki, 1929). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1997b),
Norman & Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), Jereb et al. (2014),
BEDO (2017d).
Callistoctopus macropus (Risso, 1826). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata
et al. (2019).
Callistoctopus ornatus (Gould, 1852). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata et al. (2009a,
2019), BEDO (2017d).
Cistopus chinensis Zheng, Lin, Lu & Ma, 2012. Andaman Sea. Reported by: Nabhitabhata et al. (2019).
Cistopus indicus (Rapp [in Férussac & d’Orbigny], 1835). Gulf of Thailand, Andaman Sea. Reported by:
Chotiyaputta et al. (1992), Chaitiamwong (1993), Nateewathana (1997b), Norman & Hochberg (2005),
Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019a).
Cistopus taiwanicus Liao & Lu, 2009. Andaman Sea. Reported by: Nabhitabhata et al. (2019).
Hapalochlaena maculosa (Hoyle, 1885). Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
Chaitiamwong (1993), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d), GBIF (2019b).
Octopus cyanea Gray, 1849. Gulf of Thailand, Andaman Sea. Reported by: Chotiyaputta et al. (1992),
Nateewathana (1997b), Norman & Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014),
BEDO (2017d), GBIF (2019b).
Octopus globosus Appellöf, 1886. Andaman Sea. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Reported by WoRMS Editorial Board (2019) as a taxon inquirendum.
Octopus kaurna Stranks, 1990. Gulf of Thailand. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Octopus microphthalmus Goodrich, 1896. Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019a, 2019b).
Octopus niveus Lesson, 1830. Andaman Sea. Reported by: Nateewathana (1997b), Norman & Hochberg (2005),
Nabhitabhata et al. (2009a, 2019), Hylleberg & Aungtonya (2013), Aungtonya et al. (2014), BEDO (2017d).
Reported by WoRMS Editorial Board (2019) to be a taxon inquirendum.
Octopus parvus (Sasaki, 1917). Gulf of Thailand. Reported by: Nabhitabhata et al. (2009a, 2019), BEDO (2017d).
Octopus vulgaris Cuvier, 1797. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana (1997b), Norman &
Hochberg (2005), Nabhitabhata et al. (2009a, 2019), Aungtonya et al. (2014), BEDO (2017d).
Thaumoctopus mimicus Norman & Hochberg, 2005. Gulf of Thailand, Andaman Sea. Reported by: Norman &
Hochberg (2005), BEDO (2017d), Nabhitabhata et al. (2019).
Order Vampyromorpha
Family Vampyroteuthidae
Vampyroteuthis infernalis Chun, 1903. Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al. (2019).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Class Gastropoda
Subclass Caenogastropoda
Order Littorinimorpha
Superfamily Calyptraeoidea
Family Calyptraeidae
Calyptraea chinensis (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University
Museum of Fisheries (Natural History) (KUMF), Duangdee (2007), Duangdee et al. (2007, 2011), Nabhitabhata
(2009), Sanpanich & Duangdee (2013).
Calyptraea pellucida (Reeve, 1859). Gulf of Thailand. Also reported as: Calyptraea (Calyptraea) pellucida (Reeve,
1859). Reported by: Gemert (2003), Robba et al. (2004).
Desmaulus extinctorium (Lamarck, 1822). Reported as: Calyptraea extinctorium Lamarck, 1822; Crucibulum
extinctorium Lamarck, 1822; Calyptraea morbida (Reeve, 1859); and Calyptraea marbid (Reeve). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Kurozumi et al. (1989), Swennen et al. (2001),
Gemert (2003), Robba et al. (2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Ergaea walshi (Reeve, 1859). Also reported as: Siphopatella walshi (Reeve, 1859) and Crepidula walshi Reeve,
1859. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Bussarawit (1995), Swennen et al. (2001), Gemert (2003), Robba et al. (2004), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2013), Duangdee et al. (2011), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a,
2019b), J. D. Taylor et al. (unpublished data).
Sigapatella calyptraeformis (Lamarck, 1822). Reported as: Calyptraea calyptraeformis (Lamarck, 1822).
Reported by: KUMF.
Superfamily Capuloidea
Family Capulidae
Capulus japonicus A. Adams, 1861. Andaman Sea. Reported by: Wells (2005), Duangdee et al. (2013b).
Separatista helicoides (Gmelin, 1791). Andaman Sea. Reported by: Nabhitabhata (2009).
Superfamily Cypraeoidea
Family Cypraeidae
Arestorides argus (Linnaeus, 1758). Also reported as: Arestorides argus argus (Linnaeus, 1758); Arestorides
argus contracasta Lorenz, 2012; Cypraea argus Linnaeus, 1758; and Lyncina argus (Linnaeus). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Schilder (1965), Bech (1996), Duangdee (2004), Wells (2005),
Kasetsart University (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011,
2013b), Sanpanich & Duangdee (2013), BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b).
Bistolida hirundo (Linnaeus, 1758). Also reported as: Bistolida hirundo hirundo (Linnaeus, 1758); and Cypraea
hirundo Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Gemert (2003), Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2013), Duangdee et al. (2011), BEDO (2017b), Lorenz (2018), GBIF (2019a).
Bistolida kieneri (Hidalgo, 1906). Also reported as: Cypraea kieneri Hidalgo, 1906. Gulf of Thailand, Andaman
Sea. Reported by: KUMF, Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a).
Bistolida stolida (Linnaeus, 1758). Also reported as: Cypraea stolida (Linnaeus, 1758); and Bistolida stolida
rubiginosa (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Schilder (1965), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), Lorenz (2018). GBIF
(2019a) lists two subspecies from the Andaman Sea: Bistolida kieneri depriesteri Schilder, 1933; and Bistolida
stolida rubiginosa (Gmelin, 1791). WoRMS Editorial Board (2019) regards both subspecies as valid.
Bistolida ursellus (Gmelin, 1791). Also reported as: Bistolida ursellus jomi Meyer & Lorenz, 2017; and Cypraea
ursellus Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), Lorenz (2018), GBIF (2019a, 2019b).
Blasicrura interrupta (J. E. Gray, 1824). Reported as: Cypraea interrupta (Gray, 1824); and Bistolida interrupta
(Gray, 1824). Gulf of Thailand, Andaman Sea. Reported by: Schilder (1965), Heiman (2010), BEDO (2017b),
Lorenz (2018), GBIF (2019a, 2019b).
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The Marine and Estuarine Molluscs of Thailand
Blasicrura pallidula (Gaskoin, 1849). Also reported as: Blasicrura pallidula (Gaskoin, 1849). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Callistocypraea leucodon (Broderip, 1828). Reported as: Callistocypraea leucodon leucodon (Broderip, 1828).
Andaman Sea. Reported by: Lorenz (1999), BEDO (2017b), GBIF (2019a). Two subspecies have been recorded
from Thailand, Callistocypraea leucodon leucodon (Broderip, 1828) and Callistocypraea leucodon tenuidon
(Lorenz, 1999).
Callistocypraea nivosa (Broderip, 1827). Reported as: Cypraea nivosa (Broderip, 1837); and Lyncina nivosa
(Broderip). Andaman Sea. Reported by: KUMF, Schilder (1965), Nabhitabhata (2009), BEDO (2017b), Lorenz
(2018), GBIF (2019a).
Chelycypraea testudinaria (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017b), GBIF
(2019a, 2019b).
Contradusta walkeri (G. B. Sowerby I, 1832). Reported as: Contradusta walkeri walkeri (G. B. Sowerby I, 1832);
Erronea walkeri (Sowerby); Erronea walkeri walkeri (G. B. Sowerby I, 1832); and Cypraea walkeri (Sowerby,
1832). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Schilder (1965), Gemert (2003), Lorenz (2018),
GBIF (2019a).
Cribrarula cribraria (Linnaeus, 1758). Also reported as: Cribraria cribraria (Linnaeus); Cypraea cribraria
Linnaeus, 1758; and Cypraea cribraria comma (Perry, 1811). Gulf of Thailand, Andaman Sea. Reported by:
THNHM, KUMF, Schilder (1965), Wells (2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008),
Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), GBIF
(2019a, 2019b).
Cypraea tigris Linnaeus, 1758. Also reported as: Cypraea tigris tigris forma hinnulea (Melvill, 1888). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, Schilder (1965), Tantanasiriwong (1978), Nateewathana et al.
(1981), Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1995), Duangdee (2004, 2007), Wells
(2005), Kasetsart University (2006), ONR & EPP (2006), Duangdee (2007), ONEP (2007), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018),
BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b) records the subspecies Cypraea tigris pardalis Shaw, 1794.
Eclogavena quadrimaculata (J. E. Gray, 1824). Also reported as: Blasicrura quadrimaculata (Gray, 1824) and
Cypraea quadrimaculata Gray, 1824. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a).
Erronea adusta (Lamarck, 1810). Reported as: Erronea adusta melanesiae Schilder & Schilder, 1937; and Erronea
adusta persica Schilder & Schilder, 1938. Andaman Sea. Reported by: BEDO (2017b), Lorenz (2018).
Erronea caurica (Linnaeus, 1758). Also reported as: Erronea caurica chrismeyeri Lorenz, 2017; Cypraea caurica;
and Cypraea caurica draccaena corrosa (Gronovius, 1781). Gulf of Thailand, Andaman Sea. Reported by: THNHM,
KUMF, Morlet (1889), Schilder (1965), Nateewathana et al. (1981), Way & Purchon (1981), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), Lorenz (2018), GBIF (2019a,
2019b).
Erronea cylindrica (Born, 1778). Reported as: Cypraea cylindrica Born, 1778; and Erronea cylindrica cylindrica
(Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1937, 1938), Bussarawit (1995), Gemert
(2003), Duangdee (2004), Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Duangdee
et al. (2011).
Erronea errones (Linnaeus, 1758). Also reported as: Erronea errones errones (Linnaeus, 1758); Erronea errones
forma bimaculata J. E. Gray, 1824; and Cypraea errones (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Morlet (1889), Suvatti (1937), Schilder (1965), Tantanasiriwong (1978), Nateewathana
et al. (1981), Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1995), Swennen et al. (2001), Gemert
(2003), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Sanpanich &
Duangdee (2013), BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b).
Erronea onyx (Linnaeus, 1758). Reported as: Cypraea onyx (Linnaeus, 1758); Cypraea onyx succincta Linnaeus;
and Cypraea onyx adusta (Lamarck, 1810). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet
(1889), Suvatti (1938), Schilder (1965), Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit (1995),
Carpenter & Niem (1998), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a, 2019b).
Erronea pallida (J. E. Gray, 1824). Gulf of Thailand, Andaman Sea. Reported by: Schilder (1965).
Erronea pyriformis (Gray, 1824). Reported as: Erronea pyriformis pyriformis (Gray, 1824); and Cypraea pyriformis
(Gray, 1842). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Schilder (1965), Tantanasiriwong (1978),
Nateewathana et al. (1981), Swennen et al. (2001), Gemert (2003), Robba et al. (2007), Nabhitabhata (2009),
BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b).
Erronea xanthodon J. E. Gray in G.B. Sowerby I, 1832. Gulf of Thailand. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Ficadusta pulchella (Swainson, 1823). Reported as: Cypraea pulchella (Swainson, 1832); and Ficadusta pulchella
pericalles (Melvill & Standen, 1904). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nabhitabhata (2009),
BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b).
Ipsa childreni (J. E. Gray, 1825). Andaman Sea. Reported by: Duangdee et al. (2013b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Leporicypraea geographica (Schilder & Schilder, 1933). Gulf of Thailand, Andaman Sea. Reported by: GBIF
(2019a, 2019b).
Leporicypraea mappa (Linnaeus, 1758). Reported as: Cypraea mappa (Linnaeus, 1758); Cypraea mappa
geographica (Schilder & Schilder, 1933); and Leporicypraea geographica geographica (Schilder & Schilder,
1933). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Carpenter & Niem (1998), Duangdee (2004),
Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), BEDO
(2017b), Lorenz (2018), GBIF (2019b).
Luria isabella (Linnaeus, 1758). Also reported as: Cypraea isabella Linnaeus, 1758; and Cypraea isabella
controversa (Gray, 1825). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti
(1938), Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Carpenter & Niem (1998), Wells (2005), Kasetsart University (2006), Thamrongnawasawat
et al. (2008), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF
(2019a, 2019b).
Lyncina carneola (Linnaeus, 1758). Also reported as: Cypraea carneola Linnaeus, 1758; and Cypraea carneola
leviathan (Schilder & Schilder, 1937). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet
(1889), Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Carpenter & Niem (1998), Wells (2005), Kasetsart University (2006), Duangdee et al.
(2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
Lyncina lynx (Linnaeus, 1758). Reported as: Cypraea lynx Linnaeus, 1758. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al. (1981), Way &
Purchon (1981), Bussarawit (1995), Kasetsart University (2000), Wells (2005), Thamrongnawasawat et al.
(2008), Duangdee et al. (2011, 2013b), BEDO (2017b), GBIF (2019a).
Lyncina vitellus (Linnaeus, 1758). Also reported as: Cypraea vitellus. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Suvatti (1938), Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981),
Way & Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Gemert (2003), Duangdee (2004),
Thamrongnawasawat et al. (2008), Duangdee et al. (2013b), Sanpanich & Duangdee (2013), BEDO (2017b),
Sanpanich & Duangdee (2018), GBIF (2019a).
Mauritia arabica (Linnaeus, 1758). Also reported as: Cypraea arabica Linnaeus, 1758; Cypraea arabica asiatica
forma gibba (Coen, 1949); Arabica arabica asiatica; Mauritia arabica asiatica Schilder & Schilder, 1939; and
Mauritia arabica asiatica forma gibba Coen, 1949. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Morlet (1889), Suvatti (1937, 1938), Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981),
Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Swennen et al.
(2001), Gemert (2003), Duangdee (2004), Wells (2005), Kasetsart University (2006), ON R & EPP (2006),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2013, 2018), Hylleberg & Aungtonya (2013), BEDO (2017b), Lorenz (2018), GBIF (2019a,
2019b).
Mauritia depressa (Gray, 1824). Reported as: Cypraea depressa Gray, 1824. Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Wells (2005), BEDO (2017b), GBIF (2019a).
Mauritia eglantina (Duclos, 1833). Reported as: Cypraea eglantina Duclos, 1833. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter &
Niem (1998), Kasetsart University (2000), Wells (2005), Nabhitabhata (2009), BEDO (2017b), GBIF (2019b).
Mauritia mauritiana (Linnaeus, 1758). Also reported as: Cypraea mauritania Linnaeus, 1758. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Schilder (1965), Nateewathana et al. (1981), Way & Purchon
(1981), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004), Wells (2005), Kasetsart University
(2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sørensen (2012),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b), Lorenz (2018), GBIF (2019a, 2019b).
Mauritia scurra (Gmelin, 1791). Reported as: Cypraea scurra (Gmelin, 1791); and Mauritia scurra scurra
(Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Carpenter & Niem (1998), Nabhitabhata
(2009), BEDO (2017b), Lorenz (2018), GBIF (2019a).
Melicerona felina (Gmelin, 1791). Reported as: Cypraea felina (Gmelin, 1791). Andaman Sea. Reported by: KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b).
Melicerona listeri (Gray, 1824). Andaman Sea. Reported by: GBIF (2019a).
Monetaria annulus (Linnaeus, 1758). Also reported as: Cypraea annulus Linnaeus, 1758. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Schilder (1965), Tantanasiriwong (1978),
Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Wells
(2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
Monetaria caputserpentis (Linnaeus, 1758). Also reported as: Cypraea caputserpentis Linnaeus, 1758; and
Erosaria caputserpentis (Linnaeus). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit
(1995), Carpenter & Niem (1998), Swennen et al. (2001), Wells (2005), Kasetsart University (2006), ONR &
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The Marine and Estuarine Molluscs of Thailand
EPP (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich &
Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Monetaria moneta (Linnaeus, 1758). Also reported as: Cypraea moneta Linnaeus, 1758; and Erosaria moneta
(Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Suvatti (1938), Schilder
(1965), Nateewathana et al. (1981), Carpenter & Niem (1998), Gemert (2003), Wells (2005), Kasetsart University
(2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a).
Naria erosa (Linnaeus, 1758). Also reported as: Naria erosa erosa (Linnaeus, 1758); Cypraea erosa Linnaeus,
1758; and Erosaria erosa (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet
(1889), Suvatti (1937, 1938), Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Way &
Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004, 2007), Wells (2005), Kasetsart
University (2006), ONR & EPP (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al.
(2011, 2013b), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), Lorenz
(2018), GBIF (2019a, 2019b).
Naria gangranosa (Dillwyn, 1817). Reported as: Erosaria gangranosa (Dillwyn); Naria gangranosa forma
reentsii (Dunker, 1852); Cypraea gangranosa (Dillwyn, 1817); Cypraea gangranosa gangranosa (Dillwyn, 1817);
and Cypraea gangranosa rentsii (Dunker, 1852). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Schilder (1965), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2013b), BEDO
(2017b), Lorenz (2018), Sanpanich & Duangdee (2018), GBIF (2019a).
Naria helvola (Linnaeus, 1758). Reported as: Cypraea helvola Linnaeus, 1758. Andaman Sea. Reported by: Morlet
(1889), Wells (2005).
Naria labrolineata (Gaskoin, 1848). Reported as: Cypraea labrolineata Gaskoin, 1848. Gulf of Thailand, Andaman
Sea. Reported by: THNHM, Wells (2005).
Naria lamarckii (J. E. Gray, 1825). Reported as: Naria lamarckii forma redimita (Melvill, 1888); Cypraea lamackii
lamackii (Gray, 1825); Erosaria lamarcki (Gray); and Cypraea lamackii redimita (Melvill, 1888). Gulf of
Thailand, Andaman Sea. Reported by: KUMF, THNHM, Morlet (1889), Suvatti (1938), Schilder (1965),
Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Duangdee (2004), Kasetsart
University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Taylor (2009), Sanpanich & Duangdee (2013), BEDO (2017b), Lorenz (2018), GBIF
(2019a, 2019b).
Naria ocellata (Linnaeus, 1758). Andaman Sea. Reported by: GBIF (2019a).
Naria miliaris (Gmelin, 1791). Also reported as: Cypraea miliaris Gmelin, 1791; and Erosaria miliaris (Gmelin,
1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti (1937, 1938), Schilder (1965),
Swennen et al. (2001), Gemert (2003), Robba et al. (2004), Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019b).
Nucleolaria nucleus (Linnaeus, 1758). Also reported as: Cypraea nucleus Linnaeus, 1758; and Nuclearia nucleus
(Linnaeus). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Schilder (1965), Wells (2005),
Kasetsart University (2006), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a).
Ovatipsa chinensis (Gmelin, 1791). Reported as: Cypraea chinensis Gmelin, 1791. Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Ovatipsa coloba (Melvill, 1888). Also reported as: Ovatipsa coloba coloba (Melvill, 1888). Andaman Sea. Reported
by: Lorenz (2018), GBIF (2019a).
Palmadusta asellus (Linnaeus, 1758). Also reported as: Palmadusta asellus asellus (Linnaeus, 1758); and Cypraea
asellus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1937, 1938),
Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Gemert
(2003), Duangdee (2004), Wells (2005), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013), BEDO (2017b), Lorenz (2018), GBIF (2019a).
Palmadusta clandestina (Linnaeus, 1767). Reported as: Palmadusta clandestina clandestina (Linnaeus, 1767);
and Cypraea clandestina (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b), Lorenz (2018),
GBIF (2019a).
Palmadusta contaminata (G. B. Sowerby I, 1832). Also reported as: Cypraea contaminata (Sowerby, 1832). Gulf
of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981), Gemert (2003), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), BEDO (2017b), Lorenz (2018),
GBIF (2019a).
Palmadusta ziczac (Linnaeus, 1758). Reported as: Palmadusta ziczac yzac Meyer & Lorenz, 2017; Cypraea ziczac
Linnaeus, 1758; and Cypraea ziczac undata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by:
KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Wells (2005), Nabhitabhata (2009), BEDO (2017b),
Lorenz (2018), GBIF (2019a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Perisserosa guttata (Gmelin, 1791). Also reported as: Cypraea guttata surinensis (Raybaudi, 1978). Andaman
Sea. Reported by: Thamrongnawasawat et al. (2008), BEDO (2017b), GBIF (2019a).
Purpuradusta fimbriata (Gmelin, 1791). Reported as: Purpuradusta fimbriata fimbriata (Gmelin, 1791); and
Cypraea fimbriata Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Wells (2005),
Nabhitabhata (2009), BEDO (2017b), Lorenz (2018). GBIF (2019a) records the subspecies Purpuradusta
fimbriata marmorata (Schröter, 1804) in Thailand.
Purpuradusta gracilis (Gaskoin, 1849). Also reported as: Purpuradusta gracilis gracilis (Gaskoin, 1849); and
Cypraea gracilis (Gaskoin, 1848). Gulf of Thailand, Andaman Sea. Reported by: Schilder (1965),
Tantanasiriwong (1978), Nateewathana et al. (1981), Gemert (2003), Duangdee (2007), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), GBIF (2019a, 2019b).
Purpuradusta hammondae (Iredale, 1939). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2013).
Purpuradusta microdon (J. E. Gray, 1828). Reported as: Cypraea microdon (Gray, 1828). Andaman Sea. Reported
by: KUMF, Nabhitabhata (2009), GBIF (2019a).
Purpuradusta minoridens (Melvill, 1901). Reported as: Purpuradusta minoridens minoridens (Melvill, 1901);
and Cypraea minoridens (Melvill, 1901). Andaman Sea. Reported by: Wells (2005), GBIF (2019a).
Pustularia bistrinotata Schilder & Schilder, 1937. Andaman Sea. Reported by: GBIF (2019a).
Pustularia cicercula (Linnaeus, 1758). Also reported as: Cypraea cicercula Linnaeus, 1758; Cypraea cicercula
lienardi (Jousseaume, 1874); Pustularia cicercula cicercula (Linnaeus, 1758); and Pustularia cicercula tricornis
(Jousseaume, 1874). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Schilder
(1965), Nateewathana et al. (1981), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b),
Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), Lorenz (2018), GBIF
(2019a).
Pustularia globulus (Linnaeus, 1758). Also reported as: Pustularia globulus globulus (Linnaeus, 1758); Cypraea
globulus Linnaeus, 1758; and Cypraea globulus globulus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Suvatti (1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995),
Wells (2005), Nabhitabhata (2009), Duangdee et al. (2013b), BEDO (2017b), Lorenz (2018), GBIF (2019a).
Ransoniella punctata (Linnaeus, 1771). Reported as: Ransoniella punctata bridgesi Meyer & Lorenz, 2017; and
Cypraea punctata Linnaeus, 1771. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Wells (2005), Duangdee
et al. (2013b), Sanpanich & Duangdee (2018), Lorenz (2018).
Staphylaea limacina (Lamarck, 1810). Andaman Sea. Reported by: KUMF, Wells (2005), Nabhitabhata (2009).
GBIF (2019a) uses the subspecies Staphylaea limacina limacina (Lamarck, 1810), but this subspecies is not
recognised by WoRMS Editorial Board (2019).
Staphylaea staphylaea (Linnaeus, 1758). Also reported as: Cypraea staphylaea (Linnaeus, 1758); and
Staphylaea consobrina (Garrett, 1879). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889),
Schilder (1965), Tantanasiriwong (1978), Nateewathana et al. (1981), Wells (2005), Duangdee et al. (2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Talostolida pellucens (Melvill, 1888). Reported as: Talostolida pellucens pellucens (Melvill, 1888). Reported by:
Lorenz (2018).
Talostolida teres (Gmelin, 1791). Reported as: Cypraea teres Gmelin, 1791. Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Wells (2005), Nabhitabhata (2009), GBIF (2019a).
Talparia talpa (Linnaeus, 1758). Also reported as: Talparia talpa talpa (Linnaeus, 1758); and Cypraea talpa
Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Schilder (1965), Tantanasiriwong
(1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Duangdee (2004), Wells (2005), Kasetsart
University (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013, 2018), BEDO (2017b), Lorenz (2018), GBIF (2019a).
Family Eratoidae
Alaerato mactanica (T. Cossignani & V. Cossignani, 1997). Andaman Sea. Reported by: BEDO (2017e).
Eratoena gemma (Bavay, 1917). Andaman Sea. Reported as: Cypraeerato gemma (Bavey, 1917). Reported by:
BEDO (2017b).
Eratoena sulcifera (Gray in G. B. Sowerby I, 1832). Andaman Sea. Reported by: BEDO (2017b, 2017e).
Proterato pura (Kuroda & Habe, 1971). Andaman Sea. Reported by: BEDO (2017e).
Family Ovulidae
Calcarovula longirostrata (G. B. Sowerby I, 1828). Reported as: Phenacovolva longirostrata (Sowerby, 1828).
Andaman Sea. Reported by: BEDO (2017b).
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The Marine and Estuarine Molluscs of Thailand
Calpurnus verrucosus (Linnaeus, 1758). Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011),
Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b).
Carpiscula galearis C. N. Cate, 1973. Andaman Sea. Reported by: Cate (1973), Azuma (1976), Higo et al. (1999),
Lorenz & Fehse (2009).
Crenavolva traillii (A. Adams, 1855). Reported as: Primovula striatula traillii (Adams, 1855). Gulf of Thailand,
Andaman Sea. Reported by: Robba et al. (2004), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011).
Cuspivolva tigris (Yamamoto, 1971). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Diminovula alabaster (Reeve, 1865). Andaman Sea. Reported by: Thamrongnawasawat et al. (2008).
Diminovula dautzenbergi (F. A. Schilder, 1931). Also reported as: Pseudosimnia (Diminovula) perilla C. N. Cate,
1973. Andaman Sea. Reported by: Cate (1973), Lorenz & Fehse (2009).
Globovula cavanaghi (Iredale, 1931). Andaman Sea. Reported by: GBIF (2019a).
Hiatavolva coarctata (G. B. Sowerby II [in A. Adams & Reeve], 1848). Gulf of Thailand, Andaman Sea. Reported
by: Duangdee (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Kurodavolva wakayamaensis (C. N. Cate & Azuma, 1973). Andaman Sea. Reported by: BEDO (2017b).
Margovula marginata (G. B. Sowerby I, 1828). Reported as: Pseudosimnia margarita (Sowerby, 1828). Andaman
Sea. Reported by: Thamrongnawasawat et al. (2008).
Ovula ovum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Way & Puchon (1981), Bussarawit (1995), Carpenter & Niem (1998), KURDI et al.
(2004), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Pellasimnia angasi (Reeve, 1865). Reported as: Phenacovolva haynesi (G. B. Sowerby III, 1889). Gulf of Thailand,
Andaman Sea. Reported by: Lorenz & Fehse (2009), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Pellasimnia brunneiterma (C. N. Cate, 1969). Reported as: Phenacovolva brunneiterna (Cate, 1969). Andaman
Sea. Reported by: BEDO (2017b), GBIF (2019a).
Pellasimnia improcera (Azuma & C. N. Cate, 1971). Andaman Sea. Reported by: Lorenz & Fehse (2009).
Phenacovolva birostris (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: KUMF, KURDI et al.
(2004), Duangdee (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Galli (2015). Considered
to be a nomen dubium by WoRMS Editorial Board (2019).
Phenacovolva brevirostris (Schumanher, 1817). Gulf of Thailand. Reported by: Duangdee (2007), Nabhitabhata
(2009).
Phenacovolva lahainaensis (C. N. Cate, 1969). Also reported as: Phenacovolva weaveri Cate & Azuma in Cate,
1973. Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Phenacovolva nectarea Iredale, 1930. Reported as: Phenacovolva tokioi (Cate, 1973). Gulf of Thailand, Andaman
Sea. Reported by: Duangdee (2004), Wells (2005), Nabhitabhata (2009).
Phenacovolva parvita C. N. Cate & Azuma in C. N. Cate, 1973. Andaman Sea. Reported by: Lorenz & Fehse (2009).
Phenacovolva rosea (A. Adams, 1854). Also reported as: Phenacovolva rosea schmidi (Fehse & Weise, 1993).
Andaman Sea. Reported by: Kasetsart University (2006), ONR & EPP (2006), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Phenacovolva schmidi Fehse & Wiese, 1993. Andaman Sea. Reported by: Lorenz & Fehse (2009), Galli (2015),
GBIF (2019a).
Phenacovolva subreflexa (G. B. Sowerby II, 1848). Andaman Sea. Reported by: GBIF (2019a).
Prionovolva brevis (G. B. Sowerby I, 1828). Andaman Sea. Reported by: Rosenberg (2010).
Procalpurnus lacteus (Lamarck, 1810). Reported as: Calpurnus lacteus (Lamarck, 1810). Gulf of Thailand,
Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Nabhitabhata (2009).
Prosimnia semperi (Weinkauff, 1881). Andaman Sea. Reported by: Duangdee et al. (2007, 2011), Nabhitabhata
(2009), Sanpanich & Duangdee (2013).
Volva volva (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Carpenter & Niem (1998), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
BEDO (2017b).
Family Triviidae
Cleotrivia globosa (Gray in G. B. Sowerby II, 1832). Also reported as: Cypraea globosa Gray in G. B. Sowerby II,
1832. Andaman Sea. Reported by: Morlet (1889), BEDO (2017b).
Trivellona eos (Roberts, 1913). Andaman Sea. Reported by: Duangdee et al. (2013a).
Trivellona paucicostata (Schepman, 1909). Andaman Sea. Reported by: BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Trivirostra oryza (Lamarck, 1810). Reported as: Trivia oryza. Gulf of Thailand, Andaman Sea. Reported by:
KUMF, Gemert (2003), Wells (2005), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich &
Duangdee (2013), BEDO (2017e).
Trivirostra pellucidula (Gaskoin, 1846). Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich & Duangdee (2013).
Family Velutinidae
Coriocella hibyae (Wellens, 1991). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018).
Coriocella nigra Blainville, 1824. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009, 2011),
Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Superfamily Ficoidea
Family Ficidae
Ficus ficus (Linnaeus, 1758). Also reported as: Pirula ficus (Linnaeus, 1758); and Ficus ficoides (Lamarck,
1822). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978),
Nateewathana et al. (1981), Carpenter & Niem (1998), Kasetsart University (2000), Swennen et al. (2001),
Duangdee et al. (2007), Nabhitabhata (2009), GBIF (2019a, 2019b).
Ficus gracilis (G. B. Sowerby I, 1825). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Carpenter & Niem (1998), Duangdee et al. (2009, 2011), Nabhitabhata (2009),
Sanpanich & Duangdee (2013), GBIF (2019a).
Ficus variegata (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Robba et al.
(2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Superfamily Littorinoidea
Family Littorinidae
Echinolittorina leucosticta (Philippi, 1847). Reported as: Nodilittorina leucosticta. Andaman Sea. Reported by:
Sørensen (2012).
Echinolittorina malaccana (Philippi, 1847). Reported as: Echinolittorina trochoides (Gray, 1839). Gulf of
Thailand, Andaman Sea. Reported by: Sanpanich et al. (2004), Reid (2007), Duangdee et al. (2009, 2011),
Sanpanich & Duangdee (2009, 2013, 2018), Dechruksa et al. (2014).
Echinolittorina melanacme (E. A. Smith, 1876). Reported as: Littorina ventricosa. Gulf of Thailand, Andaman
Sea. Reported by: THNHM, Reid (2007), Samakraman et al. (2010).
Echinolittorina pascua (Rosewater, 1970). Reported as: Nodilittorina trochoides (Gray, 1839). Andaman Sea.
Reported by: Gemert (2003), Sørensen (2012), J. D. Taylor et al. (unpublished data).
Echinolittorina radiata (Souleyet, 1852). Reported as: Echinolittorina radiata. Gulf of Thailand. Reported by:
Samakraman et al. (2010).
Echinolittorina reticulata (Anton, 1838). Gulf of Thailand. Reported by: Reid (2001, 2007).
Echinolittorina vidua (Gould, 1839). Also reported as: Nodilittorina vidua (Gould, 1839). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Sanpanich et al. (2004), Wells (2005), Reid (2006), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), Dechruksa et al. (2014), BEDO
(2017b), GBIF (2019a).
Littoraria articulata (Philippi, 1846). Also reported as: Littoraria (Palustorina) articulata (Philippi, 1846). Gulf
of Thailand, Andaman Sea. Reported by: KUMF, Reid (1986), Kurozumi et al. (1989), Gemert (2003), Duangdee
(2004), Sanpanich et al. (2004), Kasetsart University (2006), ONR & EPP (2006), Printrakoon et al. (2008),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), Premcharoen et al. (2016), GBIF
(2019b), J. D. Taylor et al. (unpublished data).
Littoraria bengalensis Reid, 2001. Also reported as: Littorina bengalensis Reid, 2001. Gulf of Thailand, Andaman
Sea. Reported by: Reid (2001a), Sanpanich et al. (2004), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
Galli (2015), J. D. Taylor et al. (unpublished data).
Littoraria carinifera (Menke, 1830). Also reported as: Littorina carinifera (Menke, 1830); and Littorina
(Palustorina) carinifera (Menke, 1830). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet
(1889), Rosewater (1970), Nielsen (1976b), Nateewathana et al. (1981), Way & Purchon (1981), Reid (1986),
Swennen et al. (2001), Gemert (2003), Sanpanich et al. (2004), Printrakoon et al. (2008), Duangdee et al. (2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor
et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Littoraria conica (Philippi, 1846). Andaman Sea. Reported by: Sanpanich et al. (2004), Duangdee et al. (2011),
Sanpanich & Duangdee (2013), J. D. Taylor et al. (unpublished data).
Littoraria intermedia (Philippi, 1846). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Reid (1986),
Robba et al. (2004), Sanpanich et al. (2004), Printrakoon et al. (2008), Duangdee et al. (2011), Sanpanich &
Duangdee (2013), Dechruksa et al. (2014), BEDO (2017b), GBIF (2019a).
Littoraria lutea (Philippi, 1847). Gulf of Thailand. Reported by: GBIF (2019b).
Littoraria melanostoma (Gray, 1839). Also reported as: Littorina melanostoma Gray, 1839. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Rosewater (1970), Reid (1986), Swennen et al.
(2001), Duangdee (2004), Robba et al. (2004), Sanpanich et al. (2004), Sri-aroon et al. (2005), Printrakoon et
al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Premcharoen et al. (2016), Dechruksa et al.
(2014), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Littoraria pallescens (Philippi, 1846). Also reported as: Littorina sieboldii Philippi, 1846. Gulf of Thailand,
Andaman Sea. Reported by: Morlet (1889), Reid (1986), Swennen et al. (2001), Sanpanich et al. (2004),
Printrakoon et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Littoraria scabra (Linnaeus, 1758). Also reported as: Littorina scabra (Linnaeus, 1758) and Littorina scabra
scabra. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Rosewater (1970),
Nielsen (1976b), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Reid (1986),
Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Duangdee (2004, 2007),
Sanpanich et al. (2004), Sri-aroon et al. (2005), ONR & EPP (2006), Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Sørensen (2012), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), Dechruksa et al.
(2014), BEDO (2017b), GBIF (2019a, 2019b).
Littoraria strigata (Philippi, 1846). Also reported as: Littorina strigata (Philippi, 1846); and Littorina (Palustorina)
strigata. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Reid (1986), Gemert (2003), Sanpanich et al.
(2004), Printrakoon et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2013), GBIF (2019a, 2019b).
Littoraria undulata (Gray, 1839). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Rosewater
(1970), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Subba Rao & Dey (2000),
Sanpanich et al. (2004), Sri-aroon et al. (2005, 2006), Wells (2005), Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Sørensen (2012), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
Nodilittorina pyramidalis (Quoy & Gaimard, 1833). Also reported as: Nodilittorina pyramidalis pyramidalis
(Quoy & Gaimard, 1833). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Rosewater (1970), Nateewathana
et al. (1981), Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998),
Kasetsart University (2000, 2006), Subba Rao & Dey (2000), Duangdee (2004), ONR & EPP (2006), Hylleberg
& Aungtonya (2013), BEDO (2017b), GBIF (2019a, 2019b).
Peasiella infracostata (Issel, 1869). Gulf of Thailand. Reported by: Reid (1998).
Peasiella patula Reid, 1998. Gulf of Thailand. Reported by: Reid (1998), Gemert (2003).
Peasiella roepstorffiana (Nevill, 1885). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich et al. (2004),
Duangdee et al. (2009, 2011), Nabhitabhata (2009), Samakraman et al. (2010), Sørensen (2012), Sanpanich &
Duangdee (2013, 2018), BEDO (2017e), J. D. Taylor et al. (unpublished data).
Tectarius antonii (Philippi, 1846). Also reported as: Tectarius (Echininus) nodulosus. Gulf of Thailand. Reported
by: THNHM, Nateewathana et al. (1981), Kostina et al. (2016).
Tectarius pagodus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Superfamily Naticoidea
Family Naticidae
Conuber incei (Philippi, 1853). Reported as: Polinices incei (Philippi, 1853). Andaman Sea. Reported by: Wells
(2005).
Eunaticina papilla (Gmelin, 1791). Also reported as: Eunaticina tumescens Reeve, 1864. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al.
(1981), Swennen et al. (2001), Gemert (2003), Robba et al. (2004), Wells (2005), ONEP (2007), Nabhitabhata
(2009), Duangdee et al. (2011), Torigoe & Inaba (2011), Sawatna et al. (2012), Sanpanich & Duangdee (2013),
BEDO (2017b).
Eunaticina papilla lamarckiana (Récluz, 1843). Reported as: Eunaticina lamarckiana. Gulf of Thailand. Reported
by: THNHM, Morlet (1889).
Mammilla fibrosa (Gray, 1850). Gulf of Thailand, Andaman Sea. Reported by: Torigoe & Inaba (2011), BEDO
(2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Mammilla mammata (Röding, 1798). Also reported as: Polinices mammatus (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009),
Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b).
Mammilla melanostoma (Gmelin, 1791). Reported as: Polinices melanostomus Gmelin, 1791; and Mammilla
opaca (Récluz, 1851). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Carpenter
& Niem (1998), Nabhitabhata (2009), GBIF (2019b).
Mammilla melanostomoides (Quoy & Gaimard, 1832). Also reported as: Polinices melanostomoides (Quoy &
Gaimard). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Torigoe & Inaba (2011),
BEDO (2017b).
Mammilla sebae (Récluz, 1844). Also reported as: Polinices sebae (Récluz, 1844). Gulf of Thailand, Andaman
Sea. Reported by: KUMF, Tantanasiriwong (1978), Carpenter & Niem (1998), Kilburn & Hylleberg (1998),
Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2011), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Torigoe & Inaba (2011), Sanpanich & Duangdee (2013).
Mammilla simiae (Deshayes, 1838). Reported as: Polinices simiae (Deshayes, 1838). Gulf of Thailand, Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Wells (2005), Duangdee et al. (2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Natica arachnoidea (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Kilburn & Hylleberg
(1998), Swennen et al. (2001), Nabhitabhata (2009), Duangdee et al. (2011), Torigoe & Inaba (2011), Sanpanich
& Duangdee (2013), BEDO (2017b).
Natica buriasiensis Récluz, 1844. Gulf of Thailand. Reported by: GBIF (2019b).
Natica fasciata (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et
al. (2011), Sanpanich & Duangdee (2013).
Natica maculosa Lamarck, 1822. Reported as: Natica mauelosa. Gulf of Thailand. Reported by: THNHM, Morlet
(1889).
Natica schepmani Thiele, 1925. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Natica seychellium R. B. Watson, 1886. Reported as: Natica maheensis (Reeve, 1855). Andaman Sea. Reported
by: Kilburn & Hylleberg (1998), Nabhitabhata (2009).
Natica simplex G. B. Sowerby III, 1897. Reported as: Natica simplex (Röding, 1798). Gulf of Thailand. Reported
by: Nabhitabhata (2009).
Natica sinensis X.-T. Ma & S.-P. Zhang, 1993. Reported as: Natica sinensis. Gulf of Thailand. Reported by: THNHM.
Natica spadicea (Gmelin, 1791). Gulf of Thailand. Reported by: GBIF (2019b).
Natica stellata Hedley, 1913. Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004), Sanpanich &
Duangdee (2009, 2013), Nabhitabhata (2009), Duangdee et al. (2011).
Natica vitellus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Carpenter & Niem (1998), Subba Rao & Dey (2000), Swennen et al. (2001), Robba
et al. (2004), Duangdee (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011),
Torigoe & Inaba (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019b).
Naticarius alapapilionis (Röding, 1798). Reported as: Naticarius (Naticarius) alapapilionis (Röding, 1798); and
Naticarius (Glyphepithema) alapapilionis (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM,
Nateewathana et al. (1981), Kilburn & Hylleberg (1998), Gemert (2003), Nabhitabhata (2009), Torigoe & Inaba
(2011), BEDO (2017b).
Naticarius colliei (Récluz, 1844). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Naticarius manceli (Jousseaume, 1874). Reported as: Naticarius (Naticarius) manceli (Jousseaume, 1874). Gulf
of Thailand. Reported by: Gemert (2003).
Naticarius onca (Röding, 1798). Also reported as: Natica chinensis Lamarck, 1816; and Natica onca (Röding,
1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978),
Nateewathana et al. (1981), Swennen et al. (2001), Wells (2005), ONR & EPP (2006), Duangdee et al. (2009,
2011, 2013b), Nabhitabhata (2009), Torigoe & Inaba (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Neverita didyma (Röding, 1798). Reported as: Glossaulax didyma (Röding, 1798); and Polinices didyma. Gulf of
Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981),
Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Kasetsart University (2000), Swennen
et al. (2001), Duangdee (2004), Robba et al. (2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013), Torigoe & Inaba (2011), BEDO (2017b), GBIF (2019b), J. D. Taylor et al.
(unpublished data).
Notocochlis cernica (Jousseaume, 1874). Reported as: Natica cernica (Jousseaume, 1874). Gulf of Thailand,
Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Nabhitabhata (2009), BEDO (2017b).
Notocochlis gualtieriana (Récluz, 1844). Also reported as: Natica gualtieriana (Récluz, 1844); and Natica aff.
acellus Reeve, 1855. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Carpenter & Niem
(1998), Gemert (2003), Robba et al. (2007), Sanpanich & Duangdee (2009, 2013), Nabhitabhata (2009),
Duangdee et al. (2011), BEDO (2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Notocochlis tigrina (Röding, 1798). Also reported as: Natica tigrina (Röding, 1798); Tanea tigrina (Röding,
1798); and Paratectonatica tigrina (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Nateewathana (1995), Carpenter
& Niem (1998), Swennen et al. (2001), Duangdee (2004), Robba et al. (2004), Kasetsart University (2006),
Duangdee et al. (2007, 2009, 2011), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Torigoe & Inaba
(2011), Sanpanich & Duangdee (2013), Galli (2015), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al.
(unpublished data).
Polinices albumen (Linnaeus, 1758). Also reported as: Neverita albumen (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter &
Niem (1998), Subba Rao & Dey (2000), Swennen et al. (2001), Duangdee (2004), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Duangdee et al. (2011), Torigoe & Inaba (2011), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019a, 2019b).
Polinices aurantius (Röding, 1798). Andaman Sea. Reported by: Wells (2005), GBIF (2019a).
Polinices candidissimus (Le Guillou, 1842). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Polinices flemingianus (Récluz, 1844). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Kurozumi et al. (1989), Kasetsart University (2000), Subba Rao & Dey (2000),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF
(2019a).
Polinices mammilla (Linnaeus, 1758). Also reported as: Natica piriformis Récluz, 1843; Polinices tumidus
(Swainson); and Polinices mamilla Lamarck. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889),
Suvatti (1937), Nateewathana et al. (1981), Kurozumi et al. (1989), Carpenter & Niem (1998), Kasetsart
University (2000, 2005, 2007), Subba Rao & Dey (2000), Swennen et al. (2001), Gemert (2003), Duangdee
(2004, 2007), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013b), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Torigoe & Inaba (2011),
Hylleberg & Aungtonya (2013), Kostina et al. (2016), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al.
(unpublished data).
Polinices peselephanti (Link, 1807). Also reported as: Polinices columnaris (Récluz, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Swennen et al. (2001),
Nabhitabhata (2009), Torigoe & Inaba (2011), BEDO (2017b).
Polinices powisianus (Récluz, 1844). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Swennen et al. (2001), Nabhitabhata (2009), Duangdee et al. (2011), Torigoe
& Inaba (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Sigatica pomatiella (Melvill, 1893). Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004), Chaiwathee
et al. (2007).
Sinum eximium (Reeve, 1864). Gulf of Thailand. Reported by: Swennen et al. (2001), Torigoe & Inaba (2011).
Sinum haliotoideum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Also reported as: Sinum (Ectosinum)
planulatum (Recluz, 1844). Reported by: Swennen et al. (2001), Torigoe & Inaba (2011), Sanpanich & Duangdee
(2013).
Sinum incisum (Reeve, 1864). Also reported as: Sinum undulatus. Gulf of Thailand, Andaman Sea. Reported by:
THNHM, ONR & EPP (2006), Nabhitabhata (2009), Duangdee et al. (2011), Torigoe & Inaba (2011), Sanpanich
& Duangdee (2013).
Sinum javanicum (Gray, 1834). Reported as: Sinum javanicum Griffith & Pidgeon, 1834. Gulf of Thailand.
Reported by: KUMF, Torigoe & Inaba (2011).
Sinum neritoideum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti
(1938), Swennen et al. (2001), Robba et al. (2004), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Torigoe & Inaba (2011), BEDO (2017b).
Sinum planulatum (Récluz, 1843). Gulf of Thailand. Reported by: Gemert (2003), Duangdee et al. (2007),
Nabhitabhata (2009).
Tanea areolata (Récluz, 1844). Also reported as: Natica areolata Récluz. Andaman Sea. Reported by: THNHM,
Nateewathana et al. (1981), Kilburn & Hylleberg (1998).
Tanea lineata (Röding, 1798). Also reported as: Natica lineata (Röding, 1798). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Morlet (1889), Suvatti (1937, 1938), Tantanasiriwong (1978), Nateewathana et
al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000), Subba Rao & Dey (2000),
Robba et al. (2004), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Torigoe
& Inaba (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019b).
Tanea mosaica (Sowerby, 1843). Andaman Sea. Reported by: Kilburn & Hylleberg (1998).
Tanea sagittata (Menke, 1843). Also reported as: Notocochlis sagittata (Linnaeus). Andaman Sea. Reported by:
Nateewathana et al. (1981).
Tanea undulata (Röding, 1798). Reported as: Natica euzona (Récluz, 1844); Natica undulata (Röding, 1798);
and Tanea euzona (Récluz, 1844). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Nateewathana et al. (1981), Carpenter & Niem (1998), Kilburn & Hylleberg (1998), Nabhitabhata (2009),
Duangdee et al. (2011), Torigoe & Inaba (2011), Sanpanich & Duangdee (2013, 2018), BEDO (2017b).
Superfamily Rissooidea
Family Barleeiidae
Ansola angustata (Pilsbry, 1901). Reported as: Barleeia angustata (Pilsbry, 1901). Gulf of Thailand. Reported
by: Robba et al. (2004).
Family Rissoidae
Alvania ogasawarana (Pilsbry, 1904). Reported as: Alvania (Alvania) ogasawarana (Pilsbry, 1904). Gulf of
Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Haurakia novarensis (Frauenfeld, 1867). Reported as: Alvania (Alvania) novarensis (Frauenfeld, 1867). Gulf of
Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Cingula matusimana Nomura, 1940. Reported as: Lucidesta matusimana (Nomura, 1940). Gulf of Thailand.
Reported by: Gemert (2003). This species could not be verified through WoRMS Editorial Board (2019), but is
shown on Galli (2015).
Lucidestea mundula (A. Adams, 1860). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO
(2017e).
Parashiela invisibilis (Hedley, 1899). Andaman Sea. Reported by: BEDO (2017e).
Rissoa rosea (Deshayes, 1862). Reported as: Rissoina rosea (Deshayes, 1862). Andaman Sea. Reported by: BEDO
(2017e). WoRMS Editorial Board (2019) states this is a taxon inquirendum and should be placed in the family
Pyramidellidae.
? Rissoella omphalotropis (A. Adams, 1860). Reported as: ? Voorwindia omphalotropis (A. Adams, 1860). Gulf of
Thailand. Reported by: Gemert (2003). This species could not be verified through WoRMS Editorial Board
(2019), but is discussed in Ponder (1984).
Simulamerelina wanawana (Kay, 1979). Andaman Sea. Reported by: BEDO (2017e).
Voorwindia cf. paludinoides (Yokoyama, 1927). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003),
BEDO (2017e). This species could not be verified through WoRMS Editorial Board (2019), but is recorded in
Ganmanee et al. (2003).
Family Rissoinidae
Apataxia cerithiiformis (Tryon, 1887). Reported as: Rissoina cerithiiformis Tryon, 1887. Andaman Sea. Reported
by: BEDO (2017e).
Moerchiella antoni Schwartz, 1860. Reported as: Rissoina (Moerchiella) antoni Schwartz, 1860. Gulf of Thailand.
Reported by: Gemert (2003).
Moerchiella dorbignyi (A. Adams, 1853). No Thai locality provided. Reported by: Subba Rao & Dey (2000).
Moerchiella cf. okinawensis Sleurs, 1993. Reported as: Rissoina (Moerchiella) cf. okinawensis Sleurs, 1993. Gulf
of Thailand. Reported by: Gemert (2003).
Moerchiella cf. artensis (Montrouzier in Souverbie & Montrouzier, 1872). Reported as: Rissoina cf. artensis
Montrouzier in Souverbie and Montrouzier, 1872. Andaman Sea. Reported by: BEDO (2017e).
Phosinella clathrata (A. Adams, 1853). Reported as: Rissoina (Phosinella) clathrata Adams, 1851. Gulf of
Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Phosinella dunkerina (Kuroda & Habe in Habe, 1961). Gulf of Thailand, Andaman Sea. Reported by: BEDO
(2017e).
Phosinella pura (Gould, 1861). Reported as: Rissoina (Phosinella) pura (Gould, 1861). Gulf of Thailand. Reported
by: Gemert (2003).
Rissoina ambigua (Gould, 1849). Also reported as: Rissoina (Rissoina) materinsulae Pilsbry, 1904. Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Gemert (2003), Kasetsart University (2006),
Duangdee et al. (2011), BEDO (2017e).
Rissoina costata A. Adams, 1851. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Rissoina cf. costulata (Dunker, 1860). Andaman Sea. Reported by: BEDO (2017e). Listed by WoRMS Editorial
Board (2019) as a nomen dubium.
Rissoina fimbriata Souverbie, 1872. Reported as: Rissoina (Rissolina) fimbriata Souverbie, 1872. Gulf of Thailand.
Reported by: Gemert (2003).
Rissoina plicatula Gould, 1861. Reported as: Rissoina (Rissolina) plicatula Gould, 1861. Gulf of Thailand. Reported
by: Gemert (2003).
Rissoina scolopax Souverbie, 1877. Reported as: Rissoina (Rissolina) scolopax Souverbie, 1877. Gulf of Thailand.
Reported by: Gemert (2003).
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The Marine and Estuarine Molluscs of Thailand
Rissoina tornatilis Gould, 1861. Reported as: Rissoina (Phosinella) tornatilis Gould, 1861. Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Rissoina cf. vangoethemorum Sleurs, 1994. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Sulcorissoina imbricata (A. A. Gould, 1861). Reported as Rissoina cf. imbricata (Gould, 1861). Andaman Sea.
Reported by: BEDO (2017e).
Family Zebinidae
Pandalosia ephamilla (Watson, 1886). Andaman Sea. Reported by: BEDO (2017e).
Pandalosia subulata Laseron, 1956. Reported as: Schwartziella (Pandalosia) subulata (Laseron, 1956). Gulf of
Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Stosicia annulata (Dunker, 1860). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004), BEDO (2017e).
Zebina pupiniformis (Preston, 1908). Andaman Sea. Reported by: BEDO (2017e).
Superfamily Stromboidea
Family Rostellariidae
Rimellopsis powisii Petit de la Saussaye, 1840. Reported as: Rostellaria powisii Petit de la Saussaye, 1840.
Andaman Sea. Reported by: Roussy (1991a), BEDO (2017b).
Rostellariella delicatula (G. Nevill, 1881). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991a), GBIF
(2019a, 2019b).
Tibia fusus (Linnaeus, 1758). Also reported as: Tibia fusus fusus (Linnaeus, 1758). Gulf of Thailand. Reported by:
THNHM, KUMF, Roussy (1991a), Carpenter & Niem (1998), Swennen et al. (2001), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), GBIF (2019b).
Varicospira cancellata (Lamarck, 1816). Also reported as: Rimella cancellata (Lamarck, 1816). Andaman Sea.
Reported by: Nateewathana et al. (1981), Roussy (1991), BEDO (2017b).
Family Seraphsidae
Terebellum terebellum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Jung
& Abbott (1967), Nateewathana et al. (1981), Roussy (1991a), Bussarawit (1995), Swennen et al. (2001),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013,
2018), BEDO (2017b).
Family Strombidae
Canarium betuleti (Kronenberg, 1991). Also reported as: Strombus ochroglottis betuleti. Andaman Sea. Reported
by: Dekker (1996), GBIF (2019a).
Canarium erythrinum (Dillwyn, 1817). Also reported as: Strombus erythrinus (Dillwyn, 1817) and Strombus
erythrinus erythinus (Dillwyn). Gulf of Thailand, Andaman Sea. Reported by: Abbott (1960), Tantanasiriwong
(1978), Nateewathana et al. (1981), Roussy (1991a), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013), BEDO (2017b).
Canarium labiatum (Röding, 1798). Reported as: Strombus labiatus (Röding, 1798) and Strombus labiatus
olydius (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Roussy (1991a), Subba Rao & Dey (2000), Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Sanpanich & Duangdee (2013), GBIF (2019a).
Canarium microurceus Kira, 1959. Reported as: Strombus microurceus (Kira, 1959). Andaman Sea. Reported by:
Roussy (1991a), Wells (2005).
Canarium mutabile (Swainson, 1821). Reported as: Strombus floridus Lamarck, 1822; and Strombus mutabilis
(Swainson, 1821). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Tantanasiriwong (1978),
Nateewathana et al. (1981), Roussy (1991a), Carpenter & Niem (1998), BEDO (2017b).
Canarium olydium (Duclos, 1844). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019a).
Canarium urceus (Linnaeus, 1758). Also reported as: Strombus urceus (Linnaeus, 1758); Strombus urceus form
ustulatus Schumacher; Strombus (Canarium) urceus f. ustulatus Schumacher, 1817; Strombus ustulatus; and
Canarium urceum (L.). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti
(1937, 1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Kurozumi et al.
(1989), Roussy (1991a), Bussarawit (1995), Kasetsart University (2000, 2006), Swennen et al. (2001), Gemert
(2003), Duangdee (2004, 2007), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2011), ONEP (2007),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Hylleberg
& Aungtonya (2013), BEDO (2017b), GBIF (2019a, 2019b).
Canarium wilsonorum (Abbott, 1967). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
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Conomurex decorus (Röding, 1798). Reported as: Strombus decorus (Röding, 1798); and Strombus decorus decorus
(Roeding). Gulf of Thailand, Andaman Sea. Reported by: Abbott (1960), Tantanasiriwong (1978), Nateewathana
et al. (1981), Way & Purchon (1981), Amornjaruchit (1988), Roussy (1991a), Bussarawit (1995), Wells (2005),
Kronenberg et al. (2009), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Conomurex luhuanus (Linnaeus, 1758). Also reported as: Strombus luhuanus (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti (1937, 1938), Kurozumi et al. (1989), Roussy
(1991a), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004, 2007), Kasetsart University (2006),
ONR & EPP (2006), Duangdee et al. (2007, 2009), ONEP (2007), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011, 2013b), GBIF (2019b).
Dolomena columba (Lamarck, 1822). Reported as: Strombus plicatus columba (Lamarck, 1822). Andaman Sea.
Reported by: KUMF, Nateewathana et al. (1981), Bussarawit (1995), BEDO (2017b).
Dolamena dilatatus (Swainson, 1821). Reported as: Strombus dilatatus Swainson, 1821. Gulf of Thailand,
Andaman Sea. Reported by: Swennen et al. (2001), Nabhitabhata (2009), GBIF (2019a).
Dolomena hickeyi (Willan, 2000). Gulf of Thailand, Andaman Sea. Reported by: Kronenberg & Dharma (2005).
Dolomena labiosa (Wood, 1828). Reported as: Strombus labiosus (Wood, 1828). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Roussy (1991a), Aungtonya & Hylleberg (1998), Swennen
et al. (2001), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Dolomena plicata (Röding, 1798). Also reported as: Strombus sibbaldi G. B. Sowerby II, 1842, which WoRMS
Editorial Board (2019) classifies as a subspecies of Dolomena plicata. Gulf of Thailand, Andaman Sea. Reported
by: Morlet (1889), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Dolomena pulchella (Reeve, 1851). Also reported as: Strombus plicatus pulchellus (Reeve, 1851). Gulf of Thailand,
Andaman Sea. Reported by: Roussy (1991a), Aungtonya & Hylleberg (1998), Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Dolomena variabilis (Swainson, 1820). Also reported as: Strombus variabilis (Swainson, 1820). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Abbott (1960), Roussy ( 1991a), Swennen et al.
(2001), Wells (2005), Duangdee (2007), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Sanpanich &
Duangdee (2013), GBIF (2019b). GBIF (2019a) recognises the Thai subspecies as Dolomena variabilis variabilis
(Swainson, 1820).
Doxander vittatus (Linnaeus, 1758). Reported as: Strombus (Doxander) vittatus vittatus Linnaeus, 1758;
Strombus vittatus vittatus; Strombus Doxander vittatus apicatus (Man in 't Veld & Visser, 1993); and Doxander
vittatus apicatus (Man in 't Veld & Visser, 1993). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889),
Nateewathana et al. (1981), Kronenberg (1998), Subba Rao & Dey (2000), Gemert (2003), Robba et al. (2007),
Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
Euprotomus aratrum (Röding, 1798). Reported as: Strombus aratrum (Röding, 1798). Gulf of Thailand, Andaman
Sea. Reported by: Roussy (1991a), Swennen et al. (2001), Nabhitabhata (2009), GBIF (2019b).
Euprotomus aurisdianae (Linnaeus, 1758). Reported as: Strombus aurisdianae (Linnaeus, 1758). Andaman Sea.
Reported by: Nateewathana et al. (1981), Roussy (1991a), Bussarawit (1995), Thamrongnawasawat et al. (2008),
BEDO (2017b), GBIF (2019b).
Euprotomus bulla (Röding, 1798). Reported as: Strombus bulla (Röding, 1798). Andaman Sea. Reported by:
THNHM, Roussy (1991a).
Euprotomus vomer (Röding, 1798). Gulf of Thailand. Reported by: GBIF (2019b).
Gibberulus gibberulus (Linnaeus, 1758). Also reported as: Strombus gibberulus albus (Mörch, 1850); Strombus
gibberulus gibberulus; and Strombus gibberulus gibbosus (Roeding). Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Bussarawit (1995), Nabhitabhata (2009), Duangdee et al. (2013b), BEDO (2017b).
Harpago chiragra (Linnaeus, 1758). Also reported as: Lambis chiragra (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Abbott (1961), Tantanasiriwong (1978), Nateewathana et al. (1981),
Way & Purchon (1981), Roussy (1991a), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao & Dey (2000),
Duangdee (2004), Wells (2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b).
Labiostrombus epidromis (Linnaeus, 1758). Reported as: Strombus epidromis (Linnaeus, 1758). Gulf of Thailand.
Reported by: Roussy (1991a), Carpenter & Niem (1998), Swennen et al. (2001), Nabhitabhata (2009), GBIF
(2019b).
Laevistrombus canarium (Linnaeus, 1758). Also reported as: Strombus canarium (Linnaeus, 1758); and Strombus
(Laevistrombus) canarium (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Suvatti (1937, 1938), Abbott (1960), Tantanasiriwong (1978), Nateewathana et al. (1981), Amornjaruchit (1988),
Nateewathana (1995), Carpenter & Niem (1998), Kasetsart University (2000), Gemert (2003), Duangdee
(2004, 2007), Duangdee et al. (2007), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Hylleberg & Aungtonya (2013), GBIF (2019b), J. D.
Taylor et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Laevistrombus turturella (Röding, 1798). Reported as: Strombus isabella Lamarck, 1822; and Strombus
(Laevistrombus) turturella Röding, 1798. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Kurozumi
et al. (1989), Man in 't Veld & Turck (1998), Gemert (2003).
Lambis crocata (Link, 1807). Gulf of Thailand, Andaman Sea. Reported by: Roussy (1991a), Carpenter & Niem
(1998), Nabhitabhata (2009).
Lambis lambis (Linnaeus, 1758). Also reported as: Pterocera lambis (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al. (1981), Roussy
(1991a), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao & Dey (2000), Duangdee (2004), Kasetsart
University (2006), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013,
2018), BEDO (2017b), GBIF (2019a).
Lambis millepeda (Linnaeus, 1758). Gulf of Thailand. Reported by: Bussarawit (1995), Carpenter & Niem (1998),
Nabhitabhata (2009).
Lambis scorpius (Linnaeus, 1758). Also reported as: Lambis scorpius indomaris Abbott, 1961; and Strombus
scorpius. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana
et al. (1981), Way & Purchon (1981), Roussy (1991a), Bussarawit (1995), Carpenter & Niem (1998), Wells
(2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al.
(2011, 2013a, 2013b), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b),
GBIF (2019a).
Lambis truncata (Kiener, 1843). Also reported as: Lambis truncata truncata (Kiener, 1843). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Roussy (1991a), Bussarawit
(1995), Carpenter & Niem (1998), Wells (2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b).
Lentigo lentiginosus (Linnaeus, 1758). Also reported as: Strombus lentiginosus (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti (1938), Tantanasiriwong (1978), Nateewathana et al.
(1981), Roussy (1991a), Bussarawit (1995), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Lentigo pipus (Röding, 1798). Reported as: Strombus pipus (Röding,1798). Andaman Sea. Reported by: Roussy
(1991a).
Margistrombus marginatus (Linnaeus, 1758). Also reported as: Strombus marginatus (Linnaeus, 1758); Strombus
(Dolomena) marginatus s.l.; Strombus (Dolomena) marginatus marginatus (Linnaeus, 1758); and Strombus
marginatus robustus (Sowerby, 1874). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti
(1937, 1938), Abbott (1960), Roussy (1991a), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998),
Swennen et al. (2001), Gemert (2003), ONR & EPP (2006), Duangdee (2007), Duangdee et al. (2007, 2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Taylor (2009), Hylleberg & Aungtonya
(2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b). GBIF (201 9b) also recognises
Margistrombus robustus (G. B. Sowerby II, 1875).
Margistrombus septimus (Duclos, 1844). Also reported as: Strombus marginatus septimus Duclos. Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b).
Mirabilistrombus listeri (Gray, 1852). Reported as: Strombus listeri (Gray, 1852). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Roussy (1991a), Bussarawit
(1995), Kronenberg (1998), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), BEDO (2017b), GBIF
(2019a).
Sinustrombus sinuatus ([Lightfoot], 1786). Also reported as: Strombus sinuatus (Humphrey, 1786). Andaman
Sea. Reported by: Roussy (1991a).
Terestrombus terebellatus (G. B. Sowerby II, 1842). Also reported as: Strombus terebellatus terebellatus Sowerby.
Andaman Sea. Reported by: Nateewathana et al. (1981), Roussy (1991a), Bussarawit (1995), Subba Rao & Dey
(2000), Nabhitabhata (2009), Duangdee et al. (2011, 2013a), Sanpanich & Duangdee (2013), BEDO (2017b).
Tridentarius dentatus (Linnaeus, 1758). Reported as: Strombus dentatus (Linnaeus, 1758). Andaman Sea.
Reported by: Morlet (1889), Suvatti (1937, 1938), Roussy (1991a), Carpenter & Niem (1998), Nabhitabhata
(2009).
Superfamily Tonnoidea
Family Bursidae
Bufonaria crumena (Lamarck, 1816). Reported as: Ranella crumena Lamarck, 1816; and Bursa crumena (Lamarck,
1816). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Suvatti (1937, 1938), Carpenter & Niem
(1998), Nabhitabhata (2009).
Bufonaria echinata (Link, 1807). Reported as: Bursa echinata (Link, 1807) and Bursa spinosa (Lamarck).
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Nabhitabhata (2009).
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Bufonaria elegans (G. B. Sowerby II, 1836). Andaman Sea. Reported by: KUMF, Cossignani (1994), Bussarawit
(1995), Beu (2005), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Bufonaria foliata (Broderip, 1826). Gulf of Thailand. Reported by: GBIF (2019b).
Bufonaria margaritula (Deshayes, 1832). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Cossignani (1994), Aungtonya & Hylleberg (1998), BEDO (2017b).
Bufonaria perelegans Beu, 1987. Gulf of Thailand, Andaman Sea. Reported by: Beu (1987, 1998), BEDO (2017b),
GBIF (2019a).
Bufonaria rana (Linnaeus, 1758). Also reported as: Bufonaria rana phuketensis Kira, 1955; Bufonaria rana
albivaricosa Reeve, 1844; Bursa albivaricosa (Reeve, 1844); and Bursa rana. Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Cossignani (1994), Aungtonya &
Hylleberg (1998), Carpenter & Niem (1998), Kasetsart University (2000), Swennen et al. (2001), Gemert
(2003), Duangdee (2004, 2007), Robba et al. (2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009),
Sanpanich & Duangdee (2013), Galli (2015), BEDO (2017b), GBIF (2019a, 2019b).
Bursa bufonia (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Wells
(2005), Nabhitabhata (2009).
Bursa cruentata (G. B. Sowerby II, 1835). Andaman Sea. Reported by: Duangdee et al. (2013b).
Bursa granularis (Röding, 1798). Also reported as: Colubrellina granularis (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon
(1981), Bussarawit (1995), Wells (2005), Nabhitabhata (2009), BEDO (2017b), Sanpanich & Duangdee (2018),
GBIF (2019a, 2019b).
Bursa rosa (Perry, 1811). Andaman Sea. Reported by: Nateewathana et al. (1981), Wells (2005), BEDO (2017b),
GBIF (2019a).
Tutufa bubo (Linnaeus, 1758). Also reported as: Bursa bubo (Linnaeus). Gulf of Thailand, Andaman Sea. Reported
by: THNHM, Tantanasiriwong (1978), Carpenter & Niem (1998), Kasetsart University (2000), Wells (2005),
Nabhitabhata (2009), Duangdee et al. (2013a), BEDO (2017b), Sanpanich & Duangdee (2018).
Tutufa oyamai Habe, 1973. Also reported as: Tutufa (Tutufella) oyamai (Habe, 1973). Gulf of Thailand, Andaman
Sea. Reported by: Cossignani (1994), Aungtonya & Hylleberg (1998), Nabhitabhata (2009), GBIF (2019a).
Tutufa rubeta (Linnaeus, 1758). Also reported as: Bursa rubeta (Linnaeus). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Cossignani (1994), Bussarawit
(1995), Carpenter & Niem (1998), Wells (2005), Nabhitabhata (2009), BEDO (2017b), Sanpanich & Duangdee
(2018), GBIF (2019a).
Family Cassidae
Casmaria erinaceus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Wells
(2005), Nabhitabhata (2009), Duangdee et al. (2013b).
Casmaria ponderosa (Gmelin, 1791). Also reported as: Casmaria ponderosa ponderosa (Gmelin, 1791). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981),
Bussarawit (1995), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich & Duangdee (2013, 2018),
BEDO (2017b).
Cassis cornuta (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich &
Duangdee (2013), Verbinnen et al. (2016), BEDO (2017b).
Cypraecassis rufa (Linnaeus, 1758). Andaman Sea. Reported by: THNHM, Carpenter & Niem (1998),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Galeodea alcocki (E. A. Smith, 1906). Andaman Sea. Reported by: BEDO (2017b).
Phalium areola (Linnaeus, 1758). Andaman Sea. Reported by: Abbott (1968), Nateewathana et al. (1981),
Sanpanich & Duangdee (2013), BEDO (2017b).
Phalium bandatum (Perry, 1811). Also reported as: Phalium bandatum bandatum (Perry, 1811). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009).
Phalium decussatum (Linnaeus, 1758). Gulf of Thailand. Reported by: Suvatti (1938), Nabhitabhata (2009).
Phalium fimbria (Gmelin, 1791). Andaman Sea. Reported by: Abbott (1968).
Phalium glaucum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Suvatti (1938), Abbott
(1968), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Kreipl (1997), Carpenter
& Niem (1998), Kasetsart University (2000), Swennen et al. (2001), Duangdee (2004), ONR & EPP (2006),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2013), Tan et al. (2013), Galli (2015), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Phalium muangmani Raybaudi Massilia & Prati Musetti, 1995. Andaman Sea. Reported by: Raybaudi Massilia
& Prati Musetti (1995), Kreipl (1997), Verbinnen et al. (2016), GBIF (2019a).
Semicassis bisulcata (Schubert & J. A. Wagner, 1829). Also reported as: Phalium bisulcatum (Schubert &
Wagner, 1829); Phalium bisulcatum bisulcatum (Schubert & Wagner); and Semicassis bisulcata forma diuturna
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The Marine and Estuarine Molluscs of Thailand
(Iredale, 1927). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Abbott (1968), Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1995), Kreipl (1997), Swennen et al. (2001), Duangdee (2007),
Duangdee et al. (2007, 2009, 2011), ONEP (2007), Robba et al. (2007), Nabhitabhata (2009), Taylor (2009),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Family Charoniidae
Charonia tritonis (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nugranad (2000), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), GBIF (2019a).
Family Cymatiidae
Cymatium tripus (Gmelin, 1791). Also reported as: Ranularia tripus (Lamarck). Andaman Sea. Reported by:
THNHM, Nateewathana et al. (1981), BEDO (2017b). This species could not be verified through WoRMS
Editorial Board (2019), but is listed by DiscoverLife (2020).
Gelagna succincta (Linnaeus, 1771). Reported as: Linatella (Gelagna) succincta (Linnaeus, 1771). Andaman Sea.
Reported by: Beu & Cernohorsky (1986).
Gutturnium muricinum (Röding, 1798). Reported as: Cymatium muricinum (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem
(1998), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Gyrineum aculeatum (Schepman, 1909). Reported as: Biplex aculeata (Schepman, 1909); and Biplex pulchrum
(Gray in G. B. Sowerby I, 1836). Andaman Sea. Reported by: Beu (1998), BEDO (2017b).
Gyrineum bituberculare (Lamarck, 1816). Also reported as: Argobuccinum bituberculare. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, Nateewathana et al. (1981), Aungtonya & Hylleberg (1998), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), Dechruksa et al. (2014), BEDO (2017b).
Gyrineum gyrinum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998),
Wells (2005), Kasetsart University (2006), ONR & EPP (2006), Nabhitabhata (2009), Duangdee et al. (2011,
2013a), Sanpanich & Duangdee (2013).
Gyrineum natator (Röding, 1798). Also reported as: Ranella tuberculata Broderip, 1833. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Way & Purchon (1981), Subba Rao & Dey (2000), Swennen et al.
(2001), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
Gyrineum pusillum (Broderip, 1833). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004).
Lotoria lotoria (Linnaeus, 1758). Reported as: Cymatium lotorium (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009), BEDO (2017b).
Lotoria triangularis (Perry, 1811). Reported as: Lotoria perryi (Emerson & Old, 1963); and Cymatium perryi
Emerson & Old, 1963. Andaman Sea. Reported by: THNHM, Nabhitabhata (2009), Duangdee et al. (2011, 2013b),
Sanpanich & Duangdee (2013).
Monoplex aquatilis (Reeve, 1844). Reported as: Cymatium aquatile (Reeve, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009), GBIF (2019a).
Monoplex intermedius (Pease, 1869). Reported as: Cymatium intermedium (Pease, 1869). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata (2009).
Monoplex mundus (Gould, 1849). Andaman Sea. Reported by: GBIF (2019a).
Monoplex nicobaricus (Röding, 1798). Also reported as: Cymatium nicobaricum (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem (1998),
Nabhitabhata (2009), Duangdee et al. (2013b), BEDO (2017b).
Monoplex parthenopeus (Salis Marschlins, 1793). Also reported as: Cymatium succinctum (Linnaeus, 1758). Gulf
of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Nabhitabhata (2009), Duangdee et al.
(2013b), GBIF (2019a, 2019b).
Monoplex pilearis (Linnaeus, 1758). Also reported as: Cymatium pileare aquatile (Reeve, 1844); Cymatium
(Monoplex) pileare (Linné, 1758); Septa pileare (Linnaeus, 1758); and Cymatium pileare (Linnaeus, 1758). Gulf
of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981), Beu & Kay (1988), Bussarawit
(1995), Carpenter & Niem (1998), Swennen et al. (2001), Wells (2005), Kasetsart University (2006), Hylleberg
& Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
Monoplex vespaceus (Lamarck, 1822). Reported as: Gyrineum vespaceum (Lamarck, 1822). Andaman Sea.
Reported by: Duangdee et al. (2013b).
Ranularia caudata (Gmelin, 1791). Reported as: Cymatium caudatum (Gmelin, 1791); Cymatium (Linatella)
cutaceum (Lamarck, 1816); Linatella caudata (Gmelin, 1791); Cymatium (Ranularia) caudatum (Gmelin, 1791);
and Linatella cingulata (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong
(1978), Nateewathana et al. (1981), Bussarawit (1995), Aungtonya & Hylleberg (1998), Beu (1998), Swennen
et al. (2001), Gemert (2003), Robba et al. (2004), Nabhitabhata (2009), BEDO (2017b).
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Ranularia gutturnia (Röding, 1798). Reported as: Cymatium gutturnium (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Swennen et al. (2001), GBIF (2019a).
Ranularia oboesa (Perry, 1811). Also reported as: Cymatium retusum Lamarck, 1822. Gulf of Thailand, Andaman
Sea. Reported by: KUMF, Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF
(2019a).
Ranularia pyrulum (A. Adams & Reeve, 1850). Reported as: Cymatium (Ranularia) pyrulum (A. Adams & Reeve,
1850). Andaman Sea. Reported by: Beu (1986a, 1986b, 2005), GBIF (2019a).
Ranularia pyrum (Linnaeus, 1758). Cymatium pyrum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported
by: KUMF, Carpenter & Niem (1998), Duangdee (2007), Nabhitabhata (2009).
Ranularia sarcostoma (Reeve, 1844). Andaman Sea. Reported by: GBIF (2019a).
Ranularia sinensis (Reeve, 1844). Andaman Sea. Reported by: GBIF (2019a).
Reticutriton pfeifferianus (Reeve, 1844). Reported as: Cymatium pfeifferianum (Reeve, 1844). Gulf of
Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995),
Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Septa hepatica Röding, 1798. Andaman Sea. Reported by: Nateewathana et al. (1981).
Septa mixta (Arthur & Garcia-Talavera, 1990). Andaman Sea. Reported by: Beu (1998).
Septa rubecula (Linnaeus, 1758). Also reported as: Cymatium rubeculum (Linnaeus, 1758). Andaman Sea. Reported
by: ONR & EPP (2006), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Family Personidae
Distorsio anus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF.
Distorsio reticularis (Linnaeus, 1758). Also reported as: Distorsio reticulata (Röding, 1798); and Diatorsio
reticulata. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Nateewathana et al. (1981), Kasetsart
University (2000), Swennen et al. (2001), Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019b).
Family Tonnidae
Malea pomum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Wells
(2005), Vos (2007), Nabhitabhata (2009).
Tonna allium (Dillwyn, 1817). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Tonna canaliculata (Linnaeus, 1758). Also reported as: Tonna cepa (Roeding). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Nateewathana et al. (1981), Carpenter & Niem (1998), Nabhitabhata (2009), BEDO (2017b).
Tonna dolium (Linnaeus, 1758). Also reported as: Dolium maculatum Lamarck, 1822. Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al. (1981), Way &
Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000), Duangdee et al.
(2007, 2011), Vos (2007), Nabhitabhata (2009), Taylor (2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Tonna galea (Linnaeus, 1758). Also reported as: Tonna olearium (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Nateewathana et al. (1981), Carpenter & Niem (1998), Nabhitabhata (2009),
BEDO (2017b).
Tonna lischkeana (Küster, 1857). Andaman Sea. Reported by: Swennen et al. (2001), Vos (2007), Nabhitabhata
(2009), BEDO (2017b).
Tonna melanostoma (Jay, 1839). Reported as: Tonna melenostoma. Gulf of Thailand. Reported by: THNHM.
Tonna perdix (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Tonna sulcosa (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Swennen et al. (2001), BEDO
(2017b), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Tonna tessellata (Lamarck, 1816). Also reported as: Tonna maculata (Lamarck, 1822). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Swennen et al. (2001), Nabhitabhata (2009), J. D. Taylor
et al. (unpublished data).
Superfamily Truncatelloidea
Family Assimineidae
Austropilula beddomeana (Nevill, 1880). Reported as: Austropilula beddomeana Nevill, 1880. Andaman Sea.
Reported by: J. D. Taylor et al. (unpublished data).
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Assiminea zilchi Brandt, 1974. Reported as: Assiminea (Assiminea) zilchi Brandt, 1974. Gulf of Thailand. Reported
by: Brandt (1974), Robba et al. (2007).
Optediceros breviculum (Pfeiffer, 1855). Reported as: Assiminea brevicula (Pfeiffer, 1855); and Ovassiminea
brevicula (Pfeiffer). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978),
Nateewathana et al. (1981), Kasetsart University (2000), Subba Rao & Dey (2000), Swennen et al. (2001),
Suzuki et al. (2002), Robba et al. (2004), Sri-aroon et al. (2005, 2006), Premcharoen et al. (2016), GBIF (2019a),
J. D. Taylor et al. (unpublished data).
Ovassiminea obtusa (Wattebled, 1886). Reported as: Assiminea obtusa Wattebled, 1886; and Assiminea
(Ovassiminea) obtusa Wattebled, 1886. Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004), Negri
et al. (2014).
Paludinellassiminea japonica (Pilsbry, 1901). Andaman Sea. Reported by: BEDO (2017b).
Sculptassiminea spiralis Brandt, 1974. Reported as: Assiminea (Assiminea) spiralis Brandt, 1974. Gulf of
Thailand. Reported by: Brandt (1974), Robba et al. (2007).
Family Caecidae
Caecum clarum (Lamy, 1910). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Caecum japonicum (Habe, 1978). Gulf of Thailand. Reported by: BEDO (2017e).
Caecum cf. neocaledonicum (de Folin, 1868). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Caecum sepimentum (de Folin, 1868). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Mauroceras kajiyamai (Habe, 1963). Reported as: Meioceras kajiyamai (Habe, 1963). Gulf of Thailand, Andaman
Sea. Reported by: BEDO (2017e).
Family Clenchiellidae
Clenchiella minutissima (Wattebled, 1884). Gulf of Thailand. Reported by: Ponder et al. (2014), GBIF (2019b).
Coliracemata microscopica (Nevill, 1877). Gulf of Thailand. Reported by: GBIF (2019b).
Family Iravadiidae
Hyala bella (A. Adams, 1853). Reported as: Iravadia bella (A. Adams, 1853). Gulf of Thailand. Reported by:
Gemert (2003).
Iravadia bombayana (Stoliczka, 1868). Reported as: Fairbankia bombayana Stoliczka, 1868; and Iravadia
(Fairbankia) bombayana (Blanford, 1868). Gulf of Thailand. Reported by: Sri-aroon et al. (2005), Robba et al.
(2007), Nabhitabhata (2009), GBIF (2019b).
Iravadia cochinchinensis (Bavay & Dautzenberg, 1910). Reported as: Iravadia (Fairbankia) cochinchinensis
(Bavay & Dautzenberg, 1910). Gulf of Thailand. Reported by: Robba et al. (2004), Nabhitabhata (2009).
Iravadia ornata Blanford, 1867. Reported as: Iravadia (Iravadia) ornata Blanford, 1867. Gulf of Thailand.
Reported by: Robba et al. (2004), Nabhitabhata (2009), Negri et al. (2014).
Iravadia quadrasi (O. Boettger, 1893). Gulf of Thailand. Reported by: Higo et al. (1999), Nabhitabhata (2009).
Iravadia rohdei (Brandt, 1968). Reported as: Iravadia (Fairbankia) rohdei Brandt, 1968. Gulf of Thailand.
Reported by: Robba et al. (2007), Nabhitabhata (2009).
Lantauia taylori Ponder, 1994. Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014).
Liroceratia sulcata (Boettger, 1893). Andaman Sea. Reported by: BEDO (2017e).
Pellamora densilabrum (Melvill, 1912). Reported as: Iravadia densilabrum (Melvill, 1912). Gulf of Thailand.
Reported by: Gemert (2003).
Pellamora reflecta Laseron, 1956. Reported as: Iravadia reflecta (Laseron, 1956). Andaman Sea. Reported by:
BEDO (2017e).
Pseudomerelina mahimensis (Melvill, 1893). Reported as: Iravadia (Iravadia) tuberculata Brandt, 1974. Gulf of
Thailand. Reported by: Brandt (1974), Robba et al. (2004).
Pseudonoba aristaei (Melvill, 1912). Reported as: Iravadia (Pseudonoba) aristaei (Melvill, 1912). Gulf of Thailand.
Reported by: Robba et al. (2004), Negri et al. (2014).
Pseudonoba delicata (Philippi, 1849). Reported as: Iravadia delicata (Philippi, 1849). Andaman Sea. Reported
by: BEDO (2017e).
Family Spirostyliferinidae
Spirostyliferina wareni (Moolenbeek, 2009). Reported as: Hoenselaaria wareni Moolenbeek, 2009. Andaman
Sea. Reported by: Moolenbeek (2009b).
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Family Stenothyridae
Stenothyra acuta Brandt, 1974. Gulf of Thailand. Reported by: Brandt (1974), Robba et al. (2004).
Stenothyra glabra (Adams, 1861). Gulf of Thailand. Reported by: Premcharoen et al. (2016).
Stenothyra glabrata (Adams, 1851). Gulf of Thailand. Reported by: THNHM, Robba et al. (2004).
Stenothyra monilifera (Benson, 1856). Gulf of Thailand. Reported by: Robba et al. (2004), Golding (2014).
Stenothyra polita (Adams, 1851). Gulf of Thailand. Reported by: Robba et al. (2004), Golding (2014), Negri et al.
(2014).
Stenothyra ventricosa (Quoy & Gaimard, 1834). Reported as: Stenothyra moussoni Martens, 1897. Gulf of
Thailand. Reported by: Gemert (2003).
Family Truncatellidae
Truncatella guerinii A. Villa & J. Villa, 1841. Gulf of Thailand. Reported by: Gemert (2003).
Truncatella pfeifferi (Martens, 1860). Gulf of Thailand. Reported by: BEDO (2017e). This species could not be
verified through WoRMS Editorial Board (2019) but is listed in Ruhoff (1980).
Family Vitrinellidae
Circulus cinguliferus (Adams, 1850). Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004),
Chaiwathee et al. (2007).
Circulus mortoni Ponder, 1994. Gulf of Thailand. Reported by: Robba et al. (2007), Negri et al. (2014).
Pseudoliotia asteriscus (Gould, 1859). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO
(2017e).
Pseudoliotia gowllandi (Brazier, 1875). Andaman Sea. Reported by: GBIF (2019a).
Pseudoliotia granulosa (Kuroda & Habe, 1971). Andaman Sea. Reported by: Nateewathana et al. (1981), BEDO
(2017b).
Pseudoliotia pulchella (Dunker, 1860). Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004),
Chaiwathee et al. (2007), Sawatna et al. (2012).
Pseudoliotia reeviana (Reeve, 1843). Gulf of Thailand. Reported by: Duangdee (2007).
Superfamily Vanikoroidea
Family Eulimidae
Eulima bifascialis (A. Adams, 1864). Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007),
Duangdee et al. (2011).
Eulima major G. B. Sowerby I, 1834. Andaman Sea. Reported by: GBIF (2019a).
Hypermastus echinodisci Warén, 1980. Andaman Sea. Reported by: BEDO (2017b).
Hypermastus minor (Warén, 1991). Andaman Sea. Reported by: BEDO (2017b).
Hypermastus tenuissimae Warén, 1991. Andaman Sea. Reported by: BEDO (2017b)
Melanella cf. acicula (Gould, 1849). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Melanella bovicornu (Pilsbry, 1905). Reported as: Hypermastus tenuissimae (Warén & Crossland, 1991). Gulf of
Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Melanella solidula (A. Adams & Reeve, 1850). Gulf of Thailand. Reported by: Gemert (2003).
Niso venosa G. B. Sowerby III, 1895. Gulf of Thailand. Reported by: Robba et al. (2004).
Parvioris fulvescens (A. Adams, 1866). Gulf of Thailand, Andaman Sea. Reported by: Warén (1981), BEDO
(2017b).
Pyramidelloides mirandus (A. Adams, 1861). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2007),
BEDO (2017e).
Scalenostoma cf. carinatum Deshayes, 1863. Reported as: Scalenostoma carinata Deshayes, 1863. Gulf of Thailand.
Reported by: Gemert (2003).
Sticteulima amamiensis (Habe, 1961). Gulf of Thailand. Reported by: Gemert (2003). Recorded by WoRMS
Editorial Board (2019) as a taxon inquirendum.
Sticteulima lentiginosa (A. Adams, 1861). Gulf of Thailand. Reported by: Gemert (2003).
Thyca crystallina (Gould, 1846). Reported as: Parvioris fulvescens (Sowerby, 1866). Gulf of Thailand, Andaman
Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Family Haloceratidae
Haloceras cf. millestriata (Okutani, 1964). Andaman Sea. Reported by: BEDO (2017e).
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Family Hipponicidae
Cheilea cepacea (Broderip, 1834). Andaman Sea. Reported by: Duangdee et al. (2013b).
Cheilea cicatricosa (Reeve, 1858). Andaman Sea. Reported by: Duangdee et al. (2013b), BEDO (2017b).
Cheilea dormitoria (Reeve, 1858). Andaman Sea. Reported by: BEDO (2017b).
Cheilea equestris (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Wells (2005), Duangdee (2007), Nabhitabhata (2009), Duangdee et al. (2011),
Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b).
Cheilea hipponiciformis (Reeve, 1858). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Cheilea layardi (Reeve, 1858). Andaman Sea. Reported by: Duangdee et al. (2013b).
Cheilea scutulum (Reeve, 1858). Andaman Sea. Reported by: Duangdee et al. (2013a), BEDO (2017b).
Cheilea tortilis (Reeve, 1858). Andaman Sea. Reported by: Duangdee et al. (2013a), BEDO (2017b).
Sabia conica (Schumacher, 1817). Also reported as: Hipponix conicus (Schumacher, 1817); and Hipponix acuta
(Quoy & Gaimard, 1835). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Kilburn & Hylleberg
(1998), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2013a), BEDO (2017e).
Family Vanikoridae
Caledoniella montrouzieri Souverbie, 1869. Andaman Sea. Reported by: Rosewater (1975).
Macromphalus styliferinus (Nevill, 1884). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Robba
et al. (2007), BEDO (2017e).
Macromphalus thelacme (Melvill, 1904). Gulf of Thailand. Reported by: Robba et al. (2004).
Macromphalus tornatilis (Gould, 1859). Gulf of Thailand. Reported by: Robba et al. (2004).
Vanikoro cancellata (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Way & Purchon (1981), Robba et al. (2004), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017b),
Sanpanich & Duangdee (2018), GBIF (2019a).
Vanikoro fenestrata (A. Adams, 1863). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), BEDO (2017b).
Vanikoro helicoidea (Le Guillou, 1842). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), BEDO
(2017b).
Vanikoro plicata (Récluz, 1844). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Wells
(2005), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011).
Superfamily Vermetoidea
Family Vermetidae
Dendropoma planorbis (Dunker, 1860). Gulf of Thailand. Reported by: Kostina et al. (2016).
Petaloconchus renisectus Carpenter, 1857. Gulf of Thailand. Reported by: Gemert (2003), BEDO (2017e).
Petaloconchus tokyoensis (Pilsbry, 1895). Reported as: ? Vermetus tokyoensis Pilsbry, 1895. Gulf of Thailand.
Reported by: Gemert (2003).
Thylacodes adamsii (Mörch, 1859). Also reported as: Serpulorbis imbricatus (Dunker, 1860). Gulf of Thailand.
Reported by: Gemert (2003).
Thylacodes colubrinus (Röding, 1798). Reported as: Serpulorbis colubrinus (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Kasetsart University (2000), Nabhitabhata (2009).
Thylacodes medusae Pilsbry, 1891. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Thylacodes variabilis (Hadfield & Kay, 1972). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Superfamily Xenophoroidea
Family Xenophoridae
Onustus indicus (Gmelin, 1791). Reported as: Tugurium indicum (Gmelin, 1791); and Xenophora (Onustus)
indica (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Ponder (1983), Swennen et al. (2001),
Nabhitabhata (2009).
Stellaria chinensis (Philippi, 1841). Reported as: Xenophora calculifera (Reeve, 1843); Xenophora chinensis
(Philippi, 1841); and Xenophora (Stellaria) chinensis chinensis (Philippi, 1841). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, Ponder (1983), Bussarawit (1995), Subba Rao & Dey (2000), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), GBIF (2019a, 2019b).
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Stellaria solaris (Linnaeus, 1764). Also reported as: Xenophora solaris (Linnaeus, 1764); and Xenophora (Stellaria)
solaris (Linné, 1764). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Ponder (1983), Subba Rao & Dey (2000), Swennen et al. (2001), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a).
Xenophora corrugata (Reeve, 1842). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al.
(1981), Bussarawit (1995), BEDO (2017b), GBIF (2019a).
Xenophora mekranensis (Newton, 1905). Andaman Sea. Reported by: Ponder (1983), GBIF (2019b). Ponder
(1983) and GBIF (2019a) reported the subspecies Xenophora mekranensis konoi Habe, 1953.
Xenophora pallidula (Reeve, 1842). Andaman Sea. Reported by: Ponder (1983), Thamrongnawasawat et al. (2008).
Xenophora solarioides (Reeve, 1845). Also reported as: Xenophora (Xenophora) solarioides solarioides (Reeve,
1845); and Xenophora australis (Souverbie, 1870). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong
(1978), Nateewathana et al. (1981), Ponder (1983), Subba Rao & Dey (2000), Swennen et al. (2001), Gemert
(2003), Robba et al. (2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a).
Order Neogastropoda
Superfamily Buccinoidea
Family Buccinidae
Pseudoneptunea varicosa (Röding, 1798). Reported as: Pseudoneptunea varicosa (Gmelin, 1791). Gulf of
Thailand, Andaman Sea. Reported by: Robba et al. (2004), Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich & Duangdee (2013), GBIF (2019b).
Family Colubrariidae
Colubraria muricata (Lightfoot, 1786). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a).
Colubraria nitidula (G. B. Sowerby I, 1833). Andaman Sea. Reported by: Wells (2005), GBIF (2019a).
Colubraria obscura (Reeve, 1844). Andaman Sea. Reported by: Bouchet & Perrine (1996), BEDO (2017b).
Colubraria tenera (Gray, 1839). Reported as: Colubraria cf. castanea (Kuroda & Habe, 1952). Andaman Sea.
Reported by: Wells (2005).
Kanamarua adonis (Dall, 1919). Andaman Sea. Reported by: BEDO (2017b).
Family Columbellidae
Anachis miser (G. B. Sowerby I, 1844). No Thai locality provided. Reported by: Monsecour & Köhler (2006).
Anachis sanfelipensis H. N. Lowe, 1935. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Anachis terpsichore (G. B. Sowerby II, 1822). Andaman Sea. Reported by: J. D. Taylor et al. (unpublished data).
Cotonopsis phuketensis (Kosuge, Roussy & Muangman, 1998). Reported as: Cotonopsis (Cotonopsis) phuketensis
(Kosuge, Roussy & Muangman, 1998); and Cotonopsis (Cotonopsis) vanwalleghemi Kronenberg & Dekker, 1998.
Andaman Sea. Reported by: Kosuge et al. (1998), Kronenberg & Dekker (1998, 1999), Galli (2015), GBIF (2019a).
Euplica deshayesii (Crosse, 1859). Reported as: Pyrene deshayesii (Crosse, 1859). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b).
Euplica livescens (Reeve, 1859). Reported as: Mitrella livescens (Reeve, 1859). Andaman Sea. Reported by:
Duangdee et al. (2013b).
Euplica scripta (Lamarck, 1822). Also reported as: Pyrene versicolor (Sowerby, 1832); and Euplica versicolor
(G. B. Sowerby I, 1832). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Kurozumi
et al. (1989), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Kostina
et al. (2016), GBIF (2019a).
Euplica turturina (Lamarck, 1822). Also reported as: Pyrene turturina (Lamarck, 1822). Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b),
Sanpanich & Duangdee (2013).
Euplica varians (Sowerby, 1832). Reported as: Pyrene varians (Sowerby, 1832). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b).
Graphicomassa ligula (Duclos, 1840). Also reported as: Mitrella ligula (Duclos, 1840); and Mitrella albina (Kiener,
1841). Andaman Sea. Reported by: Wells (2005), Duangdee et al. (2013b).
Indomitrella puella (G. B. Sowerby I, 1844). Gulf of Thailand. Reported by: Kostina et al. (2016).
Mitrella essingtonensis (Reeve, 1859). Gulf of Thailand. Reported by: Kostina et al. (2016).
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The Marine and Estuarine Molluscs of Thailand
Mitrella scripta (Linnaeus, 1758). Reported as: Pyrene scripta (Linnaeus, 1758); and Columbella scripta (Lamarck,
1816). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Carpenter & Niem (1998),
Gemert (2003), Wells (2005), Nabhitabhata (2009), BEDO (2017b).
Mokumea cf. albovittata (Lopes, Coelho & Cardoso, 1965). Andaman Sea. Reported by: BEDO (2017e).
Mokumea divaricata (Pilsbry, 1904). Gulf of Thailand. Reported by: Robba et al. (2007).
Pardalinops testudinaria (Link, 1807). Reported as: Pyrene testudinaria (Link, 1807). Gulf of Thailand, Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Swennen et al. (2001), Gemert (2003),
Wells (2005), Duangdee (2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b).
Pictocolumbella ocellate (Link, 1807). Reported as: Columbella fulgurans Lamarck, 1819; and Pyrene ocellata
(Link, 1807). Gulf of Thailand. Reported by: Morlet (1889), Duangdee et al. (2007), Nabhitabhata (2009).
Pyrene flava (Bruguière, 1789). Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2013), Duangdee et al. (2011).
Pyrene punctata (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011).
Pyrene splendidula (G. B. Sowerby I, 1844). Gulf of Thailand. Reported by: Kostina et al. (2016).
Pyreneola semipicta (G. B. Sowerby III, 1894). Andaman Sea. Reported by: Monsecour & Köhler (2006).
Seminella comistea (Melvill, 1906). Reported as: Zafra comistea (Melvill, 1906). Gulf of Thailand. Reported by:
Robba et al. (2007).
Seminella peasei (Martens & Langkavel, 1871). Reported as: Zafra (Seminella) peasei (Martens & Langkavel,
1871). Gulf of Thailand. Reported by: Gemert (2003).
Zafra hervieri (Pace, 1902). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Zafra mitriformis A. Adams, 1860. Reported as: Zafra (Zafra) mitriformis A. Adams, 1860. Gulf of Thailand.
Reported by: Gemert (2003).
Zafra pumila (Dunker, 1858). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004), Negri et al. (2014),
BEDO (2017e).
Zafra troglodytes (Souverbie in Souverbie & Montrouzier, 1866). Reported as: Zafra (Zafra) troglodytes
(Souverbie, 1866). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Family Fasciolariidae
Benimakia lanceolata (Reeve, 1847). Also reported as: Latirus lanceolatus (Reeve, 1847). Gulf of Thailand,
Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Wells (2005), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018).
Filifusus filamentosus (Röding, 1798). Also reported as: Pleuroploca filamentosa (Röding, 1798); and Fasciolaria
filamentosa (Roeding). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Wells (2005),
ONR & EPP (2006), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
Fusinus colus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Swennen et al. (2001),
Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Fusinus stannum Callomon & Snyder, 2008. Andaman Sea. Reported as: Fusinus spectrum (A. Adams & Reeve,
1848). Reported by: Callomon & Snyder (2008), GBIF (2019a).
Hemipolygona aldeynzeri (Garcia, 2001). Reported as: Latirus aldeynzeri Garcia, 2001. Andaman Sea. Reported
by: Garcia (2001).
Latirolagena smaragdulus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Kilburn & Hylleberg (1998),
Nabhitabhata (2009), Duangdee et al. (2013b), BEDO (2017b), GBIF (2019a).
Latirus nassoides (Reeve, 1847). Gulf of Thailand. Reported by: BEDO (2017b).
Latirus pictus (Reeve, 1847). Andaman Sea. Reported by: Duangdee et al. (2013b).
Latirus polygonus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Marmorofusus nicobaricus (Röding, 1798). Reported as: Fusinus nicobaricus (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem
(1998), Nabhitabhata (2009), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a).
Nodolatirus nodatus (Gmelin, 1791). Reported as: Latirus nassoides (Reeve, 1847); and Latirus nodatus (Gmelin,
1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nabhitabhata (2009).
Peristernia nassatula (Lamarck, 1822). Andaman Sea. Reported by: Wells (2005).
Peristernia neglecta (A. Adams, 1855). Reported as: Peristernia pulchella neglecta (A. Adams, 1854). Gulf of
Thailand. Reported by: BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Pleuroploca trapezium (Linnaeus, 1758). Also reported as: Fasciolaria trapezium (Linnaeus, 1758). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Way &
Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000), ONR & EPP (2006),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Hylleberg & Aungtonya
(2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
Polygona filosa (Schubert & J. A. Wagner, 1829). Reported as: Fusinus filosus (Schubert & J. A. Wagner, 1829).
Gulf of Thailand. Reported by: Kostina et al. (2016).
Turrilatirus turritus (Gmelin, 1791). Reported as: Latirus turritus (Gmelin, 1791). Andaman Sea. Reported by:
Wells (2005).
Family Melongenidae
Brunneifusus carinifer (Habe & Kosuge, 1966). Reported as Hemifusus carinifer (Habe & Kosuge, 1966). Gulf of
Thailand, Andaman Sea. Reported by: BEDO (2017b), GBIF (2019b).
Brunneifusus ternatanus (Gmelin, 1791). Reported as: Hemifusus ternatanus (Gmelin, 1791); and Pugilina
ternatana (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Kasetsart University (2000), Swennen et al. (2001),
Gemert (2003), Robba et al. (2004), Duangdee (2007), Duangdee et al. (2007, 2011, 2013b), Nabhitabhata
(2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Lenifusus elongatus (Lamarck, 1822). Reported as: Pugilina elongata (Lamarck, 1822). Gulf of Thailand. Reported
by: Robba et al. (2004).
Volegalea cochlidium (Linnaeus, 1758). Reported as: Pugilina cochlidium (Linnaeus, 1758); Volema (Pugilina)
cochlidium (Linnaeus, 1758); Hemifusus cochlidium (Linnaeus, 1758); and Melongena pugilina Born. Gulf of
Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981), D’Asaro (1997), Carpenter & Niem
(1998), Kasetsart University (2000), Gemert (2003), Robba et al. (2004), Duangdee (2007), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), BEDO (2017b), GBIF (2019a, 2019b),
J. D. Taylor et al. (unpublished data).
Volema myristica Röding, 1798. Reported as: Purpura hippocastanum (Linnaeus, 1758); and Thais hippocastanum
(Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti (1937, 1938),
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991, 1995), Kasetsart University (2000),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), GBIF (2019a, 2019b).
Family Nassariidae
Nassaria acuminata (Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1981), BEDO
(2017b).
Nassaria pusilla Röding, 1798. Also reported as: Nassaria (Nassaria) pusilla Röding, 1798. Gulf of Thailand,
Andaman Sea. Reported by: Cernohorsky (1981), Kilburn & Hylleberg (1998), Gemert (2003), Robba et al. (2004),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019b).
Nassaria sinensis G. B. Sowerby II, 1859. Reported as: Hindsia sinensis Sowerby, 1887. Andaman Sea. Reported
by: Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b).
Nassarius acuminatus (Marrat, 1880). Reported as: Nassarius (Zeuxis) acuminatus (Marrat, 1880). Gulf of
Thailand, Andaman Sea. Reported by: Cernohorsky (1984), BEDO (2017b).
Nassarius albescens (Dunker, 1846). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), Kilburn
& Hylleberg (1998), Dekker & Kool (1999), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Duangdee et al. (2013a), BEDO (2017b).
Nassarius arcularius (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), Cernohorsky (1984), Carpenter & Niem (1998), Nabhitabhata (2009), Hylleberg & Aungtonya (2013),
BEDO (2017b).
Nassarius biendongensis Kool, 2003. Gulf of Thailand. Reported by: Kool (2003).
Nassarius cadeei Kool, 2006. Andaman Sea. Reported by: Kool (2006).
Nassarius caelatus (A. Adams, 1852). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019b).
Nassarius castus (Gould, 1850). Also reported as: Nassarius (Zeuxis) castus (Gould, 1850). Gulf of Thailand,
Andaman Sea. Reported by: Cernohorsky (1984), Robba et al. (2004), Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), Negri et al. (2014).
Nassarius celebensis (Schepman, 1907). Gulf of Thailand. Reported by: Robba et al. (2004).
Nassarius clarus (Marrat, 1877). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), Robba et al.
(2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018).
Nassarius comptus (A. Adams, 1852). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Cernohorsky (1984), Bussarawit (1995), Swennen et al. (2001), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019b).
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The Marine and Estuarine Molluscs of Thailand
Nassarius concinnus (Powys, 1835). Also reported as: Nassa (Zuexis) concentrica Marrat, 1874. Gulf of Thailand,
Andaman Sea. Reported by: Cernohorsky (1984), Gemert (2003), Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich & Duangdee (2013), BEDO (2017b).
Nassarius conoidalis (Deshayes, 1832). Also reported as: Nassarius conoidalis gemmulatus (G. P. Deshayes,
1832). Andaman Sea. Reported by: Galli (2015), BEDO (2017b).
Nassarius coronatus (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Cernohorsky (1984), Carpenter & Niem (1998),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017b), GBIF (2019a).
Nassarius crebricostatus (Schepman, 1911). Andaman Sea. Reported by: BEDO (2017b).
Nassarius crematus (Hinds, 1844). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), Carpenter
& Niem (1998), Duangdee (2004), Robba et al. (2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013), BEDO (2017b).
Nassarius crenoliratus (A. Adams, 1852). Reported as: Nassarius (Niotha) crenoliratus (A. Adams, 1852).
Andaman Sea. Reported by: Cernohorsky (1984), Gemert (2003), J. D. Taylor et al. (unpublished data).
Nassarius deshayesianus (Issel, 1866). Reported as: Nassarius deshayesi. Gulf of Thailand, Andaman Sea.
Reported by: THNHM.
Nassarius distortus (A. Adams, 1852). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University (2006),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Nassarius dorsatus (Röding, 1798). Also reported as: Nassarius unicolor (Hombron & Jacquinot, 1848). Gulf of
Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1995), Carpenter & Niem (1998), Kasetsart University (2000), Duangdee (2004), ONR & EPP (20 06),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013), BEDO (2017b).
Nassarius echinatus (A. Adams, 1852). Gulf of Thailand. Reported by: Cernohorsky (1984).
Nassarius foveolatus (Dunker, 1847). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Robba et al. (2004), Meyer et al. (2008), Nabhitabhata (2009), BEDO (2017b).
Nassarius fuscolineatus (E. A. Smith, 1875). Reported as: Reticunassa fucolineata (E. A. Smith, 1875). Gulf of
Thailand. Reported by: Gemert (2003).
Nassarius gaudiosus (Hinds, 1844). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Nassarius glans (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Cernohorsky (1984),
Carpenter & Niem (1998), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), BEDO (2017b).
Nassarius globosus (Quoy & Gaimard, 1833). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Cernohorsky (1984), Bussarawit
(1995), Duangdee (2004), Kasetsart University (2006), ONR & EPP (2006), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013),
BEDO (2017b), J. D. Taylor et al. (unpublished data).
Nassarius granifer (Kiener, 1834). Andaman Sea. Reported by: Wells (2005), Thamrongnawasawat et al. (2008),
BEDO (2017b).
Nassarius graphiterus (Hombron & Jacquinot, 1848). Reported as: Nassarius luridus (Gould, 1850). Gulf of
Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Cernohorsky
(1984), Bussarawit (1995), Nabhitabhata (2009), BEDO (2017b).
Nassarius hirtus (Kiener, 1834). Andaman Sea. Reported by: BEDO (2017b).
Nassarius horridus (Dunker, 1847). Reported as: Hebra horrida (Dunker, 1847). Andaman Sea. Reported by:
Nateewathana et al. (1981), Cernohorsky (1984), BEDO (2017b).
Nassarius idyllius (Melvill & Standen, 1901). Gulf of Thailand. Reported by: Robba et al. (2004).
Nassarius jacksonianus (Quoy & Gaimard, 1833). Gulf of Thailand, Andaman Sea. Reported by: KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Cernohorsky (1984), Swennen et al. (2001), Duangdee
et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Galli (2015), BEDO (2017b), J. D.
Taylor et al. (unpublished data).
Nassarius leptospirus (A. Adams, 1852). Reported as: Plicarcularia leptospira (A. Adams, 1852). Andaman Sea.
Reported by: THNHM, Dechruksa et al. (2014).
Nassarius livescens (Philippi, 1849). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Swennen et al. (2001), Duangdee (2007), Duangdee et al. (2007, 2009, 2011),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a), J. D. Taylor et al. (unpublished data).
Nassarius macrocephalus (Schepman, 1911). Andaman Sea. Reported by: BEDO (2017b).
Nassarius margaritifer (Dunker, 1847). Also reported as: Nassarius margaritiferus (Dunker, 1847). Gulf of
Thailand, Andaman Sea. Reported by: Duangdee (2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Nassarius marratii (E. A. Smith, 1876). Also reported as: Nassa ferruginea Marrat, 1880. Andaman Sea. Reported
by Kool (2013b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Nassarius micans (A. Adams, 1852). Also reported as: Nassarius (Zeuxis) micans (A. Adams, 1852). Gulf of
Thailand. Reported by: Cernohorsky (1984), Robba et al. (2004).
Nassarius niger (Hombron & Jacquinot, 1848). Reported as: Hebra corticata (A. Adams, 1852); Hebra nigra
(Rousseau, 1854); and Nassarius nigrus (Hombron & Jacquinot, 1848). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), Dechruksa et al. (2014), BEDO
(2017b), GBIF (2019a).
Nassarius nodiferus (Powys, 1835). Also reported as: Nassarius (Nassarius) nodiferus (Powys, 1835). Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Cernohorsky (1984), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Nassarius olivaceus (Bruguière, 1789). Also reported as: Zeuxis olivaceus (Bruguière, 1789). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Cernohorsky (1984),
Bussarawit (1995), Swennen et al. (2001), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a, 2019b).
Nassarius optimus (G. B. Sowerby III, 1903). Andaman Sea. Reported by: BEDO (2017b)
Nassarius papillosus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), Wells
(2005), GBIF (2019a).
Nassarius protrusidens (Melvill, 1918). Andaman Sea. Reported by: BEDO (2017b).
Nassarius pseudoconcinnus (E. A. Smith, 1895). Andaman Sea. Reported by: BEDO (2017b).
Nassarius psila (R. B. Watson, 1882). Andaman Sea. Reported by: BEDO (2017b).
Nassarius pullus (Linnaeus, 1758). Also reported as: Arcularia thersites; Nassa thersites; Nassarius
(Plicarcularia) pullus (Linnaeus, 1758); and Plicarcularia pullus (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: KUMF, THNHM, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al. (1981), Way &
Purchon (1981), Cernohorsky (1984), Kurozumi et al. (1989), Swennen et al. (2001), Gemert (2003), Duangdee
(2004), Robba et al. (2004), ONR & EPP (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Sanpanich & Duangdee (2013), Dechruksa et al. (2014), BEDO (2017b), GBIF (2019a), J. D. Taylor et al.
(unpublished data).
Nassarius siquijorensis (A. Adams, 1852). Also reported as: Zeuxis canaliculatas (Lamarck, 1822); and Nassa
(Zeuxis) cingenda Marrat, 1880. Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), Gemert
(2003), Robba et al. (2004), Kool (2007, 2013a), Galli (2015), GBIF (2019a).
Nassarius splendidulus (Dunker, 1846). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1984), BEDO
(2017b).
Nassarius stolatus (Gmelin, 1791). Also reported as: Nassarius (Niotha) stolatus (Gmelin, 1791). Gulf of Thailand,
Andaman Sea. Reported by: Cernohorsky (1984), Swennen et al. (2001), Robba et al. (2004), Meyer et al. (2008),
Nabhitabhata (2009), Negri et al. (2014).
Nassarius succinctus (A. Adams, 1852). Also reported as: Nassarius (Zeuxis) succinctus (A. Adams, 1852).
Andaman Sea. Reported by: GBIF (2019a).
Nassarius sufflatus (Gould, 1860). Andaman Sea. Reported by: THNHM, GBIF (2019a).
Nassarius teretiusculus (A. Adams, 1852). Gulf of Thailand. Reported by: Cernohorsky (1984).
Nassarius venustus (Dunker, 1847). Andaman Sea. Reported by: BEDO (2017b).
Nassarius vittatus (A. Adams, 1853). Gulf of Thailand. Reported by: Cernohorsky (1984).
Phos roseatus (Hinds, 1844). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1995), Nabhitabhata (2009), BEDO (2017b).
Phos sculptilis R. B. Watson, 1886. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Phos senticosus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Bussarawit (1995),
Swennen et al. (2001), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), GBIF
(2019a, 2019b).
Phos textus (Gmelin, 1791). Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009).
Reticunassa paupera (Gould, 1850). Reported as: Nassarius pauperus (Gould); Nassarius (Hima) pauperus (Gould,
1850); and Reticunassa pauperus (Gould, 1850). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky
(1984), Kurozumi et al. (1989), Kilburn & Hylleberg (1998), Gemert (2003).
Reticunassa cf. rotunda (Melvill & Standen, 1896). Reported as: Nassarius (Hima) cf. rotundus (Melvill &
Standen, 1896). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003).
Reticunassa thailandensis Galindo, Kool & Dekker, 2017. Gulf of Thailand. Reported by: Galindo et al. (2017).
Family Pisaniidae
Cantharus tranquebaricus (Gmelin, 1791). Reported as: Cantharus tranquebarica Martini. Andaman Sea.
Reported by: KUMF, Tantanasiriwong (1978), Nabhitabhata (2009), BEDO (2017b).
Clivipollia incarnata (Deshayes, 1834). Reported as: Engina incarnata (Deshayes, 1834). Andaman Sea.
Reported by: Wells (2005).
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Clivipollia pulchra (Reeve, 1846). Reported as: Pollia pulchra (Reeve, 1846) and Cantharus pulcher (Reeve,
1846). Andaman Sea. Reported by: Wells (2005), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009),
Sanpanich & Duangdee (2013).
Engina alveolata (Kiener, 1836). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Wells
(2005), Nabhitabhata (2009), BEDO (2017b).
Engina armillata (Reeve, 1846). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Engina curtisiana (E. A. Smith, 1884). Andaman Sea. Reported by: Wells (2005).
Engina lineata (Reeve, 1846). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana
et al. (1981), Bussarawit (1995), Wells (2005), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO
(2017b).
Engina mendicaria (Linnaeus, 1758). Also reported as: Pusiostoma mendicaria (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Kilburn & Hylleberg (1998), Wells (2005), Kasetsart University (2006), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Sørensen (2012), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a).
Engina phasinola (Duclos, 1840). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), BEDO (2017b).
Engina pulchra (Reeve, 1846). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee et al. (2009),
Nabhitabhata (2009).
Engina zonalis (Lamarck, 1822). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Bussarawit (1995), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Pisania fasciculata (Reeve, 1846). Andaman Sea. Reported by: Wells (2005).
Pisania ignea (Gmelin, 1791). Andaman Sea. Reported by: Gemert (2003), Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Pollia bednalli (G. B. Sowerby III, 1895). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Pollia fumosa (Dillwyn, 1817). Also reported as: Cantharus fumosus (Dillwyn, 1817). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Gemert (2003), Wells (2005), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2013, 2018), Duangdee et al. (2011), Kostina et al. (2016), BEDO (2017b), GBIF (2019a).
Pollia rubiginosa (Reeve, 1846). Gulf of Thailand. Reported by: Nabhitabhata (2009).
Pollia undosa (Linnaeus, 1758). Also reported as: Cantharus undosus (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1995), Carpenter & Niem (1998), Gemert (2003), Duangdee (2004), Wells (2005), Kasetsart University (2006),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), BEDO (2017b), GBIF (2019a).
Solenosteira gatesi Berry, 1963. Gulf of Thailand. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee
(2013).
Superfamily Conoidea
Family Borsoniidae
Maoritomella vallata (Gould, 1869). Gulf of Thailand. Reported by: Robba et al. (2004). This species could not
be verified through WoRMS Editorial Board (2019), but is recorded in Ruhoff (1980).
Family Clathurellidae
Etrema crassilabrum (Reeve, 1843). Andaman Sea. Reported by: BEDO (2017b).
Etrema cf. scalarina (Deshayes, 1863). Andaman Sea. Reported by: BEDO (2017e).
Etrema spurca (Hinds, 1843). Gulf of Thailand. Reported by: Robba et al. (2007).
Etrema streptonotus (Pilsbry, 1904). Reported as: Etrema (Etremopa) streptonotus (Pilsbry, 1904). Gulf of
Thailand. Reported by: Gemert (2003).
Etrema cf. tenera (Hedley, 1899). Andaman Sea. Reported by: BEDO (2017e).
Etremopsis albata (E. A. Smith, 1882). Andaman Sea. Reported by: Galli (2015), GBIF (2019a).
Lienardia caelata (Garrett, 1873). Reported as: Lienardia (Acrista) cf. caelata (Garrett, 1873). Gulf of Thailand.
Reported by: Gemert (2003).
Lienardia cincta (Dunker, 1871, Reported by: BEDO (2017e).
Lienardia crassicostata (Pease, 1860). Andaman Sea. Reported by: BEDO (2017e).
Lienardia cf. lischkeana (Pilsbry, 1904). Reported as: Lienardia (Lienardia) cf. lischkeana (Pilsbry, 1904). Gulf
of Thailand. Reported by: Gemert (2003).
Lienardia mighelsi Iredale & Tomlin, 1917. Reported as: Lienardia (Lienardia) mighelsi Iredale & Tomlin, 1917.
Gulf of Thailand. Reported by: Gemert (2003).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Lienardia purpurata (Souverbie, 1860). Andaman Sea. Reported by: BEDO (2017e).
Lienardia rubida (Hinds, 1843). Reported as: Lienardia (Lienardia) rubida (Hinds, 1843). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Nannodiella acricula (Hedley, 1922). Reported as: Etrema acricula Hedley, 1922. Gulf of Thailand. Reported by:
Robba et al. (2007).
Paraclathurella clothonis Hedley, 1922. Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014).
Pseudoetrema fortilirata (E. A. Smith, 1879). Gulf of Thailand. Reported by: Robba et al. (2004).
Pseudorhaphitoma ditylota (Melvill, 1912). Reported as: Etrema ditylota (Melvill, 1912). Gulf of Thailand.
Reported by: Robba et al. (2004).
Family Clavatulidae
Turricula javana (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Powell (1969),
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Swennen
et al. (2001), Robba et al. (2004), Duangdee et al. (2007, 2011), Sanpanich & Duangdee (2013), BEDO (2017b),
GBIF (2019a, 2019b).
Turricula nelliae (E. A. Smith, 1877). Andaman Sea. Reported by: BEDO (2017b).
Turricula tornata (Dillwyn, 1817). Also reported as: Turricula tornata tornata (Dillwyn, 1817). Andaman Sea.
Reported by: Powell (1969), Tantanasiriwong (1978), Nateewathana et al. (1981), BEDO (2017b), GBIF (2019a).
Family Conidae
Conasprella aculeiformis (Reeve, 1844). Reported as: Conus aculeiformis Reeve, 1844. Gulf of Thailand, Andaman
Sea. Reported by: Lim & Wee (1992).
Conasprella articulata (G. B. Sowerby II, 1873). Andaman Sea. Reported by: BEDO (2017b).
Conasprella comatosa (Pilsbry, 1904). Reported as: Conus comatosa (Pilsbry, 1904). Andaman Sea. Reported by:
Lim & Wee (1992).
Conasprella dictator (Melvill, 1898). Andaman Sea. Reported by: GBIF (2019a).
Conasprella insculptus (Kiener, 1847). Reported as: Conus insculptus Kiener, 1847. Andaman Sea. Reported by:
da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995), Carpenter & Niem (1998), Higo et al. (1999),
Duangdee (2004), Kasetsart University (2006).
Conasprella longurionis (Kiener, 1847). Gulf of Thailand, Andaman Sea. Reported by: Röckel et al. (1995), Higo
et al. (1999), BEDO (2017b), GBIF (2019a, 2019b).
Conasprella memiae (Habe & Kosuge, 1970). Reported as: Conus memiae (Habe & Kosuge, 1970). Gulf of
Thailand. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992).
Conasprella orbignyi (Audouin, 1831). Reported as: Conus orbignyi (Audouin, 1831). Gulf of Thailand, Andaman
Sea. Reported by: Lim & Wee (1992).
Conasprella otohimeae (Kuroda & Itô, 1961). Reported as: Conus otohimae (Kuroda & Itô, 1961). Gulf of
Thailand. Reported by: Lim & Wee (1992).
Conasprella saecularis (Melvill, 1898). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019a).
Conasprella viminea (Reeve, 1849). Reported as: Conus vimineus (Reeve, 1843). Gulf of Thailand, Andaman Sea.
Reported by: da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995), BEDO (2017b), GBIF (2019a).
Conus abbas Hwass in Bruguière, 1792. Reported as: Conus abbas (Gmelin, 1792). Andaman Sea. Reported by:
Lim & Wee (1992), Nabhitabhata (2009).
Conus achatinus Gmelin, 1791. Also reported as: Conus ranunculus Hwass in Bruguière, 1792; and Conus monachus
achatinus (Gmelin, 1792). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Tantanasiriwong
(1978), da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Kasetsart University (2000),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus aculeiformis (Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009).
Conus acutangulus Lamarck, 1810. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), da
Motta & Lenavat (1979), Lim & Wee (1992), Kasetsart University (2006), Nabhitabhata (2009), GBIF (2019a,
2019b).
Conus adenensis E. A. Smith, 1891. Andaman Sea. Reported by: GBIF (2019a).
Conus amadis Gmelin, 1791. Also reported as: Conus amadis amadis neptunus Kiener, 1843; Conus arbornatalis
da Motta, 1978; Conus amadis castaneofasciatus (da Motta, 1978); and Leptoconus amadis (Gmelin, 1791). Gulf
of Thailand, Andaman Sea. Reported by: KUMF, da Motta & Lenavat (1979), Coomans et al. (1980), Lim &
Wee (1992), Bussarawit (1995), Röckel et al. (1995), Subba Rao & Dey (2000), Kasetsart University (2006),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Galli (2015), BEDO (2017b), J. D. Taylor et al.
(unpublished data).
Conus ammiralis Linnaeus, 1758. Also reported as: Conus (Cylinder) ammiralis ammiralis architalassus (var.)
[Lightfoot, J.], 1786. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim & Wee
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The Marine and Estuarine Molluscs of Thailand
(1992), Röckel et al. (1995), Higo et al. (1999), Nabhitabhata (2009), Duangdee et al. (2013b), BEDO (2017b),
GBIF (2019a).
Conus anabathrum Crosse, 1865. Reported as: Conus aureolus Sowerby, 1834. Gulf of Thailand, Andaman Sea.
Reported by: Suvatti (1937, 1938), Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee (1992),
Bussarawit (1995), Carpenter & Niem (1998), Duangdee et al. (2007), Nabhitabhata (2009).
Conus andamanensis E. A. Smith, 1879. Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019a, 2019b).
Conus araneosus [Lightfoot], 1786. Also reported as: Conus araneosus nicobaricus (Hwass & Bruguière, 1792).
Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), BEDO (2017b).
Conus arenatus Hwass in Bruguière, 1792. Also reported as: Conus arenatus Hwass, 1792. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Nateewathana et al. (1981), Lim & Wee (1992),
Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO
(2017b), GBIF (2019a).
Conus aristophanes (G. B. Sowerby II, 1857). Reported as: Miliariconus aristophanes (G. B. Sowerby II, 1857).
Andaman Sea. Reported by: Sørensen (2012).
Conus armadillo Shikama, 1971. Reported as: Conus australis armadillo (Shikama, 1971). Andaman Sea.
Reported by: Suvatti (1938), Lim & Wee (1992).
Conus articulatus G. B. Sowerby III, 1873. Andaman sea. Reported by: Röckel et al. (1995).
Conus asiaticus da Motta, 1985. Andaman Sea. Reported by: Lim & Wee (1992), Nabhitabhata (2009).
Conus augur Lightfoot, 1786. Andaman Sea. Reported by: KUMF, Lim & Wee (1992), Röckel et al. (1995),
Nabhitabhata (2009).
Conus aulicus Linnaeus, 1758. Also reported as: Conus aulicus aurantia Dautzenberg, 1937; and Conus aulicus
propenudus Melvill, 1900. Gulf of Thailand, Andaman Sea. Reported by: KUMF, da Motta & Lenavat (1979),
Lim & Wee (1992), Bussarawit (1995), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee
et al. (2011, 2013b), Sanpanich & Duangdee (2013, 2018), GBIF (2019a).
Conus aureus Hwass & Bruguière, 1792. Reported as: Conus paulucciae Sowerby, 1876. Andaman Sea. Reported
by: da Motta & Lenavat (1979), Roussy (1987), Lim & Wee (1992).
Conus auricomus Hwass & Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: Lim & Wee (1992).
Conus australis Holten, 1802. Reported as: Conus australis Röding, 1798. Andaman Sea. Reported by: da Motta &
Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995), Thamrongnawasawat et al. (2008), BEDO (2017b),
GBIF (2019a).
Conus balteatus G. B. Sowerby I, 1833. Andaman Sea. Reported by: GBIF (2019a).
Conus bandanus Hwass in Bruguière, 1792. Reported as: Conus marmoreus bandanus (Hwass, 1792). Andaman
Sea. Reported by: Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee (1992), Kasetsart University
(2006).
Conus bengalensis (Okutani, 1968). Also reported as: Cylinder bengalensis (Okutani, 1968). Andaman Sea.
Reported by: Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee (1992), Thamrongnawasawat
et al. (2008), Galli (2015), BEDO (2017b), GBIF (2019a).
Conus betulinus Linnaeus, 1758. Also reported as: Conus (Dendroconus) betulinus Linnaeus, 1758. Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Nateewathana et al. (1981), Lim & Wee (1992), Aungtonya &
Hylleberg (1998), Gemert (2003), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Sanpanich
& Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus biliosus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), da Motta
& Lenavat (1979), Lim & Wee (1992).
Conus blanfordianus Crosse, 1867. Andaman Sea. Reported by: da Motta & Lenavat (1979).
Conus bullatus Linnaeus, 1758. Also reported as: Conus bullatus pongo (Shikama & Oishi, 1977). Gulf of Thailand,
Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995).
Conus buxeus (Röding, 1798). Reported as: Conus loroisii Kiener, 1847; and Conus figulinus loroisii (Kiener, 1848).
Andaman Sea. Reported by: Kohn (1978), da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee
(1992), GBIF (2019a).
Conus canonicus Hwass in Bruguière, 1792. Reported as: Conus canonicus Hwass, 1792. Andaman Sea. Reported
by: da Motta & Lenavat (1979), GBIF (2019a).
Conus capitanellus Fulton, 1938. Gulf of Thailand. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992).
Conus capitaneus Linnaeus, 1758. Also reported as: Conus capitanicus. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Morlet (1889), Lim & Wee (1992), Duangdee et al. (2011, 2013b), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a).
Conus capreolus Röckel, 1985. Reported as: Kioconus capreolus (Röckel, 1985). Andaman Sea. Reported by: Galli
(2015).
Conus caracteristicus Fischer von Waldheim, 1807. Gulf of Thailand, Andaman Sea. Reported by: THNHM,
KUMF, Morlet (1889), da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Gemert
(2003), Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), BEDO
(2017b), GBIF (2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Conus catus Hwass in Bruguière, 1792. Andaman Sea. Reported by: THNHM, KUMF, Nateewathana et al. (1981),
Lim & Wee (1992), Thamrongnawasawat et al. (2008), BEDO (2017b), GBIF (2019a).
Conus chaldeus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, da Motta & Lenavat
(1979), Nateewathana et al. (1981), Lim & Wee (1992), Wells (2005), BEDO (2017b).
Conus circumactus Iredale, 1929. Andaman Sea. Reported by: da Motta & Lenavat (1979).
Conus circumcisus Born, 1778. Andaman Sea. Reported by: GBIF (2019a).
Conus cinereus Hwass in Bruguière, 1792. Gulf of Thailand. Reported by: Morlet (1889).
Conus clarus E. A. Smith, 1881. Andaman Sea. Reported by: THNHM.
Conus cocceus Reeve, 1844. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Conus coelinae Crosse, 1858. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim &
Wee (1992), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus coffeae Gmelin, 1791. Reported as: Conus scabriusculus (Dillwyn, 1817). Gulf of Thailand, Andaman Sea.
Reported by: Lim & Wee (1992), Röckel et al. (1995), Higo et al. (1999), GBIF (2019a).
Conus collisus Reeve, 1849. Also reported as: Conus collisus stigmanicus A. Adams, 1853; Asprella collisus Reeve,
1849; and Phasmoconus collisus (L. A. Reeve, 1849). Andaman Sea. Reported by: KUMF, Suvatti (1937, 1938),
Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee (1992), Bussarawit (1995), Duangdee (2004),
Kasetsart University (2006), ONR & EPP (2006), Galli (2015), BEDO (2017b), GBIF (2019a).
Conus consors G. B. Sowerby I, 1833. Also reported as: Conus consors turschi (da Motta, 1985); and Conus magus
anceps (A. Adams, 1853). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Tantanasiriwong
(1978), da Motta & Lenavat (1979), Lim & Wee (1992), Bussarawit (1995), BEDO (2017b), GBIF (2019a).
Conus coronatus Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981),
Way & Purchon (1981), Lim & Wee (1992), Wells (2005), Duangdee et al. (2009, 2011, 2013b), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Conus crocatus Lamarck, 1810. Reported as: Conus thailandis (da Motta, 1978); Darioconus crocatus thailandis
da Motta, 1978; and Conus crocatus thailandis da Motta, 1978. Andaman Sea. Reported by: da Motta (1978), Lim
& Wee (1992), Röckel et al. (1995), Higo et al. (1999), Thamrongnawasawat et al. (2008), Galli (2015), GBIF
(2019a). WoRMS Editorial Board (2019) lists Conus crocatus thailandis da Motta, 1978, as a valid subspecies.
Conus cumingii Reeve, 1848. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim &
Wee (1992), Röckel et al. (1995), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a).
Conus cyanostoma A. Adams, 1855. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Conus daucus Hwass in Bruguière, 1792. Reported as: Conus daucus daucus Hwass in Bruguière, 1792. Andaman
Sea. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus dayriti Röckel & da Motta, 1983. Andaman Sea. Reported by: BEDO (2017b).
Conus distans Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), da Motta & Lenavat (1979), Lim & Wee (1992), Duangdee (2004), Wells (2005), Thamrongnawasawat
et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a).
Conus ebraeus Linnaeus, 1758. Andaman Sea, Gulf of Thailand. Reported by: Nateewathana et al. (1981), Subba
Rao & Dey (2000), Wells (2005), Sørensen (2012), BEDO (2017b), GBIF (2019a).
Conus eburneus Hwass in Bruguière, 1792. Also reported as: Conus eburneus polyglotta (Weinkauff, 1874). Gulf
of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), da Motta &
Lenavat (1979), Nateewathana et al. (1981), Way & Purchon (1981), Coomans et al. (1986), Lim & Wee (1992),
Carpenter & Niem (1998), Subba Rao & Dey (2000), Wells (2005), Thamrongnawasawat et al. (2008),
Duangdee et al. (2011, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Conus emaciatus Reeve, 1849. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), da
Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Duangdee et al. (2011, 2013b),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Conus episcopatus da Motta, 1982. Also reported as: Conus magnificus episcopatus (da Motta, 1982). Gulf of
Thailand, Andaman Sea. Reported by: KUMF, Lim & Wee (1992), GBIF (2019a).
Conus eximius Reeve, 1849. Also reported as: Calamiconus eximius Reeve, 1849. Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Tantanasiriwong (1978), da Motta & Lenavat (1979), Coomans et al. (1986), Lim & Wee
(1992), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004), Duangdee et al. (2011), Sanpanich &
Duangdee (2013, 2018), Galli (2015), BEDO (2017b), GBIF (2019b).
Conus figulinus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Lim & Wee (1992),
Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
Conus fischoederi Röckel & da Motta, 1983. Andaman Sea. Reported by: Röckel et al. (1995), GBIF (2019a).
Conus flavidus Lamarck, 1810. Also reported as: Virgiconus flavidus. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, Nateewathana et al. (1981), Lim & Wee (1992), Wells (2005), Duangdee et al. (2011), Sørensen
(2012), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Conus floridulus A. Adams & Reeve, 1848. Also reported as: Conus (Lividoconus) floridulus A. Adams & Reeve,
1848. Andaman Sea. Reported by: da Motta & Lenavat (1979), BEDO (2017b).
Conus frigidus Reeve, 1848. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979),
Nateewathana et al. (1981), Lim & Wee (1992), Wells (2005), BEDO (2017b), GBIF (2019a).
Conus fulmen Reeve, 1843. Andaman Sea. Reported by: THNHM.
Conus fulmineus Gmelin, 1791. Andaman Sea. Reported by: GBIF (2019a). Reported by WoRMS Editorial Board
(2019) as a nomen dubium.
Conus furvus Reeve, 1843. Reported as: Conus albicans (Sowerby, 1857). Gulf of Thailand. Reported by: Lim &
Wee (1992), Nabhitabhata (2009), GBIF (2019b).
Conus generalis Linnaeus, 1767. Also reported as: Conus generalis krabiensis (da Motta, 1982). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Morlet (1889), da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim
& Wee (1992), Higo et al. (1999), Wells (2005), Thamrongnawasawat et al. (2008), Duangdee et al. (2011),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus geographus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889),
Nateewathana et al. (1981), Lim & Wee (1992), Thamrongnawasawat et al. (2008), Duangdee et al. (2013b),
BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a).
Conus glans Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979),
Lim & Wee (1992), Wells (2005).
Conus glaucus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim &
Wee (1992), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus grangeri G. B. Sowerby III, 1900. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Lim & Wee (1992), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus granum Röckel & Fischöder, 1985. Andaman Sea. Reported by: Röckel et al. (1995), Higo et al. (1999),
GBIF (2019a).
Conus hyaena Hwass in Bruguière, 1792. Reported as: Conus hyaena Hwass, 1792. Andaman Sea. Reported by:
da Motta & Lenavat (1979), GBIF (2019a).
Conus imperialis Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Lim & Wee
(1992), Thamrongnawasawat et al. (2008), Sanpanich & Duangdee (2018), GBIF (2019a).
Conus inscriptus Reeve, 1843. Also reported as: Conus inscriptus maculospira Pilsbry & Johnson, 1911. Andaman
Sea. Reported by: KUMF, Tantanasiriwong (1978), da Motta & Lenavat (1979), Nateewathana et al. (1981),
Lim & Wee (1992), Bussarawit (1995), Röckel et al. (1995), Carpenter & Niem (1998), Thamrongnawasawat
et al. (2008), BEDO (2017b), GBIF (2019a, 2019b).
Conus janus Hwass in Bruguière, 1792. Andaman Sea. GBIF (2019a).
Conus kintoki Habe & Kosuge, 1970. Andaman Sea. Reported by: GBIF (2019a).
Conus legatus Lamarck, 1810. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1938), da Motta & Lenavat
(1979), Lim & Wee (1992), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus leopardus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Suvatti (1937, 1938),
Tantanasiriwong (1978), da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Carpenter
& Niem (1998), Kasetsart University (2006), BEDO (2017b), GBIF (2019a).
Conus litoglyphus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), da Motta & Lenavat
(1979), Lim & Wee (1992), Wells (2005), BEDO (2017b).
Conus litteratus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978),
da Motta & Lenavat (1979), Lim & Wee (1992), Subba Rao & Dey (2000), Thamrongnawasawat et al. (2008),
Duangdee et al. (2011, 2013b), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO
(2017b), GBIF (2019a, 2019b).
Conus lividus Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889),
Nateewathana et al. (1981), Lim & Wee (1992), Wells (2005), Duangdee et al. (2011, 2013b), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Conus locumtenens Blumenbach, 1791. Reported as: Conus schech Weinkauff, 1875. Andaman Sea. Reported by:
da Motta & Lenavat (1979).
Conus luteus G. B. Sowerby I, 1833. Andaman Sea. Reported by: Lim & Wee (1992), GBIF (2019a).
Conus lynceus G. B. Sowerby II, 1858. Andaman Sea. Reported by: da Motta & Lenavat (1979), Nateewathana et
al. (1981), Lim & Wee (1992), Röckel et al. (1995), BEDO (2017b), GBIF (2019a).
Conus magnificus Reeve, 1843. Andaman Sea. Reported by: da Motta & Lenavat (1979), GBIF (2019a).
Conus magus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Lim & Wee
(1992), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019b).
Conus malacanus Hwass in Bruguière, 1792. Andaman Sea. Reported by: da Motta & Lenavat (1979).
Conus marmoreus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Wells (2005),
Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus miles Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Nateewathana
et al. (1981), Lim & Wee (1992), Duangdee (2004), Wells (2005), Thamrongnawasawat et al. (2008), Duangdee
et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Conus miliaris Hwass in Bruguière, 1792. Also reported as: Conus fulgetrum (Sowerby, 1834). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee (1992), Bussarawit
(1995), Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Conus mitratus Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat
(1979), Lim & Wee (1992), GBIF (2019a).
Conus monachus Linnaeus, 1758. Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee
(2013).
Conus monile Hwass in Bruguière, 1792. Reported as: Conus monile (Bruguière, 1792). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Lim & Wee (1992), Röckel et al. (1995), BEDO (2017b), GBIF
(2019a).
Conus morleti Crosse, 1858. Andaman Sea. Reported by: BEDO (2017b).
Conus muriculatus G. B. Sowerby I, 1833. Gulf of Thailand, Andaman Sea. Reported by: Lim & Wee (1992), Kasetsart
University (2006), Duangdee (2007), Duangdee et al. (2013b).
Conus musicus Hwass in Bruguière, 1792. Also reported as: Conus ceylanensis Hwass, 1792. Gulf of Thailand,
Andaman Sea. Reported by: Kohn (1978), da Motta & Lenavat (1979), Lim & Wee (1992), Wells (2005), GBIF
(2019a, 2019b).
Conus mustelinus Hwass in Bruguière, 1792. Also reported as: Conus mustelinus (Hwass, 1792). Gulf of Thailand,
Andaman Sea. Reported by: Lim & Wee (1992), Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee
(2013, 2018).
Conus nigropunctatus G. B. Sowerby II, 1857. Andaman Sea. Reported by: Tantanasiriwong (1978), da Motta &
Lenavat (1979), Lim & Wee (1992), Bussarawit (1995), Kasetsart University (2006), ONEP (2007).
Conus nodulosus G. B. Sowerby II, 1864. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Conus nussatella Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Nateewathana
et al. (1981), Lim & Wee (1992), Wells (2005), BEDO (2017b), Sanpanich & Duangdee (2013, 20 18), GBIF
(2019a, 2019b).
Conus obscurus G. B. Sowerby I, 1833. Andaman Sea. Reported by: Lim & Wee (1992).
Conus omaria Hwass in Bruguière, 1792. Also reported as: Conus omaria patonganus (da Motta, 1982). Gulf of
Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995),
Thamrongnawasawat et al. (2008), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a).
Conus parvatus Walls, 1979. Andaman Sea. Reported by: Röckel et al. (1995), GBIF (2019a).
Conus patamakanthini Delsaerdt, 1998. Andaman Sea. Reported by: Lim & Wee (1992), Bussarawit (1995),
Thamrongnawasawat et al. (2008).
Conus pennaceus Born, 1778. Also reported as: Conus culubrinus; Conus episcopus (Hwass in Bruguière, 1792);
and Conus pennaceus forma episcopus (Hwass, 1792). Gulf of Thailand, Andaman Sea. Reported by: KUMF,
da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Duangdee et al. (2011, 2013a),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a).
Conus pertusus Hwass in Bruguière, 1792. Reported as: Conus pertusus (Bruguière, 1792). Andaman Sea.
Reported by: Suvatti (1938), Lim & Wee (1992), GBIF (2019a).
Conus pica A. Adams & Reeve, 1848. Andaman Sea. Reported by: GBIF (2019a).
Conus planorbis Born, 1778. Also reported as: Conus vitulinus Hwass in Bruguière, 1792. Gulf of Thailand,
Andaman Sea. Reported by: Lim & Wee (1992), Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), GBIF (2019a, 2019b).
Conus praecellens A. Adams, 1855. Also reported as: Conus sowerbii Reeve, 1849. Gulf of Thailand, Andaman Sea.
Reported by: da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995), BEDO (2017b).
Conus pretiosus G. Nevill & H. Nevill, 1874. Reported as: Conus phuketensis da Motta, 1978; Graphiconus pretiosus
(G. Nevill & H. Nevill, 1874); and Conus lynceus phuketensis (da Motta, 1978). Andaman Sea. Reported by: da
Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al. (1995), Carpenter & Niem (1998), Thamrongnawasawat
et al. (2008), Galli (2015), BEDO (2017b), GBIF (2019a).
Conus pseudocedonulli Blainville, 1818. Reported as: Conus ammiralis blainvillii Vignard, 1829. Andaman Sea.
Reported by: Coomans et al. (1980), GBIF (2019a).
Conus pulcher [Lightfoot], 1786. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Conus pulicarius Hwass in Bruguière, 1792. Reported as: Conus pulicarius (Hwass, 1792). Gulf of Thailand,
Andaman Sea. Reported by: Lim & Wee (1992), GBIF (2019a).
Conus queenslandis da Motta, 1984. Gulf of Thailand. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992),
Carpenter & Niem (1998), Swennen et al. (2001).
Conus quercinus [Lightfoot], 1786. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Lim & Wee
(1992), Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a,
2019b).
Conus radiatus Gmelin, 1791. Gulf of Thailand. Reported by: THNHM, Lim & Wee (1992).
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The Marine and Estuarine Molluscs of Thailand
Conus ranonganus da Motta, 1978. Also reported as: Graphiconus ranongana (da Motta, 1978). Andaman Sea.
Reported by: KUMF, Suvatti (1937, 1938), Tantanasiriwong (1978), da Motta & Lenavat (1979), Lim & Wee
(1992), Bussarawit (1995), Röckel et al. (1995), Aungtonya & Hylleberg (1998), Swennen et al. (2001), Duangdee
(2004), ONR & EPP (2006), Duangdee et al. (2007), Thamrongnawasawat et al. (2008), Galli (2015), BEDO
(2017b), GBIF (2019a).
Conus rattus Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: THNHM, da Motta &
Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Wells (2005), Duangdee et al. (2013a), BEDO
(2017b), Sanpanich & Duangdee (2018), GBIF (2019a).
Conus rawaiensis da Motta, 1978. Also reported as: Rhizoconus rawaiensis (A. J. da Motta, 1978). Andaman Sea.
Reported by: da Motta (1978), Lim & Wee (1992), Röckel et al. (1995), Thamrongnawasawat et al. (2008), Galli
(2015), GBIF (2019a).
Conus recluzianus Bernardi, 1853. Also reported as: Kioconus recluzianus simanoki Tenorio, Poppe & Tagaro,
2007. Gulf of Thailand, Andaman Sea. Reported by: Lim & Wee (1992), Tenorio et al. (2007), Galli (2015), BEDO
(2017b), GBIF (2019a, 2019b). WoRMS Editorial Board (2019) lists Kioconus recluzianus (Bernardi, 1853)
as an alternate representation.
Conus rufimaculosus Macpherson, 1959. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al.
(2011), Sanpanich & Duangdee (2013).
Conus sanguinolentus Quoy & Gaimard, 1834. Andaman Sea. Reported by: da Motta & Lenavat (1979).
Conus spectrum Linnaeus, 1758. Gulf of Thailand. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee
(2013).
Conus sponsalis Hwass in Bruguière, 1792. Also reported as: Conus puncturatus (Gmelin, 1791); and Conus
sponsalis. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Lim & Wee (1992), Wells (2005).
Conus striatellus Link, 1807. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), da Motta &
Lenavat (1979), Lim & Wee (1992).
Conus striatus Linnaeus, 1758. Also reported as: Conus floridus G. B. Sowerby II, 1858; Conus floridus chusakida
(da Motta, 1978); and Conus chusaki da Motta, 1978. Gulf of Thailand, Andaman Sea. Reported by: THNHM,
KUMF, Suvatti (1937, 1938), Tantanasiriwong (1978), da Motta & Lenavat (1979), Nateewathana et al. (1981),
Lim & Wee (1992), Röckel et al. (1995), Wells (2005), Thamrongnawasawat et al. (2008), Duangdee et al.
(2009, 2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus striolatus Kiener, 1848. Also reported as: Pionoconus striolatus (Kiener, 1848). Gulf of Thailand, Andaman
Sea. Reported by: Röckel et al. (1995), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), Galli
(2015), GBIF (2019a).
Conus subulatus Kiener, 1847. Andaman Sea. Reported by: GBIF (2019a). Reported by WoRMS Editorial Board
(2019) as a nomen dubium.
Conus sukhadwalai Röckel & da Motta, 1983. Also reported as: Stellaconus sukhadwalai Röckel & da Motta,
1983. Andaman Sea. Reported by: Galli (2015), GBIF (2019a).
Conus sulcatus Hwass in Bruguière, 1792. Reported as: Conus sulcatus Hwass, 1791; and Conus sulcatus bocki
Sowerby, 1881. Andaman Sea. Reported by: KUMF, da Motta & Lenavat (1979), Lim & Wee (1992), Röckel et al.
(1995), Carpenter & Niem (1998), BEDO (2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Conus suratensis Hwass in Bruguière, 1792. Andaman Sea. Reported by: Lim & Wee (1992), Thamrongnawasawat
et al. (2008), GBIF (2019a).
Conus tenuistriatus G. B. Sowerby II, 1858. Andaman Sea. Reported by: Lim & Wee (1992).
Conus terebra Born, 1778. Also reported as: Virgiconus terebra Born, 1778. Gulf of Thailand, Andaman Sea.
Reported by: da Motta & Lenavat (1979), Nateewathana et al. (1981), Lim & Wee (1992), Carpenter & Niem
(1998), Swennen et al. (2001), Duangdee (2004), Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee
(2013, 2018), BEDO (2017b), GBIF (2019a).
Conus tessulatus Born, 1778. Gulf of Thailand, Andaman Sea. Reported by: Lim & Wee (1992), Wells (2005),
Thamrongnawasawat et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a, 2019b).
Conus textile Linnaeus, 1758. Also reported as: Conus textile babelonica (Linnaeus, 1758); Conus textile auriger
(Röding, 1798); Conus textile textile Linnaeus, 1758; Conus textile forma undulatus Solander, 1786; Conus textile
forma verriculum Reeve, 1843; and Conus textile forma auriger (Röding, 1798). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, Morlet (1889), da Motta & Lenavat (1979), Nateewathana et al. (1981), Way & Purchon
(1981), Lim & Wee (1992), Thamrongnawasawat et al. (2008), Sanpanich & Duangdee (2009, 2018), BEDO
(2017b), GBIF (2019a).
Conus urashimanus Kuroda & Itô, 1961. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979),
Lim & Wee (1992), Hylleberg & Aungtonya (2013).
Conus varius Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim &
Wee (1992), GBIF (2019a).
Conus vexillum Gmelin, 1791. Also reported as: Conus sumatrensis Hwass, 1792. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, da Motta & Lenavat (1979), Lim & Wee (1992), Duangdee (2004, 2007), Wells
(2005), Kasetsart University (2006), Thamrongnawasawat et al. (2008), Duangdee et al. (2011, 2013a, 2013b),
Sanpanich & Duangdee (2013), GBIF (2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Conus vicweei Old, 1973. Also reported as: Textilia vicweei (Old, 1973). Andaman Sea. Reported by: da Motta &
Lenavat (1979), Lim & Wee (1992), Galli (2015), BEDO (2017b), GBIF (2019a).
Conus vidua Reeve, 1843. Reported as: Conus marmoreus vidua (Reeve, 1849). Andaman Sea. Reported by: Suvatti
(1937, 1938), da Motta & Lenavat (1979), Lim & Wee (1992), Duangdee (2004, 2007), Kasetsart University
(2006), Sanpanich & Duangdee (2013).
Conus viola Cernohorsky, 1977. Andaman Sea. Reported by: da Motta & Lenavat (1979), Röckel et al. (1995).
Conus virgo Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, da Motta & Lenavat
(1979), Nateewathana et al. (1981), Lim & Wee (1992), Thamrongnawasawat et al. (2008), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus voluminalis Reeve, 1843. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), da Motta
& Lenavat (1979), Lim & Wee (1992), Bussarawit (1995), Carpenter & Niem (1998), Duangdee et al. (2007),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Conus zeylanicus Gmelin, 1791. Andaman Sea. Reported by: da Motta & Lenavat (1979), Lim & Wee (1992).
Conus zonatus Hwass in Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: da Motta & Lenavat
(1979), Lim & Wee (1992), Thamrongnawasawat et al. (2008), Duangdee et al. (2011, 2013b), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a).
Pygmaeconus visseri (Delsaerdt, 1990). Also reported as: Pseudolilliconus visseri (Delsaerdt, 1990). Andaman
Sea. Reported by: GBIF (2019a).
Family Drilliidae
Clathrodrillia flavidula (Lamarck, 1822). Also reported as: Ptychobela flavidula (Lamarck, 1822). Gulf of
Thailand, Andaman Sea. Reported by: Robba et al. (2004), Sanpanich & Duangdee (2013).
Clavus exasperatus (Reeve, 1843). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Clavus humilis (E. A. Smith, 1879). Reported as: Clavus (Clavus) humilis (E. A. Smith, 1879). Gulf of Thailand.
Reported by: Gemert (2003).
Cymatosyrinx impolita Kuroda, Habe & Oyama, 1971. Gulf of Thailand, Andaman Sea. Reported by: Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Plagiostropha quintuplex (Melvill, 1927). Andaman Sea. Reported by: BEDO (2017e).
Ptychobela nodulosa (Gmelin, 1791). Also reported as: Drillia crenularis (Lamarck, 1816). Gulf of Thailand.
Reported by: Morlet (1889), Duangdee et al. (2007).
Ptychobela suturalis (Gray, 1838). Also reported as: Drilla suturalis (Gray, 1838). Andaman Sea. Reported by:
Tantanasiriwong (1978), Hylleberg & Aungtonya (2013), BEDO (2017b).
Family Horaiclavidae
Ceritoturris thailandica Robba, Di Geronimo, Chaimanee, Pietro Negri & Sanfilippo, 2007. Gulf of Thailand.
Reported by: Robba et al. (2007).
Epideira multiseriata (E. A. Smith, 1877). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2007), BEDO
(2017b).
Graciliclava costata (Hedley, 1922). Gulf of Thailand. Reported by: Robba et al. (2007).
Paradrillia inconstans (E. A. Smith, 1875). Gulf of Thailand. Reported by: Robba et al. (2004).
Paradrillia melvilli Powell, 1969. Gulf of Thailand. Reported by: Robba et al. (2004).
Family Mangeliidae
Benthomangelia trophonoidea (Schepman, 1913). Andaman Sea. Reported by: BEDO (2017b).
Eucithara angela (Adams & Angas, 1864). Gulf of Thailand. Reported by: Gemert (2003).
Eucithara arenivaga Hedley, 1922. Andaman Sea. Reported by: Nateewathana et al. (1981).
Eucithara caledonica (E. A. Smith, 1882). Andaman Sea. Reported by: GBIF (2019a).
Eucithara coronata (Hinds, 1843). Also reported as: Eucithara semizonata (Hervier, 1897). Gulf of Thailand.
Reported by: Gemert (2003).
Eucithara funebris (Reeve, 1846). Gulf of Thailand. Reported by: Gemert (2003).
Eucithara cf. marginelloides (Reeve, 1846). Gulf of Thailand. Reported by: Gemert (2003).
Eucithara stromboides (Reeve, 1846). Gulf of Thailand. Reported by: Gemert (2003).
Eucithara vexillum (Reeve, 1846). Andaman Sea. Reported by: GBIF (2019a).
Guraleus himerodes (Melvill & Standen, 1896). Gulf of Thailand. Reported by: Gemert (2003).
Guraleus kamakuranus (Pilsbry, 1904). Gulf of Thailand. Reported by: Gemert (2003).
Ithycythara apicodenticulata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2007. Gulf of Thailand.
Reported by: Robba et al. (2007).
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The Marine and Estuarine Molluscs of Thailand
Ithycythara funicostata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2007. Gulf of Thailand. Reported
by: Robba et al. (2007), Galli (2015).
Mangelia tanabensis J. C. Melvill, 1923. Gulf of Thailand. Reported by: Gemert (2003).
Pseudorhaphitoma crudelis Hedley, 1922. Gulf of Thailand. Reported by: Robba et al. (2004).
Pseudorhaphitoma fuscescens (Thiele, 1925). Gulf of Thailand. Reported by: Robba et al. (2004).
Pseudorhaphitoma heptagona (Dunker, 1871). Gulf of Thailand. Reported by: Robba et al. (2007).
Family Mitromorphidae
Mitromorpha alba (Petterd, 1879). Reported as: Anarithma alba (Pallarad, 1889). Gulf of Thailand, Andaman
Sea. Reported by: BEDO (2017e).
Family Pseudomelatomidae
Crassispira sinensis (Hinds, 1843). Andaman Sea. Reported by: BEDO (2017b).
Funa jeffreysii (E. A. Smith, 1875). Reported as: Inquisitor jeffreysii (E. A. Smith, 1875). Gulf of Thailand, Andaman
Sea. Reported by: BEDO (2017b).
Funa tayloriana (Reeve, 1846). Reported as: Drillia tayloriana (Reeve, 1846). Gulf of Thailand. Reported by:
Morlet (1889).
Inquisitor latifasciata (G. B. Sowerby II, 1870). Gulf of Thailand. Reported by: Robba et al. (2004).
Inquisitor vulpionis Kuroda & Oyama, 1971. Gulf of Thailand. Reported by: Robba et al. (2004).
Family Raphitomidae
Daphnella aureola (Reeve, 1845). Gulf of Thailand. Reported by: GBIF (2019b).
Daphnella axis (Reeve, 1846). Reported as: Hemidaphne axis (Reeve, 1846). Gulf of Thailand. Reported by: Robba
et al. (2007).
Daphnella celebensis Schepman, 1913. Reported as: Antimitra celebensis (Schepman, 1913). Gulf of Thailand.
Reported by: Robba et al. (2007).
Eucyclotoma lactea (Reeve, 1843). Gulf of Thailand. Reported by: Gemert (2003).
Hemilienardia apiculata (Montrouzier, 1864). Reported as: Lienardia (Lienardia) apiculata (Montrouzier in
Souverbie & Montrouzier, 1864). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Hemilienardia goubini (Hervier, 1896). Andaman Sea. Reported by: BEDO (2017e).
Hemilienardia ocellata (Jousseaume, 1883). Andaman Sea. Reported by: BEDO (2017e).
Hemilienardia rubicunda (Gould, 1860). Andaman Sea. Reported by: BEDO (2017e).
Hemilienardia thyridota (Melvill & Standen, 1896). Andaman Sea. Reported by: BEDO (2017e).
Kermia aniani Kay, 1979. Reported as: ? Kermia aniani Kay, 1979. Gulf of Thailand. Reported by: Gemert (2003).
Kermia canistra (Hedley, 1922). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Kermia daedalea (Garrett, 1873). Reported as: Philbertia (Pseudodaphnella) phaedra (Hervier, 1897). Gulf of
Thailand. Reported by: Gemert (2003).
Kermia cf. felina (Hinds, 1843). Andaman Sea. Reported by: BEDO (2017e).
Kermia tokyoensis (Pilsbry, 1895). Gulf of Thailand. Reported by: Gemert (2003).
Kuroshiodaphne supracancellata (Schepman, 1913). Andaman Sea. Reported by: BEDO (2017b).
Microdaphne cf. morrisoni Rehder, 1980. Andaman Sea. Reported by: BEDO (2017e).
Microdaphne trichodes (Dall, 1919). Gulf of Thailand. Reported by: BEDO (2017e).
Pseudodaphnella barnardi (Brazier, 1876). Reported as: Kermia barnardi (Brazier, 1876). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Pseudodaphnella excellens (G. B. Sowerby III, 1913). Reported as: Philbertia (Pseudodaphnella) excellens (G. B.
Sowerby III, 1913). Gulf of Thailand. Reported by: Gemert (2003).
Pseudodaphnella tincta (Reeve, 1846). Reported as: Philbertia (Pseudodaphnella) tincta (Reeve, 1846). Gulf of
Thailand. Reported by: Gemert (2003).
Raphitoma japonica (Melvill, 1895). Reported as: Philbertia japonica (Melvill, 1895). Gulf of Thailand. Reported
by: Gemert (2003).
Taranidaphne amphitrites (Melvill & Standen, 1903). Gulf of Thailand. Reported by: Robba et al. (2007).
Tritonoturris macandrewi (E. A. Smith, 1882). Andaman Sea. Reported by: GBIF (2019a).
Tritonoturris scalaris (Hinds, 1843). Gulf of Thailand. Reported by: Li et al. (2010).
Veprecula vepratica (Hedley, 1903). Andaman Sea. Reported by: BEDO (2017e).
Vepridaphne cestrum (Hedley, 1922). Gulf of Thailand. Reported by: Robba et al. (2007), GBIF (2019a).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Family Terebridae
Cinguloterebra anilis (Röding, 1798). Reported as: Dimidacus stramineus (Gray, 1834). Andaman Sea. Reported
by: Tantanasiriwong (1978), Nateewathana et al. (1981).
Cinguloterebra boucheti (Bratcher, 1981). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Cinguloterebra caddeyi (Bratcher & Cernohorsky, 1982). Gulf of Thailand, Andaman Sea. Reported by: Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Cinguloterebra commaculata (Gmelin, 1791). Reported as: Terebra commaculata (Gmelin, 1791). Gulf of
Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Swennen et al.
(2001), Terryn (2007), BEDO (2017b), GBIF (2019a).
Cinguloterebra pretiosa (Reeve, 1842). Andaman Sea. Reported by: GBIF (2019a).
Duplicaria duplicata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Gemert (2003), Terryn (2007), Taylor (2009), BEDO (2017b).
Duplicaria dussumierii (Kiener, 1837). Reported as: Noditerebra dussumierii (Kiener, 1839). Andaman Sea.
Reported by: KUMF, BEDO (2017b).
Duplicaria evoluta (Deshayes, 1859). Andaman Sea. Reported by: BEDO (2017b).
Duplicaria teramachii Burch, 1965. Andaman Sea. Reported by: BEDO (2017b).
Euterebra tantilla (E. A. Smith, 1873). Gulf of Thailand. Reported by: Robba et al. (2004).
Granuliterebra bathyrhaphe (E. A. Smith, 1875). Gulf of Thailand. Reported by: Robba et al. (2004).
Granuliterebra tricincta (E. A. Smith, 1877). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004),
BEDO (2017b).
Hastula celidonota (Melvill & Sykes, 1898). Andaman Sea. Reported by: Terryn (2007).
Hastula hectica (Linnaeus, 1758). Also reported as: Impages hectica (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Terryn (2007).
Hastula inconstans (Hinds, 1844). Reported as: Impages confusa (Smith). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), BEDO (2017b).
Hastula rufopunctata (E. A. Smith, 1877). Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019b).
Hastulopsis conspersa (Hinds, 1844). Andaman Sea. Reported by: BEDO (2017b).
Impages bacillus (Deshayes, 1859). Andaman Sea. Reported by: Terryn (2007), BEDO (2017b), GBIF (2019a).
Myurella affinis (Gray, 1834). Reported as: Terebra affinis (Gray, 1834). Andaman Sea. Reported by: Morlet (1889),
Wells (2005).
Myurella cf. columellaris (Hinds, 1844). Reported as: Terebra cf. columellaris (Hinds, 1844). Andaman Sea.
Reported by: Wells (2005).
Myurella nebulosa (G. B. Sowerby I, 1825). Reported as: Terebra nebulosa (Sowerby, 1825). Andaman Sea.
Reported by: Kilburn & Hylleberg (1998), Terryn (2007).
Myurella undulata (Gray, 1834). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Oxymeris areolata (Link, 1807). Reported as: Terebra areolata (Link, 1807). Gulf of Thailand, Andaman Sea.
Reported by: Carpenter & Niem (1998), Duangdee (2007).
Oxymeris chlorata (Lamarck, 1822). Reported as: Terebra chlorata (Lamarck, 1822). Andaman Sea. Reported
by: Wells (2005).
Oxymeris crenulata (Linnaeus, 1758). Reported as: Terebra crenulata (Linnaeus, 1758); and Terebra crenulata
interlineata. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Bussarawit (1995), Wells
(2005), Thamrongnawasawat et al. (2008).
Oxymeris dimidiata (Linnaeus, 1758). Reported as: Terebra dimidiata (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Wells (2005), Duangdee et al. (2013b), BEDO (2017b).
Oxymeris felina (Dillwyn, 1817). Reported as: Terebra felina (Dillwyn, 1817). Andaman Sea. Reported by: Wells
(2005).
Oxymeris maculata (Linnaeus, 1758). Reported as: Terebra maculata (Linnaeus, 1758). Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti (1938), Tantanasiriwong (1978), Nateewathana et al.
(1981), Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004), Wells (2005), Thamrongnawasawat
et al. (2008), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Pristiterebra miranda (E. A. Smith, 1873). Also reported as: Terebra miranda (E. A. Smith, 1873). Gulf of
Thailand. Reported by: Bratcher & Cernohorsky (1987), Robba et al. (2004), Terryn (2007).
Punctoterebra nitida (Hinds, 1844). Reported as: Strioterebrum nitidum (Hinds, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Strioterebrum plumbeum (Quoy & Gaimard, 1833). Also reported as: Terebra plumbeum Quoy & Gaimard, 1833.
Andaman Sea. Reported by: Gemert (2003), BEDO (2017b).
Terebra amanda Hinds, 1844. Andaman Sea. Reported by: GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Terebra babylonia Lamarck, 1822. Gulf of Thailand, Andaman Sea. Reported by: Morlet (1889), Duangdee et al.
(2009, 2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Terebra cingulifera Lamarck, 1822. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Terebra guttata (Röding, 1798). Andaman Sea. Reported by: Wells (2005).
Terebra helichrysum Melvill & Standen, 1903. Andaman Sea. Reported by: BEDO (2017b).
Terebra laevigata Gray, 1834. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Terebra quoygaimardi Cernohorsky & Bratcher, 1976. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich
& Duangdee (2009, 2013), Duangdee et al. (2011).
Terebra spectabilis (Hinds, 1844). Reported as: Duplicaria spectabilis (Hinds, 1843). Gulf of Thailand. Reported
by: Robba et al. (2004). WoRMS Editorial Board (2019) lists Terebra spectabilis as a taxon inquirendum.
Terebra subulata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Wells (2005), Kasetsart University (2006), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018).
Terebra textilis Hinds, 1844. Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2007), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Terenolla pygmaea (Hinds, 1844). Andaman Sea. Reported by: Wells (2005), BEDO (2017b).
Triplostephanus triseriatus (Gray, 1834). Also reported as: Terebra triseriata (Gray, 1834). Andaman Sea.
Reported by: Wells (2005), BEDO (2017b).
Family Turridae
Gemmula graeffei (Weinkauff, 1875). Andaman Sea. Reported by: GBIF (2019a).
Gemmula kieneri (Doumet, 1840). Andaman Sea. Reported by: BEDO (2017b).
Gemmula rosario Shikama & Hayashi, 1977. Andaman Sea. Reported by: GBIF (2019a).
Gemmula speciosa (Reeve, 1843). Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Nateewathana
et al. (1981), BEDO (2017b), GBIF (2019a).
Lophiotoma acuta (Perry, 1811). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), BEDO
(2017b), GBIF (2019a).
Polystira nobilis (Hinds, 1843). No Thai locality provided. Reported by: THNHM.
Turridrupa bijubata (Reeve, 1845). Andaman Sea. Reported by: Wells (2005).
Turris annulata (Reeve, 1843). Andaman Sea. Reported by: Powell (1964), Subba Rao & Dey (2000), Kilburn
et al. (2012).
Turris babylonia (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998).
Turris crispa (Lamarck, 1816). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Kilburn et al. (2012), BEDO (2017b).
Turris nadaensis Azuma, 1973. Andaman Sea. Reported by: Kilburn et al. (2012).
Turris nodifera (Lamarck, 1822). Reported as: Pleurotoma nodifera Lamarck, 1822. Gulf of Thailand. Reported
by: Morlet (1889).
Turris spectabilis (Reeve, 1843). Gulf of Thailand, Andaman Sea. Reported by: Kilburn et al. (2012).
Unedogemmula indica (Röding, 1798). Reported as: Lophiotoma indica (Röding, 1798). Gulf of Thailand, Andaman
Sea. Reported by: KUMF, Powell (1964), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1995), Aungtonya & Hylleberg (1998), Wells (2005), Duangdee (2013), BEDO (2017b).
Unedogemmula unedo (Kiener, 1839). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Superfamily Mitroidea
Family Mitridae
Domiporta circula (Kiener, 1838). Reported as: Neocancilla circula (Kiener, 1838); and Subcancilla circula (Kiener,
1838). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Cernohorsky (1991), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a).
Domiporta granatina (Lamarck, 1811). Also reported as: Neocancilla granatina (Lamarck, 1811). Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Cernohorsky (1991), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Domiporta praestantissima (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011)
Sanpanich & Duangdee (2013).
Gemmulimitra avenacea (Reeve, 1845). Also reported as: Mitra (Nebularia) avenacea (Reeve, 1845). Gulf of
Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Gemmulimitra duplilirata (Reeve, 1845). Reported as: Ziba duplilirata (Reeve, 1845). Gulf of Thailand. Reported
by: Cernohorsky (1991).
Gemmulimitra rubiginosa (Reeve, 1844). Reported as: Mitra rubiginosa (Reeve, 1844); and Mitra (Nebularia)
rubiginosa (Reeve, 1844). Andaman Sea. Reported by: Cernohorsky (1976a), Higo et al. (1999).
Imbricaria bacillum (Lamarck, 1811). Reported as: Ziba bacillum (Lamarck, 1811). Andaman Sea. Reported by:
Cernohorsky (1991).
Imbricaria flammea (Quoy & Gaimard, 1833). Reported as: Ziba flammea (Quoy & Gaimard, 1833). Gulf of
Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), GBIF (2019b).
Imbricaria insculpta (A. Adams, 1853). Reported as: Ziba insculpta (A. Adams, 1853). Andaman Sea. Reported
by: Cernohorsky (1991).
Imbricaria interlirata (Reeve, 1844). Reported as: Subcancilla interlirata (Reeve, 1844); and Ziba interlirata
(Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Cernohorsky (1991), Bussarawit (1995), Kilburn & Hylleberg (1998), Swennen et al. (2001), Duangdee (2004),
Robba et al. (2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Imbricaria pretiosa (Reeve, 1844). Andaman Sea. Reported by: GBIF (2019a).
Imbricaria rufilirata (A. Adams & Reeve, 1850). Andaman Sea. Reported by: GBIF (2019a).
Imbricariopsis punctata (Swainson, 1821). Reported as: Imbricaria punctata (Swainson, 1821). Andaman Sea.
Reported by: Wells (2005).
Isara swainsonii (Broderip, 1836). Reported as: Mitra swainsonii (Broderip, 1836). Andaman Sea. Reported by:
BEDO (2017b).
Mitra mitra (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Cernohorsky (1976a),
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Wells
(2005), Nabhitabhata (2009), BEDO (2017b), Sanpanich & Duangdee (2018).
Mitra stictica (Link, 1807). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Carpenter & Niem (1998),
Nabhitabhata (2009).
Mitra turgida Reeve, 1845. Andaman Sea. Reported by: BEDO (2017b).
Nebularia acuminata (Swainson, 1824). Andaman Sea. Reported by: Cernohorsky (1976a).
Nebularia aegra (Reeve, 1845). Reported as: Cancilla aegra (Reeve, 1845). Gulf of Thailand, Andaman Sea.
Reported by: Cernohorsky (1991), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Nebularia chrysostoma (Broderip, 1836). Gulf of Thailand, Andaman Sea. Also reported as: Mitra chrysostoma
(Broderip, 1836). Reported by: Cernohorsky (1976a), Wells (2005), Duangdee et al. (2013b).
Nebularia contracta (Swainson, 1820). Also reported as: Mitra contracta (Swainson, 1820). Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Duangdee et al.
(2013a).
Nebularia eremitarum (Röding, 1798). Reported as: Mitra eremitarium Röding, 1798. Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Wells (2005), Nabhitabhata (2009).
Nebularia fastigium (Reeve, 1845). Reported as: Mitra fastigium (Reeve, 1845). Gulf of Thailand, Andaman Sea.
Reported by: Cernohorsky (1976a), Wells (2005).
Nebularia pellisserpentis (Reeve, 1844). Andaman Sea. Reported by: Cernohorsky (1976a).
Nebularia ustulata (Reeve, 1844). Reported as: Mitra ustulata (Reeve, 1844). Gulf of Thailand, Andaman Sea.
Reported by: Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Neocancilla clathrus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Cernohorsky (1991), Nabhitabhata (2009), Duangdee et al. (2011, 2013a), Sanpanich
& Duangdee (2013), BEDO (2017b), GBIF (2019a).
Neocancilla papilio (Link, 1807). Andaman Sea. Reported by: Wells (2005).
Pseudonebularia atjehensis (Oostingh, 1939). Reported as: Mitra atjehensis (Oostingh, 1939); and Mitra
(Nebularia) atjehensis (Oostingh, 1939). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1976a),
Higo et al. (1999), GBIF (2019a).
Pseudonebularia chrysalis (Reeve, 1844). Reported as: Mitra chrysalis Reeve, 1844. Gulf of Thailand, Andaman
Sea. Reported by: Cernohorsky (1976a), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013).
Pseudonebularia cucumerina (Lamarck, 1811). Reported as: Mitra cucumerina (Lamarck, 1811). Andaman Sea.
Reported by: Cernohorsky (1976a), Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009),
Duangdee et al. (2013b), BEDO (2017b), GBIF (2019a).
Pseudonebularia doliolum (Küster, 1839). Reported as: Mitra doliolum Kuster, 1839. Andaman Sea. Reported
by: Duangdee et al. (2013b).
Pseudonebularia fraga (Quoy & Gaimard, 1833). Reported as: Mitra fraga Quoy & Gaimard, 1833; and Mitra
(Nebularia) fraga Quoy & Gaimard, 1833. Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1976a),
Tantanasiriwong (1978), Nateewathana et al. (1981), Gemert (2003), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2013), Duangdee et al. (2011).
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The Marine and Estuarine Molluscs of Thailand
Pseudonebularia proscissa (Reeve, 1844). Reported as: Mitra proscissa (Reeve, 1844). Gulf of Thailand, Andaman
Sea. Reported by: Cernohorsky (1976a), Nateewathana et al. (1981), Wells (2005), Nabhitabhata (2009),
Hylleberg & Aungtonya (2013), BEDO (2017b).
Pseudonebularia rubritincta (Reeve, 1844). Also reported as: Mitra rubritincta Reeve, 1844. Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Duangdee et al. (2013b).
Pseudonebularia tabanula (Lamarck, 1811). Reported as: Mitra tabanula (Lamarck, 1811). Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013).
Pterygia crenulata (Gmelin, 1791). Andaman Sea. Reported by: Nateewathana et al. (1981).
Pterygia dactylus (Linnaeus, 1767). Andaman Sea. Reported by: Cernohorsky (1991).
Pterygia sinensis (Reeve, 1844). Andaman Sea. Reported by: BEDO (2017b).
Roseomitra rosacea (Reeve, 1845). Reported as: Mitra rosacea (Reeve, 1845); and Mitra (Nebularia) rosacea
(Reeve, 1845). Andaman Sea. Reported by: Cernohorsky (1976a), Higo et al. (1999).
Scabricola coriacea (Reeve, 1845). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1991), GBIF
(2019a).
Scabricola desetangsii (Kiener, 1838). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Cernohorsky (1991), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017b),
GBIF (2019a, 2019b).
Scabricola olivaeformis (Swainson, 1821). Reported as: Imbricaria olivaeformis (Swainson, 1821). Andaman Sea.
Reported by: Wells (2005).
Strigatella ambigua (Swainson, 1829). Reported as: Mitra ambigua Swainson, 1829. Gulf of Thailand, Andaman
Sea. Reported by: Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Strigatella aurantia (Gmelin, 1791). Reported as: Mitra aurantia (Gmelin, 1791); and Mitra (Nebularia) aurantia
(Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti (1937, 1938),
Cernohorsky (1976a), Tantanasiriwong (1978), Nateewathana et al. (1981), Gemert (2003), Duangdee (2004),
Wells (2005), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013), Hylleberg & Aungtonya (2013), BEDO (2017b), GBIF (2019a).
Strigatella colombelliformis (Kiener, 1838). Reported as: Mitra colombelliformis Kiener, 1838. Gulf of Thailand,
Andaman Sea. Reported by: Kasetsart University (2006), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013), GBIF (2019b).
Strigatella coronata (Lamarck, 1811). Reported as: Mitra coronata (Lamarck, 1811). Gulf of Thailand, Andaman
Sea. Reported by: Cernohorsky (1976a), Wells (2005), Nabhitabhata (2009).
Strigatella fulvescens (Broderip, 1836). Andaman Sea. Reported by: GBIF (2019a).
Strigatella litterata (Lamarck, 1811). Reported as: Mitra litterata (Lamarck, 1811). Gulf of Thailand, Andaman
Sea. Reported by: Cernohorsky (1976a), Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata
(2009), BEDO (2017b).
Strigatella paupercula (Linnaeus, 1758). Reported as: Mitra paupercula (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Strigatella pica (Dillwyn, 1817). Andaman Sea. Reported by: Cernohorsky (1976a).
Strigatella retusa (Lamarck, 1811). Reported as: Mitra retusa (Lamarck, 1811). Gulf of Thailand, Andaman Sea.
Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b).
Strigatella scutulata (Gmelin, 1791). Reported as: Mitra scutulata (Gmelin, 1791); Mitra (Strigatella) scutulata
f. amphorella Lamarck, 1811; and Mitra scutula. Gulf of Thailand, Andaman Sea. Reported by: THNHM,
Cernohorsky (1976a), Nateewathana et al. (1981), Way & Purchon (1981), Gemert (2003), Duangdee (2004,
2013), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich
& Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Swainsonia fissurata (Lamarck, 1811). Reported as: Scabricola fissurata (Lamarck, 1811). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Swainsonia fusca (Swainson, 1824). Reported as: Scabricola (Swainsonia) fusca (Swainson, 1824). Gulf of
Thailand. Reported by: Cernohorsky (1991).
Swainsonia ocellata (Swainson, 1831). Also reported as: Scabricola (Swainsonia) ocellata ocellata (Swainson,
1831). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1991), Nabhitabhata (2009), Duangdee et
al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a, 2019b).
Superfamily Muricoidea
Family Muricidae
Attiliosa houarti Vokes, 1999. Andaman Sea. Reported by: Merle et al. (2011).
Attiliosa orri (Cernohorsky, 1976). Also reported as: Muricopsis orri (Cernohorsky, 1976). Gulf of Thailand,
Andaman Sea. Reported by: Cernohorsky (1976b), Bussarawit (1995), Middelfa rt (1997), Aungtonya &
Hylleberg (1998), Nabhitabhata (2009), Merle et al. (2011), BEDO (2017b).
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Babelomurex indicus (E. A. Smith, 1899). Gulf of Thailand. Reported by: GBIF (2019b).
Chicoreus axicornis (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Bussarawit (1991,
1995), Houart (2001), Nabhitabhata (2009), BEDO (2017b).
Chicoreus banksii (G. B. Sowerby II, 1841). Gulf of Thailand, Andaman Sea. Reported by: Middelfart (1997),
Swennen et al. (2001), Wells (2005), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a, 2019b).
Chicoreus brunneus (Link, 1807). Also reported as: Chicoreus (Triplex) brunneus (Link, 1807); Chicoreus burnneus
(Link); and Murex adustus. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1992, 1995), Middelfart (1997),
Carpenter & Niem (1998), Kasetsart University (2000, 2006), Gemert (2003), Duangdee (2004, 2007), ONR &
EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Merle et al. (2011), Hylleberg & Aungtonya (2013),
BEDO (2017b), GBIF (2019a, 2019b).
Chicoreus capucinus (Lamarck, 1822). Also reported as: Murex capucinus; and Naquetia capucina (Lamarck).
Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Nielsen (1976b), Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1991, 1992), Middelfart (1996, 1997), Kasetsart University (2000),
Sri-aroon et al. (2005), Meyer et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011),
Nabhitabhata (2009), Sanpanich & Duangdee (2013), Dechruksa et al. (2014), BEDO (2017b), GBIF (2019a,
2019b), J. D. Taylor et al. (unpublished data).
Chicoreus microphyllus (Lamarck, 1816). Also reported as: Chicoreus (Triplex) microphyllus (Lamarck, 1816).
Gulf of Thailand, Andaman Sea. Reported by: Houart (1992), Bussarawit (1995), Wells (2005), Nabhitabhata
(2009), Merle et al. (2011), GBIF (2019a, 2019b).
Chicoreus palmarosae (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1991, 1995),
Middelfart (1997), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), GBIF (2019a).
Chicoreus ramosus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Bussarawit (1991, 1992, 1995), Hylleberg & Nateewathana (1992), Bech (1996), Middelfart (1997),
Carpenter & Niem (1998), Swennen et al. (2001), Duangdee (2004), KURDI et al. (2004), Wells (2005), Kasetsart
University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b),
GBIF (2019a, 2019b).
Chicoreus saulii (G. B. Sowerby II, 1841). Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit
(1991).
Chicoreus torrefactus (G. B. Sowerby II, 1841). Also reported as: Chicoreus (Triplex) torrefactus (G. B. Sowerby II,
1841). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Way & Purchon (1981), Bussarawit (1991,
1992, 1995), Hylleberg & Nateewathana (1992), Middelfart (1997), Carpenter & Niem (1998), Gemert (2003),
Duangdee (2004, 2007), KURDI et al. (2004), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al.
(2007, 2009, 2011, 2013b), ONEP (2007), Houart (2008), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Merle et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018), BEDO (2017b),
GBIF (2019a).
Coralliophila costularis (Lamarck, 1816). Andaman Sea, Gulf of Thailand. Reported by: Wells (2005), Printrakoon
et al. (2008).
Coralliophila fearnleyi (Emerson & D’Attilio, 1965). Also reported as: Coralliophila (Bachelomurex) fearnleyi
(Emerson & D’Attilio, 1965). Gulf of Thailand, Andaman Sea. Reported by: Duangdee (2004), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Coralliophila rubrococcinea Melvill & Standen, 1901. Gulf of Thailand. Reported by: Robba et al. (2007).
Coralliophila violacea (Kiener, 1836). Also reported as: Coralliophila neritoidea (Lamarck, 1816). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Wells (2005), Nabhitabhata (2009), Duangdee et al.
(2013b), BEDO (2017b), GBIF (2019a).
Cytharomorula lefevreiana (Tapparone Canefri, 1880). Andaman Sea. Reported by: Bussarawit (1991).
Cytharomorula paucimaculata (G. B. Sowerby III, 1903). Reported as: Pascula paucimaculata (Sowerby). Gulf
of Thailand. Reported by: Kurozumi et al. (1989).
Drupa morum Röding, 1798. Also reported as: Drupa morum morum Roeding. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Emerson & Cernohorsky (1973), Tantanasiriwong (1978), Nateewathana et al.
(1981), Bussarawit (1991, 1995), Houart & Vilvens (1997), Middelfart (1997), Wells (2005), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Drupa ricinus (Linnaeus, 1758). Also reported as: Drupa ricinus ricinus (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Emerson & Cernohorsky (1973), Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Wells (2005), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Drupa rubusidaeus Röding, 1798. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Bussarawit
(1995), Middelfart (1997), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2013b), GBIF (2019a).
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The Marine and Estuarine Molluscs of Thailand
Drupella cornus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1991), Middelfart
(1997), Wells (2005), GBIF (2019a).
Drupella eburnea (Küster, 1862). Gulf of Thailand. Reported by: Gemert (2003).
Drupella margariticola (Broderip in Broderip & Sowerby, 1833). Also reported as: Drupa magaviticola;
Cronia margariticola (Broderip, 1833); Morula margariticola (Broderip, 1832); Morula (Cronia) margariticola
(Broderip, 1833); and Ergalatax margariticola (Broderip, 1833). Gulf of Thailand, Andaman Sea. Reported by:
Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1991), Middelfart (1996, 1997), Swennen et al.
(2001), Gemert (2003), Duangdee (2004, 2007), Wells (2005), Kasetsart University (2006), Duangdee et al.
(2007, 2009, 2011), ONEP (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), Sørensen (2012), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013, 2018),
Kostina et al. (2016), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Drupella rugosa (Born, 1778). Also reported as: Drupella concatenata (Lamarck); and Thais rugosa Born, 1778.
Gulf of Thailand, Andaman Sea. Reported by: KUMF, Kurozumi et al. (1989), Bussarawit (1991), Middelfart
(1996, 1997), Gemert (2003), Robba et al. (2004), Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013, 2018), Duangdee et al. (2011), BEDO (2017b), GBIF (2019a, 2019b).
Drupina lobata (Blainville, 1832). Also reported as: Drupa lobata (Blainville, 1832). Gulf of Thailand, Andaman
Sea. Reported by: Emerson & Cernohorsky (1973), Bussarawit (1991), Middelfart (1997), Subba Rao & Dey
(2000), Wells (2005), Duangdee et al. (2013b), BEDO (2017b), GBIF (2019a).
Ergalatax contracta (Reeve, 1846). Also reported as: Cronia (Ergalatax) contracta (Reeve, 1846). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Kurozumi et al. (1989),
Bussarawit (1991), Middelfart (1997), Subba Rao & Dey (2000), Swennen et al. (2001), Gemert (2003), Robba
et al. (2004), Duangdee (2007), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013), BEDO (2017b), GBIF (2019a, 2019b).
Favartia maculata (Reeve, 1845). Andaman Sea. Reported by: Houart & Gori (2011).
Favartia salmonea (Melvill & Standen, 1899). Andaman Sea. Reported by: Bussarawit (1991), Houart (2016).
Favartia sykesi (Preston, 1904). Andaman Sea. Reported by: Houart (2016).
Haustellum haustellum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Bussarawit (1991),
Carpenter & Niem (1998), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Merle et al. (2011).
Homalocantha anatominca (Perry, 1811). Andaman Sea. Reported by: KUMF, Nabhitabhata (2009), GBIF (2019a).
Homalocantha scorpio (Linnaeus, 1758). Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Homalocantha zamboi (Burch & Burch, 1960). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1991),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Indothais blanfordi (Melvill, 1893). Andaman Sea. Reported by: Houart & Delemarre (2013).
Indothais gradata (Jonas, 1846). Reported as: Thais gradata (Jonas, 1846). Andaman Sea. Reported by: THNHM,
Bussarawit (1991), Middelfart (1997), Meyer et al. (2008), GBIF (2019a).
Indothais javanica (Philippi, 1848). Reported as: Thais javanica (Philippi, 1848); and Stramonita javanica
(Philippi, 1848). Andaman Sea. Reported by: THNHM, Tan & Sigurdsson (1996), Middelfart (1997), Houart &
Delemarre (2013), Negri et al. (2014), GBIF (2019a).
Indothais lacera (Born, 1778). Also reported as: Cuma lacera (Born, 1778); Thais lacera (Born, 1778); Thais
carinifera (Lamarck, 1822); and Thais mutabilis (Link, 1807). Gulf of Thailand, Andaman Sea. Reported by: KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991, 1992), Middelfart (1997), Carpenter
& Niem (1998), Swennen et al. (2001), Robba et al. (2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b).
Indothais malayensis (Tan & Sigurdsson, 1996). Also reported as: Thais malayensis (Tan, 1996). Gulf of Thailand,
Andaman Sea. Reported by: Tan & Sigurdsson (1996), Middelfart (1997), Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Indothais rufotincta (Tan & Sigurdsson, 1996). Also reported as: Thais rufotincta (Tan & Sigurdsson, 1996).
Gulf of Thailand, Andaman Sea. Reported by: Middelfart (1997), Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Indothais sacellum (Gmelin, 1791). Also reported as: Thais sacellum (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: Middelfart (1997), Gemert (2003), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019a).
Lataxiena blosvillei (Deshayes, 1832). Also reported as: Bedeva blosvillei (Deshayes, 1832). Gulf of Thailand,
Andaman Sea. Reported by: Bussarawit (1991, 1992), Middelfart (1997), Gemert (2003), Robba et al. (2004),
Nabhitabhata (2009), Sanpanich & Duangdee (2013), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Lataxiena cf. bombayana (Melvill, 1893). Reported as: Muricopsis cf. bombayanus (Melvill, 1893). Andaman Sea.
Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991), Carpenter & Niem (1998).
Lataxiena fimbriata (Hinds, 1844). Also reported as: Trophon fimbriatus. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, Gemert (2003), Robba et al. (2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich &
Duangdee (2013), GBIF (2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Lataxiena solenosteiroides Houart, Fraussen & Barbier, 2013. Andaman Sea. Reported by: Houart et al. (2013),
GBIF (2019a).
Leptoconchus incrassa A. Gittenberger & E. Gittenberger, 2011. Andaman Sea. Reported by: Gittenberger &
Gittenberger (2011).
Leptoconchus massini A. Gittenberger & E. Gittenberger, 2011. Andaman Sea. Reported by: Gittenberger &
Gittenberger (2011).
Maculotriton serriale (Deshayes, 1834). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Bussarawit (1991), Middelfart (1997), Duangdee et al. (2013b), BEDO (2017b).
Mancinella alouina (Röding, 1798). Also reported as: Mancinella mancinella (Linnaeus, 1758); Thais mancinella
(Linnaeus, 1758); and Thais alouina (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: THNHM,
KUMF, Nateewathana et al. (1981), Bussarawit (1991, 1995), Middelfart (1997), Carpenter & Niem (1998),
Duangdee (2004), Wells (2005), Duangdee et al. (2007, 2009, 2011), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Mancinella armigera Link, 1807. Reported as: Thais armigera (Link, 1907). Andaman Sea. Reported by: THNHM,
KUMF, Tantanasiriwong (1978), Middelfart (1997), Carpenter & Niem (1998), Nabhitabhata (2009), BEDO
(2017b).
Mancinella echinata (Blainville, 1832). Reported as: Thais echinata (Blainville, 1832); and Purpura (Mancinella)
echinata. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978),
Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1991, 1995), Middelfart (1997), Gemert (2003),
Duangdee (2004), Wells (2005), Kasetsart University (2006), ONR & EPP (2006), Duangdee (2007), Duangdee
et al. (2007, 2009, 2011, 2013b), ONEP (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013), Hylleberg & Aungtonya (2013), BEDO (2017b), GBIF (2019a, 2019b).
Mancinella echinulata (Lamarck, 1822). Also reported as: Thais echinulata (Lamarck, 1822). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Kostina et al. (2016), BEDO (2017b), GBIF
(2019a).
Menathais intermedia (Kiener, 1836). Reported as: Thais intermedia (Kiener, 1835). Gulf of Thailand, Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997),
Nabhitabhata (2009), BEDO (2017b).
Menathais tuberosa (Röding, 1798). Also reported as: Thais tuberosa (Röding, 1798); Mancinella tuberosa; and
Mancinella (Menethais) tuberosa (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Bussarawit
(1991), Middelfart (1997), Carpenter & Niem (1998), Duangdee (2004), Wells (2005), Nabhitabhata (2009),
Duangdee et al. (2011), Sørensen (2012), Sanpanich & Duangdee (2013), GBIF (2019a).
Morula biconica (Blainville, 1832). Also reported as: Cronia biconica (Blainville, 1832); and Morula
(Habromorula) biconica (Blainville, 1832). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1991),
Duangdee (2004, 2007), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011,
2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018).
Morula nodicostata (Pease, 1868). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Wells (2005), Nabhitabhata (2009), BEDO
(2017b), GBIF (2019a).
Morula spinosa (H. Adams & A. Adams, 1855). Also reported as: Habromorula spinosa (H. Adams & A. Adams,
1855); and Morula (Habromorula) spinosa (H. Adams & A. Adams, 1853). Gulf of Thailand, Andaman Sea. Reported
by: Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Gemert (2003), Houart (2004), Wells
(2005), Sanpanich & Duangdee (2009, 2013, 2018), Duangdee et al. (2011), BEDO (2017b), GBIF (2019a, 2019b).
Morula uva (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Middelfart (1997), Wells (2005), GBIF
(2019a).
Murex aduncospinosus Sowerby, 1841. Also reported as: Murex (Murex) aduncospinosus Sowerby, 1841. Gulf of
Thailand, Andaman Sea. Reported by: Ponder & Vokes (1988), Bussarawit (1992), Gemert (2003), Merle et al.
(2011), Houart (2014), J. D. Taylor et al. (unpublished data).
Murex altispira Ponder & Vokes, 1988. Also reported as: Murex (Murex) altispira Ponder & Vokes, 1988. Gulf of
Thailand, Andaman Sea. Reported by: Ponder & Vokes (1988), Bussarawit (1991), Swennen et al. (2001),
Nabhitabhata (2009), Houart (2010, 2014), Merle et al. (2011), GBIF (2019b).
Murex carbonnieri (Jousseaume, 1881). Andaman Sea. Reported by: KUMF, D’Asaro (1992), Middelfart (1997),
Nabhitabhata (2009), BEDO (2017b).
Murex concinnus Reeve, 1845. Gulf of Thailand. Reported by: Gemert (2003).
Murex djarianensis K. Martin, 1895. Also reported as: Murex djarianensis poppei Houart, 1979; Murex (Murex)
djarianensis poppei Houart, 1979; and Murex djarianensis (Deshayes, 1844). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Ponder & Vokes (1988), Bussarawit (1991, 1992), Middelfart (1997), Aungtonya & Hylleberg
(1998), Kasetsart University (2000, 2006), Duangdee (2004), ONR & EPP (2006), Duangdee et al. (2007, 2009,
2011), ONEP (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Merle et al. (2011), Sanpanich
& Duangdee (2013), Houart (2014), BEDO (2017b), GBIF (2019a, 2019b).
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The Marine and Estuarine Molluscs of Thailand
Murex occa G. B. Sowerby II, 1834. Also reported as: Murex (Murex) occa Sowerby, 1834. Gulf of Thailand,
Andaman Sea. Reported by: Ponder & Vokes (1988), Bussarawit (1991, 1992), Middelfart (1996, 1997), Subba
Rao & Dey (2000), Swennen et al. (2001), Nabhitabhata (2009), Houart (2010, 2014), Merle et al. (2011), BEDO
(2017b), GBIF (2019a, 2019b).
Murex pecten Lightfoot, 1786. Also reported as: Murex ternispina Lamarck, 1822. Gulf of Thailand, Andaman Sea.
Reported by: Bussarawit (1991, 1992), Middelfart (1997), Carpenter & Niem (1998), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009).
Murex scolopax Dillwyn, 1817. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Murex spectabilis Ponder & Vokes, 1988. Andaman Sea. Reported by: Merle et al. (2011).
Murex tenuirostrum Lamarck, 1822. Gulf of Thailand, Andaman Sea. Reported by: Ponder & Vokes (1988),
Bussarawit (1991), Subba Rao & Dey (2000), Merle et al. (2011).
Murex ternispina Lamarck, 1822. Gulf of Thailand. Reported by: Subba Rao & Dey (2000), GBIF (2019b).
Murex trapa Röding, 1798. Also reported as: Murex martinianus Reeve, 1845; and Murex (Murex) trapa. Gulf of
Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Tantanasiriwong (1978), Nateewathana
et al. (1981), Ponder & Vokes (1988), Kurozumi et al. (1989), Bussarawit (1991, 1992, 1995), Middelfart (1997),
Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Kasetsart University (2000), Subba Rao & Dey (2000),
Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), Robba et al. (2004), Duangdee et al. (2007, 2009,
2011), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Taylor (2009), Sanpanich & Duangdee (2013),
Houart (2014), BEDO (2017b), GBIF (2019b).
Murex tribulus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Ponder & Vokes (1988),
Bussarawit (1991), Carpenter & Niem (1998), Nabhitabhata (2009), Duangdee et al. (2011), Merle et al. (2011),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Murex troscheli Lischke, 1868. Andaman Sea. Reported by: Gemert (2003), Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich & Duangdee (2013, 2018).
Murichorda fiscellum (Gmelin, 1791). Reported as: Morula fiscella (Gmelin, 1791). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Nabhitabhata
(2009), BEDO (2017b).
Muricodrupa anaxares (Kiener, 1836). Reported as: Morula anaxares (Kiener, 1836); and Morula anexares. Gulf
of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1991, 1995), Middelfart (1997), Kasetsart University (2006), ONEP (2007), Thamrongnawasawat et al. (2008),
Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sørensen (2012), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Naquetia barclayi (Reeve, 1858). Andaman Sea. Reported by: Bussarawit (1991).
Naquetia triqueter (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1992), Middelfart
(1997), Wells (2005), GBIF (2019a).
Nassa francolina (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1991), Middelfart (1997), Carpenter & Niem (1998), Wells (2005),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2013a), BEDO (2017b).
Nassa serta (Bruguière, 1798). No Thai locality provided. Reported by: Bussarawit (1991), Carpenter & Niem
(1998), Nabhitabhata (2009).
Neothais marginatra (Blainville, 1832). Reported as: Semiricinula marginatra; and Thais marginatra. Andaman
Sea. Reported by: Duangdee et al. (2009), Sørensen (2012).
Nucella emarginata (Deshayes, 1839). Reported as: Thais marginata (Blainville, 1832). Gulf of Thailand. Reported
by: Nabhitabhata (2009), Duangdee et al. (2011).
Oppomorus funiculatus (Reeve, 1846). Reported as: Morula funiculus (Wood, 1828). Andaman Sea. Reported by:
Bussarawit (1991).
Oppomorus purpureocinctus (Preston, 1909). Reported as: Morula (Morula) purpureocincta (Preston, 1909).
Andaman Sea. Reported by: Houart (2002), GBIF (2019a).
Orania ficula (Reeve, 1848). Also reported as: Lataxiena ficula (Reeve, 1848). Gulf of Thailand, Andaman Sea.
Reported by: Gemert (2003), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
GBIF (2019b).
Orania livida (Reeve, 1846). Andaman Sea. Reported by: Middelfart (1997).
Orania nodulosus (Pease, 1869). Reported as: Ergalatax nodulosus (Pease, 1869). Andaman Sea. Reported by:
Bussarawit (1991).
Pascula ochrostoma (Blainville, 1832). Reported as: Drupella ochrostoma (Blainville, 1832). Andaman Sea.
Reported by: Nateewathana et al. (1981), Bussarawit (1991), Hylleberg & Aungtonya (2013).
Pterynotus alatus (Röding, 1798). Also reported as: Pterynotus pinnatus (Swainson, 1822). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991),
Middelfart (1997), Nabhitabhata (2009), BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Pterynotus barclayanus (H. Adams, 1873). Reported as: Marchia purpureus (Azuma, 1976). Andaman Sea.
Reported by: Bussarawit (1991).
Pterynotus bipinnatus (Reeve, 1845). Also reported as: Pterymarchia bipinnata (Reeve, 1845). Andaman Sea.
Reported by: Wells (2005), GBIF (2019a).
Pterynotus pellucidus (Reeve, 1845). Reported as: Pterymarchia pellucida (Reeve, 1845). Andaman Sea. Reported
by: THNHM.
Pterynotus tripterus (Born, 1778). Reported as: Pterymarchia triptera (Born, 1778). Andaman Sea. Reported by:
Middelfart (1997), BEDO (2017b), GBIF (2019a).
Purpura bufo Lamarck, 1822. Reported as: Thais bufo (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported
by: Morlet (1889), Bussarawit (1991), Carpenter & Niem (1998), Nabhitabhata (2009), J. D. Taylor et al.
(unpublished data).
Purpura persica (Linnaeus, 1758). Also reported as: Purpura (Mancinella) persica. Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991,
1995), Middelfart (1997), Carpenter & Niem (1998), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
BEDO (2017b), GBIF (2019a).
Rapana rapiformis (Born, 1778). Also reported as: Rapana repiformis. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991, 1995), Middelfart
(1997), Carpenter & Niem (1998), Swennen et al. (2001), Duangdee et al. (2007, 2011), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Reishia bitubercularis (Lamarck, 1822). Reported as: Purpura undata Lamarck, 1822; Thais bitubercularis
(Lamarck, 1822); Mancinella bitubercularis (Lamarck, 1822); Purpura kieneri (Deshayes, 1844); and Thais kieneri
(Deshayes, 1844). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Tantanasiriwong (1978),
Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit (1991), Middelfart (1996, 1997), Kasetsart
University (2000, 2006), Swennen et al. (2001), Duangdee (2004), ONR & EPP (2006), Duangdee et al. (2007,
2009, 2011), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013),
Sørensen (2012), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Reishia bronni (Dunker, 1860). Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Reishia clavigera (Küster, 1860). Reported as: Thais clavigera. Gulf of Thailand. Reported by: Samakraman et al.
(2010).
Reishia luteostoma (Holten, 1803). Andaman Sea. Reported by: Morlet (1889), BEDO (2017b).
Semiricinula fusca (Küster, 1862). Reported as: Morula fusca (Kuster); and Morulina fusca (Kuster). Gulf of
Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991),
Nabhitabhata (2009), BEDO (2017b).
Semiricinula konkanensis (Melvill, 1893). Andaman Sea. Reported by: Raven (2016).
Semiricinula muricina (Blainville, 1832). Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich &
Duangdee (2013).
Semiricinula muricoides (Blainville, 1832). Also reported as: Thais muricoides (Blainville, 1832). Andaman Sea.
Reported by: Gemert (2003), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Semiricinula squamosa (Pease, 1868). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013), GBIF (2019a, 2019b).
Semiricinula tissoti (Petit de la Saussaye, 1852). Reported as: Thais tissoti (Petit de la Saussaye, 1852). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, Middelfart (1996), Nabhitabhata (2009), GBIF (2019b).
Semiricinula turbinoides (Blainville, 1832). Andaman Sea. Reported by: GBIF (2019a).
Spinidrupa infans (Smith, 1884). Reported as: Muricopsis infans (E. A. Smith, 1884). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Nabhitabhata (2009), BEDO (2017b).
Stramonita biserialis (Blainville, 1832). Reported as: Purpura bicostalis Lamarck, 1816. Gulf of Thailand. Reported
by: Morlet (1889).
Stramonita delessertiana (d’Orbigny, 1841). Reported as: Purpura blainvillei Deshayes, 1844. Gulf of Thailand.
Reported by: Morlet (1889).
Taurasia striata (Blainville, 1832). Andaman Sea. Reported by: Raven (2016).
Tenguella ceylonica (Dall, 1923). Reported as: Morula ceylonica (Dall, 1923). Andaman Sea. Reported by: J. D.
Taylor et al. (unpublished data).
Tenguella granulata (Duclos, 1832). Also reported as: Morula granulata (Duclos, 1832). Gulf of Thailand, Andaman
Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991, 1992),
Middelfart (1996, 1997), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), Wells (2005), Kasetsart
University (2006), Printrakoon et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee
et al. (2011), Sørensen (2012), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b), J. D.
Taylor et al. (unpublished data).
Tenguella marginalba (Blainville, 1832). Reported as: Morula marginalba (Blainville, 1832). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991, 1995),
Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013).
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The Marine and Estuarine Molluscs of Thailand
Tenguella musiva (Kiener, 1835). Reported as: Morula musiva (Kiener, 1835). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Kurozumi et al. (1989), Bussarawit (1992), Middelfart (1997), Gemert (2003),
Duangdee (2004, 2007), Kasetsart University (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Sørensen
(2012), Kostina et al. (2016), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Thais costata (Blainville, 1832). Gulf of Thailand. Reported by: Robba et al. (2004). Reported by WoRMS Editorial
Board (2019) as a taxon inquirendum.
Tylothais aculeata (Deshayes, 1844). Reported as: Thalessa aculeata (Deshayes, 1844); and Thais aculeata (Link,
1807). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Kurozumi et al. (1989), Bussarawit (1991),
Carpenter & Niem (1998), Kasetsart University (2000, 2006), Duangdee (2004), Wells (2005), ONR & EPP
(2006), Duangdee et al. (2007, 2011, 2013b), ONEP (2007), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), Kostina et al. (2016), GBIF (2019a).
Tylothais savignyi (Deshayes, 1844). Reported as: Thais savignyi (Deshayes, 1844). Andaman Sea. Reported by:
Way & Purchon (1981).
Tylothais virgata (Dillwyn, 1817). Reported as: Thais distinguenda (Dunker, 1866); Thais virgatus; Thalessa
virgata (Dillwyn, 1817); Mancinella virgatus (Dillwyn, 1817); and Mancinella (Menethais) distinguenda (Dunker
& Zelebor). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1991), Middelfart (1997), Bech (1998),
Duangdee (2004), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2009), Nabhitabhata
(2009), Sørensen (2012), BEDO (2017b), Houart (2017), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Vexilla vexillum (Gmelin, 1791). Also reported as: Purpura vexillum (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Carpenter & Niem (1998), Nabhitabhata (2009).
Vokesimurex dolichourus (Ponder & Vokes, 1988). Reported as: Haustellum dolichourus Ponder & Vokes, 1988.
Gulf of Thailand, Andaman Sea. Reported by: Ponder & Vokes (1988), Bussarawit (1991), Middelfart (1997),
Aungtonya & Hylleberg (1998), Nabhitabhata (2009), Merle et al. (2011), Houart (2014), BEDO (2017b), GBIF
(2019a).
Vokesimurex gallinago (G. B. Sowerby III, 1903). Reported as: Haustellum gallinago (G. B. Sowerby III, 1903).
No Thai locality provided. Reported by: Higo et al. (1999).
Vokesimurex kiiensis (Kira, 1959). Reported as: Murex kiiensis (Kuroda, 1959); and Murex kiiensis (Kira). Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1991), Nabhitabhata (2009).
Vokesimurex rectirostris (G. B. Sowerby II, 1841). Andaman Sea. Reported by: GBIF (2019a).
Superfamily Olivoidea
Family Ancillariidae
Ancilla ampla (Gmelin, 1791). Also reported as: Ancilla ampla cylindrica (G. B. Sowerby II, 1859). Andaman Sea.
Reported by: BEDO (2017b).
Turrancilla glans (E. A. Smith, 1899). Andaman Sea. Reported by: GBIF (2019a).
Family Olividae
Agaronia gibbosa (Born, 1778). Also reported as: Oliva gibbosa (Born, 1778); and Olivancillaria gibbosa (Born,
1778). Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1938), Tantanasiriwong (1978), Nateewathana
et al. (1981), Way & Purchon (1981), Kasetsart University (2000), Subba Rao & Dey (2000), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a), Tan et al. (2019).
Felicioliva kaleontina (Duclos, 1835). Reported as: Oliva kaleontina Duclos, 1835. Gulf of Thailand. Reported by:
Morlet (1889).
Oliva amethystina (Röding, 1798). Andaman Sea. Reported by: BEDO (2017b).
Oliva cf. angustata Marrat, 1868. Also reported as: Oliva scripta Lamarck, 1811. Andaman Sea. Reported by:
Burch & Burch (1967), Raven & Recourt (2018).
Oliva annulata (Gmelin, 1791). Also reported as: Oliva (Annulatoliva) annulata forma mantichora Duclos, 1840.
Gulf of Thailand, Andaman Sea. Reported by: da Motta (1981), Carpenter & Niem (1998), Wells (2005), Hunon
et al. (2009), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011). Listed by
WoRMS Editorial Board (2019) as a nomen dubium.
Oliva buelowi G. B. Sowerby III, 1889. Reported as: Oliva buloui (Sowerby, 1887). Gulf of Thailand, Andaman Sea.
Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Hunon et al. (2009),
Nabhitabhata (2009), Taylor (2009), Galli (2015), BEDO (2017b), Raven & Recourt (2018). GBIF (2019a) lists
the subspecies Oliva buelowi phuketensis Tursch, Germain & Greifeneder, 1986.
Oliva caerulea (Röding, 1798). Also reported as: Oliva (Miniaceoliva) caerulea (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: Carpenter & Niem (1998), Gemert (2003), Wells (2005), Hunon et al. (2009),
Nabhitabhata (2009), BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Oliva carneola (Gmelin, 1791). Reported as: Oliva carneola (Gmelin, 1791) forma andamanensis Bridgman, 1909.
Andaman Sea. Reported by: da Motta (1981), Sterba (2004).
Oliva concavospira G. B. Sowerby III, 1914. Gulf of Thailand. Reported by: THNHM.
Oliva dubia Schepman, 1904. Andaman Sea. Reported by: BEDO (2017b).
Oliva elegans Lamarck, 1811. Andaman Sea. Reported by: BEDO (2017b).
Oliva guttata Fischer von Waldheim, 1808. Reported as: Oliva guttata (Röding, 1798). Gulf of Thailand, Andaman
Sea. Reported by: Wells (2005), Sanpanich & Duangdee (2018).
Oliva hanleyorum Petuch & Sargent, 1986. Also reported as: Oliva indomalaysica (Petuch & Sargent, 1986); Oliva
vidua macleay Duclos, 1840; and Oliva (Viduoliva) indomalaysica (Petuch & Sargent, 1986). Gulf of Thailand,
Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Sterba (2004), Hunon et al. (2009), Nabhitabhata (2009),
Galli (2015), Raven & Recourt (2018).
Oliva hirasei Kuroda & Habe, 1952. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2007, 2011),
Nabhitabhata (2009), Sanpanich & Duangdee (2013), GBIF (2019b).
Oliva irisans Lamarck, 1811. Reported as: Oliva (Miniaceoliva) concinna kremerorum Petuch & Sargent, 1986.
Andaman Sea. Reported by: Hunon et al. (2009).
Oliva mantichora Duclos, 1840. Andaman Sea. Reported by: Raven & Recourt (2018), GBIF (2019a).
Oliva miniacea (Röding, 1798). Also reported as: Miniaceoliva (Miniaceoliva) miniacea miniacea (Röding, 1798).
Gulf of Thailand, Andaman Sea. Reported by: KUMF, da Motta (1981), Tursch & Greifeneder (1996), Carpenter
& Niem (1998), Swennen et al. (2001), Hunon et al. (2009), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2013), Duangdee et al. (2011), GBIF (2019, 2019b).
Oliva mucronata Marrat, 1871. Andaman Sea. Reported by: THNHM.
Oliva mustelina Lamarck, 1811. Gulf of Thailand, Andaman Sea. Reported by: Suvatti (1938), Tursch & Machbaete
(1995), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Oliva neostina Duclos, 1840. Gulf of Thailand. Reported by: Morlet (1889).
Oliva oliva (Linnaeus, 1758). Also reported as: Oliva (Oliva) oliva forma gratiosa Vanatta, 1915. Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Tursch et al. (1992),
Tursch & Machbaete (1995), Carpenter & Niem (1998), Swennen et al. (2001), Duangdee (2004), Sterba (2004),
Kasetsart University (2006), Duangdee et al. (2007, 2011), ONEP (2007), Hunon et al. (2009), Nabhitabhata
(2009), Sanpanich & Duangdee (2013), BEDO (2017b), Raven & Recourt (2018), GBIF (2019a, 2019b).
Oliva ornata Marrat, 1867. Also reported as: Oliva lignaria. Gulf of Thailand, Andaman Sea. Reported by: THNHM,
da Motta (1981).
Oliva ozodona Duclos, 1835. Reported as: Oliva (Omogymna) nitidula sandwicensis Pease, 1860. Andaman Sea.
Reported by: Sterba (2004), Hunon et al. (2009).
Oliva pacifica Marrat, 1870. Also reported as: Oliva (Arctoliva) pacifica Marrat, 1870; Oliva ziegleri da Motta
(1981); and Oliva arctata (Marrat, 1871). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
da Motta (1981), Nateewathana et al. (1981), Bussarawit (1995), Tursch & Machbaete (1995), Sterba (2004),
Hunon et al. (2009), Nabhitabhata (2009), Galli (2015), BEDO (2017b), GBIF (2019a).
Oliva panniculata Duclos, 1835. Andaman Sea. Reported by: Burch & Burch (1967).
Oliva picta Reeve, 1850. Andaman Sea. Reported by: Galli (2015).
Oliva reticulata (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Carpenter & Niem (1998),
Nabhitabhata (2009), GBIF (2019a).
Oliva scripta Lamarck, 1811. Andaman Sea. Reported by: da Motta (1981).
Oliva sericea (Röding, 1798). Also reported as: Oliva (Miniacea) sericea (Röding, 1798). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Hunon et al. (2009),
Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Oliva sidelia Duclos, 1840. Reported as: Oliva faba Marrat, 1867; and Oliva smithi Bridgman, 1906. Andaman Sea.
Reported by: Tursch & Machbaete (1995), Sterba (2004), Hunon et al. (2009).
Oliva tigridella Duclos, 1835. No Thai locality given. Reported by: Hunon et al. (2009).
Oliva todosina Duclos, 1840. Also reported as: Oliva lepida Duclos, 1840; and Oliva (Galeola) todosina Duclos,
1840. Gulf of Thailand, Andaman Sea. Reported by: Sterba (2004), Hunon et al. (2009), Raven & Recourt (2018),
GBIF (2019a, 2019b).
Oliva tricolor Lamarck, 1811. Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata
(2009), GBIF (2019a).
Oliva vidua (Röding, 1798). Andaman Sea. Reported by: THNHM, da Motta (1981), Way & Purchon (1981), BEDO
(2017b), GBIF (2019a).
Olivella micula (Marrat, 1871). Also reported as: Olivella cf. plana (Marrat, 1871). Gulf of Thailand. Reported by:
Swennen et al. (2001), Raven & Recourt (2018).
Olivella spretoides Yokoyama, 1922. Gulf of Thailand. Reported by: Robba et al. (2004).
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The Marine and Estuarine Molluscs of Thailand
Superfamily Turbinelloidea
Family Costellariidae
Atlantilux rubra (Broderip, 1836). Andaman Sea. Reported by: BEDO (2017e).
Pusia microzonias (Lamarck, 1811). Reported as: Vexillum microzonias (Lamarck, 1811). Andaman Sea. Reported
by: Wells (2005).
Thala exilis (Reeve, 1845). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Vexillum acuminatum (Gmelin, 1791). Also reported as: Mitra crebrilirata Reeve, 1844. Andaman Sea. Reported
by: Morlet (1889), Nateewathana et al. (1981), Wells (2005), BEDO (2017b), GBIF (2019a).
Vexillum adamsi (Dohrn, 1861). Reported as: Pusia adamsi (Dohrn, 1861). Andaman Sea. Reported by: BEDO
(2017b).
Vexillum cadaverosum (Reeve, 1844). Andaman Sea. Reported by: Wells (2005), Duangdee et al. (2013b).
Vexillum citrinum (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019a).
Vexillum coccineum (Reeve, 1844). Gulf of Thailand. Reported by: GBIF (2019b).
Vexillum curviliratum (G. B. Sowerby II, 1874). Also reported as: Vexillum (Vexillum) curviliratum (G. B. Sowerby
II & III, 1874). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017b), GBIF (2019a, 2019b).
Vexillum delicatum (A. Adams, 1853). Gulf of Thailand. Reported by: GBIF (2019b).
Vexillum dilectissimum (Melvill & Sykes, 1899). Reported as: Costellaria dilectissimum Melvill & Standen, 1899.
Andaman Sea. Reported by: Galli (2015).
Vexillum discolorium (Reeve, 1845). Reported as: Vexillum coronatum (Lamarck, 1811). Andaman Sea. Reported
by: Wells (2005), BEDO (2017e).
Vexillum festum (Reeve, 1845). Also reported as: Vexillum (Pusia) festum (Reeve, 1845). Andaman Sea. Reported
by: Nateewathana et al. (1981), Gemert (2003), Wells (2005), BEDO (2017b).
Vexillum formosense (G. B. Sowerby III, 1889). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019a).
Vexillum funereum (Reeve, 1844). Andaman Sea. Reported by: J. D. Taylor et al. (unpublished data).
Vexillum lyratum (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Galli
(2015), GBIF (2019b).
Vexillum malcolmense (Melvill & Standen, 1901). Reported as: Vexillum (Costellaria) malcolmense (Melvill &
Standen, 1901). Gulf of Thailand. Reported by: Robba et al. (2007).
Vexillum mica (Reeve, 1845). Reported as: Vexillum lyratum (Lamarck, 1811). Andaman Sea. Reported by: Kilburn
& Hylleberg (1998), Nabhitabhata (2009).
Vexillum mirabile (A. Adams, 1853). Gulf of Thailand. Reported as: Mitra pica (Dillwyn, 1817). Reported by:
Kasetsart University (2006), ONEP (2007), Nabhitabhata (2009), Sørensen (2012).
Vexillum obeliscus (Reeve, 1844). Reported as: Vexillum (Costellaria) obeliscus (Reeve, 1844). Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Vexillum plicarium (Linnaeus, 1758). Reported as: Vexillum (Vexillum) plicarium (Linnaeus, 1758). Andaman
Sea. Reported by: Wells (2005), Sanpanich & Duangdee (2013), GBIF (2019a).
Vexillum radix (G. B. Sowerby II, 1874). Reported as: Vexillum (Costellaria) radix (G. B. Sowerby II, 1874). Gulf
of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Vexillum rugosum (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009).
Vexillum cf. rusticum (Reeve, 1845). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Vexillum sanguisuga (Linnaeus, 1758). Andaman Sea. Reported by: BEDO (2017b).
Vexillum scitulum (A. Adams, 1853). Andaman Sea. Reported by: BEDO (2017b), GBIF (2019a).
Vexillum subdivisum (Gmelin, 1791). Andaman Sea. Reported by: Wells (2005), Nabhitabhata (2009).
Vexillum taeniatum (Lamarck, 1811). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al.
(1981), Swennen et al. (2001), BEDO (2017b), GBIF (2019a, 2019b).
Vexillum vulpecula (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Vexillum woldemarii (Kiener, 1838). Reported as: Zierliana woldemarii (Kiener, 1838). Gulf of Thailand, Andaman
Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Sørensen (2012), BEDO (2017b).
Family Turbinellidae
Vasum ceramicum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Wells (2005), Nabhitabhata (2009).
Vasum turbinellus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al.
(1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Duangdee (2004),
KURDI et al. (2004), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP
(2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018), BEDO
(2017b), GBIF (2019a).
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Superfamily Volutoidea
Family Cancellariidae
Merica asperella (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Robba et al. (2007),
Nabhitabhata (2009).
Merica elegans (G. B. Sowerby I, 1822). Reported as: Cancellaria elegans Sowerby, 1822. Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Robba et al. (2004, 2007), BEDO (2017b).
Merica oblonga (G. B. Sowerby I, 1825). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017b).
Merica sinensis (Reeve, 1856). Reported as: Cancellaria sinensis (Reeve, 1856). Gulf of Thailand, Andaman Sea.
Reported by: Nabhitabhata (2009).
Scalptia crenifera (G. B. Sowerby I, 1832). Also reported as: Trigonostoma crenifera (Sowerby). Andaman Sea.
Reported by: Suvatti (1938), Nateewathana et al. (1981), Kilburn & Hylleberg (1998), Verhecken (2011), BEDO
(2017b).
Scalptia scalariformis (Lamarck, 1822). Gulf of Thailand. Reported by: Robba et al. (2004).
Scalptia textilis (Kiener, 1841). Andaman Sea. Reported by: BEDO (2017b).
Trigonostoma bicolor (Hinds, 1843). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998),
Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Tritonoharpa antiquata (Reeve, 1844). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Nabhitabhata
(2009).
Tritonoharpa pseudangasi Beu & P. A. Maxwell, 1987. No Thai locality given. Reported by: Verhecken (2011).
Family Cystiscidae
Crithe cossinea (T. Cossignani, 1997). Andaman Sea. Reported by: BEDO (2017e).
Cystiscus sandwicensis (Pease, 1860). Reported as: Gibberula sandwicensis (Pease, 1860). Gulf of Thailand.
Reported by: Gemert (2003).
Family Volutidae
Cymbiola nobilis (Lightfoot, 1786). Also reported as: Cymbiola (Cymbiola) nobilis (Lightfoot, 1786); and
Voluta scapha Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Nateewathana et al. (1981), Kasetsart University (2000, 2006), Swennen et al. (2001), Gemert (2003),
Duangdee (2004, 2007), Duangdee et al. (2007, 2011), ONEP (2007), Robba et al. (2007), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Cymbiola vespertilio (Linnaeus, 1758). Reported as: Voluta vespertilio (Linnaeus, 1758). Gulf of Thailand.
Reported by: Morlet (1889), Kurozumi et al. (1989).
Melo melo (Lightfoot, 1786). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Suvatti (1937, 1938),
Tantanasiriwong (1978), Nateewathana et al. (1981), Amornjaruchit (1988), Bussarawit (1995), Nateewathana
(1995), Aungtonya & Hylleberg (1998), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Swennen
et al. (2001), Duangdee (2004, 2007), Duangdee et al. (2007, 2011), ONEP (2007), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Superfamily [unassigned] Neogastropoda (temporary name)
Family Babyloniidae
Babylonia ambulacrum (G. B. Sowerby I, 1825). Also reported as: Babylonia canaliculata. Andaman Sea. Reported
by: THNHM, BEDO (2017b).
Babylonia areolata (Link, 1807). Also reported as: Babylonia areolata forma areolata (Link, 1807). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1937), Altena & Gittenberger (1981), Amornjaruchit (1988),
Nateewathana (1995), Carpenter & Niem (1998), Swennen et al. (2001), Duangdee et al. (2007, 2011), Robba
et al. (2007), Hualkasin et al. (2008), Nabhitabhata (2009), Fraussen & Stratmann (2013), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a, 2019b).
Babylonia lutosa (Lamarck, 1816). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Carpenter & Niem
(1998), Nabhitabhata (2009).
Babylonia perforata (G. B. Sowerby II, 1870). Andaman Sea. Reported by: Fraussen & Stratmann (2013).
Babylonia spirata (Linnaeus, 1758). Also reported as: Babylonia spirata forma spirata (Linnaeus, 1758) and
Babylonia canaliculata. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Nateewathana et al. (1981),
Nabhitabhata (2009), Duangdee et al. (2011), Fraussen & Stratmann (2013), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019b), J. D. Taylor et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Family Harpidae
Harpa amouretta Röding, 1798. Reported as: Harpa amouretta minor Lamarck, 1822. Andaman Sea. Reported
by: Hylleberg & Aungtonya (2013), BEDO (2017b).
Harpa articularis (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Rehder (1973),
Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Dance & Poppe (1999), Swennen
et al. (2001), Duangdee (2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b).
Harpa davidis Röding, 1798. Andaman Sea. Reported by: THNHM, Rehder (1973), Subba Rao & Dey (2000), Merle
& Pacaud (2003).
Harpa harpa (Linnaeus, 1758). Also reported as: Harpa hurpa. Gulf of Thailand, Andaman Sea. Reported by:
THNHM, Carpenter & Niem (1998), KURDI et al. (2004), Nabhitabhata (2009).
Harpa major (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019b).
Morum ninomiyai Emerson, 1986. Andaman Sea. Reported by: Emerson (1986), Galli (2015), BEDO (2017b),
GBIF (2019a).
Family Marginellidae
Cryptospira bridgettae Wakefield, 2010. Andaman Sea. Reported by: Wakefield (2010).
Cryptospira dactylus (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Cossignani (2006), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Cryptospira elegans (Gmelin, 1791). Also reported as: Persicula elegans Gmelin; and Marginella elegans. Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Aungtonya & Hylleberg
(1998), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Wakefield (2010), Sanpanich & Duangdee (2013),
Galli (2015), BEDO (2017b), GBIF (2019a).
Cryptospira fischeri (Bavay, 1903). Andaman Sea. Reported by: Wakefield (2010), BEDO (2017b).
Cryptospira glauca Jousseaume, 1875. Andaman Sea. Reported by: BEDO (2017b).
Cryptospira marchii Jousseaume, 1875. Andaman Sea. Reported by: Wakefield (2010).
Cryptospira mccleeryi Wakefield, 2010. Gulf of Thailand. Reported by: Wakefield (2010).
Cryptospira praecallosa (Higgins, 1876). Andaman Sea. Reported by: Wakefield (2010), Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Cryptospira strigata (Dillwyn, 1817). Also reported as: Cryptospira strigata forma unicolor; and Cryptospira
strigata unicolor Bert, 1985. Andaman Sea. Reported by: Cossignani (2006), Wakefield (2010), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), Galli (2015), GBIF (2019a).
Cryptospira tricincta (Hinds, 1844). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2013),
BEDO (2017b).
Cryptospira ventricosa (Fischer von Waldheim, 1807). Also reported as: Marginella ventricosa G. Fisher. Gulf
of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Swennen et al. (2001),
Nabhitabhata (2009), Wakefield (2010), BEDO (2017b), GBIF (2019a).
Heterocithara himerta (Melvill & Standen, 1896). Andaman Sea. Reported by: BEDO (2017e).
Volvarina angustata (G. B. Sowerby II, 1846). Reported as: Marginella angustata (Sowerby, 1846). Andaman
Sea. Reported by: Nateewathana et al. (1981), BEDO (2017b).
Volvarina hirasei (Bavay, 1917). Andaman Sea. Reported by: BEDO (2017e).
Volvarina philippinarum (Redfield, 1848). Gulf of Thailand. Reported by: Robba et al. (2004).
Order [unassigned] Caenogastropoda
Superfamily Campaniloidea
Family Plesiotrochidae
Plesiotrochus souverbianus P. Fischer, 1878. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO
(2017e).
Superfamily Cerithioidea
Family Batillariidae
Batillaria sordida (Gmelin, 1791). Reported as: Cerithium bornii G. B. Sowerby II, 1855. Gulf of Thailand. Reported
by: Morlet (1889).
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Family Cerithiidae
Bittium glareosum Gould, 1861. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Tanamai &
Nabhitabhata (2016), BEDO (2017e).
Bittium impendens (Hedley, 1899). Andaman Sea. Reported by: BEDO (2017e).
Bittium reticulatum (da Costa, 1778). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Bittium simplex (Jeffreys, 1867). Andaman Sea. Reported by: Sawatna et al. (2012), Tanamai & Nabhitabhata
(2016).
Cerithidium cerithinum (Philippi, 1849). Also reported as: Fenella cerithine (Philippi, 1848). Gulf of Thailand.
Reported by: Gemert (2003), Robba et al. (2004), Sawatna et al. (2012).
Cerithidium perparvulum (R. B. Watson, 1886). Gulf of Thailand. Reported by: Gemert (2003).
Cerithidium submammillatum (De Rayneval & Ponzi, 1854). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Cerithium alutaceum (Gould, 1861). Also reported as: Cerithium stigmosum Gould, 1861. Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Tanamai & Nabhitabhata (2016), BEDO (2017e).
Cerithium citrinum G. B. Sowerby II, 1855. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013), Kostina et al. (2016).
Cerithium columna G. B. Sowerby I, 1834. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Houbrick (1992), Subba Rao & Dey (2000), Nabhitabhata (2009), Duangdee et al. (2011), Hylleberg &
Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Cerithium coralium Kiener, 1841. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Suvatti (1938),
Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao & Dey (2000), Swennen
et al. (2001), Gemert (2003), Duangdee (2004), Wells (2005), Kasetsart University (2006), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011), Meyer et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013), BEDO (2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Cerithium dialeucum Philippi, 1849. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), GBIF (2019a,
2019b).
Cerithium echinatum (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Wells (2005), Kasetsart University (2006), ONEP (2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Cerithium lifuense Melvill & Standen, 1895. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011, 2013b), Sanpanich & Duangdee (2013).
Cerithium lissum R. B. Watson, 1880. Gulf of Thailand. Reported by: Gemert (2003), Nabhitabhata (2009).
Cerithium munitum G. B. Sowerby II, 1855. Gulf of Thailand. Reported by: Nabhitabhata (2009).
Cerithium nesioticum Pilsbry & Vanetta, 1906. Also reported as: Semivertagus nesioticus (Pilsbry & Vannatta,
1906). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Wells (2005), Nabhitabhata (2009),
Duangdee et al. (2013b).
Cerithium nodulosum Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet
(1889), Suvatti (1938), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao
& Dey (2000), Wells (2005), Kasetsart University (2006), ONR & EPP (2006), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a).
Cerithium novaehollandiae G. B. Sowerby II, 1855. Gulf of Thailand, Andaman Sea. Reported by: Houbrick (1992),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Cerithium punctatum Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: Subba Rao & Dey (2000),
Kostina et al. (2016), GBIF (2019a).
Cerithium rostratum Sowerby, 1855. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Cerithium salebrosum Sowerby, 1855. Andaman Sea. Reported by: Houbrick (1992), Wells (2005).
Cerithium tenellum Sowerby, 1855. Reported as: Cerithium tenuifilosum (Sowerby, 1866). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Wells (2005), Hylleberg & Aungtonya (2013), BEDO (2017b).
Cerithium torresi Smith, 1884. Gulf of Thailand, Andaman Sea. Reported by: Houbrick (1992), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011, 2013a), Sanpanich & Duangdee (2013), GBIF (2019a).
Cerithium traillii G. B. Sowerby II, 1855. Also reported as: Clypeomorus traillii (Sowerby). Gulf of Thailand,
Andaman Sea. Reported by: Suvatti (1938), Kurozumi et al. (1989), Subba Rao & Dey (2000), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013,
2018), BEDO (2017b).
Cerithium tuberculatum (Linnaeus, 1767). Reported as: Clypeomorus tuberculatus. Gulf of Thailand, Andaman
Sea. Reported by: Sørensen (2012).
Cerithium zebrum Kiener, 1841. Andaman Sea. Reported by: BEDO (2017b, 2017e).
Cerithium zonatum (Wood, 1828). Also reported as: Cerithium purpurascens (Sowerby). Gulf of Thailand,
Andaman Sea. Reported by: Houbrick (1985), Kurozumi et al. (1989), Subba Rao & Dey (2000), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011), Sørensen (2012), Sanpanich & Duangdee (2013), GBIF (2019a,
2019b).
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The Marine and Estuarine Molluscs of Thailand
Clypeomorus batillariaeformis (Habe & Kosuge, 1966). Also reported as: Clypeomorus moniliferus (Kiener,
1841). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981), Way & Purchon (1981),
Houbrick (1985), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Subba Rao & Dey (2000),
Gemert (2003), Duangdee (2004), Wells (2005), ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013, 2018), Kostina et al. (2016), BEDO (2017b), GBIF
(2019a, 2019b).
Clypeomorus bifasciata (G. B. Sowerby II, 1855). Reported as: Cerithium concisus (Hombron & Jacquinot, 1854)
and Clypeomorus humilis (Dunker). Gulf of Thailand, Andaman Sea. Reported by: KUMF, THNHM, Morlet (1889),
Houbrick (1985), Kurozumi et al. (1989), Subba Rao & Dey (2000), Swennen et al. (2001), Gemert (2003),
Duangdee (2004), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Clypeomorus brevis (Quoy & Gaimard, 1834). Also reported as: Cerithium breve Quoy & Gaimard, 1834. Gulf of
Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995), Nabhitabhata (2009).
Clypeomorus irrorata (Gould, 1849). Also reported as: Cerithium obesum Sowerby, 1855. Gulf of Thailand,
Andaman Sea. Reported by: Way & Purchon (1981), Houbrick (1985).
Clypeomorus nympha Houbrick, 1985. Gulf of Thailand. Reported by: GBIF (2019b).
Clypeomorus pellucida (Hombron & Jacquinot, 1848). Also reported as: Cerithium patulum Sowerby, 1855; and
Clypeomorus patulum (Sowerby). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Nateewathana et al. (1981),
Houbrick (1985), Bussarawit (1995), Duangdee (2004), Thamrongnawasawat et al. (2008), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), Dechruksa
et al. (2014), BEDO (2017b), J. D. Taylor et al. (unpublished data).
Clypeomorus petrosa (Wood, 1828). Also reported as: Clypeomorus petrosa chemnitzianum (Pilsbry, 1901); and
Cerithium chemnitzianum (Pilsbry, 1901). Gulf of Thailand, Andaman Sea. Reported by: Way & Purchon (1981),
Houbrick (1985), Kurozumi et al. (1989), Subba Rao & Dey (2000), Nabhitabhata (2009), Duangdee et al. (2011,
2013a), Sørensen (2012), Sanpanich & Duangdee (2013), GBIF (2019a).
Clypeomorus purpurastoma Houbrick, 1985. Gulf of Thailand, Andaman Sea. Reported by: Houbrick (1985),
Subba Rao & Dey (2000), Gemert (2003), Sørensen (2012), Duangdee et al. (2013b).
Pseudovertagus aluco (Linnaeus, 1758). Reported as: Vertagus aluco (Linnaeus, 1758). Gulf of Thailand. Reported
by: Morlet (1889).
Rhinoclavis articulata (Adams & Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Swennen et al. (2001), Duangdee et al. (2007, 2013), Nabhitabhata (2009), GBIF (2019a, 2019b).
Rhinoclavis aspera (Linnaeus, 1758). Also reported as: Rhinoclavis (Rhinoclavis) aspera (Linnaeus, 1758); and
Rhinoclavis asper (L., 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Houbrick (1978),
Way & Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao & Dey (2000), Gemert (2003),
Duangdee (2004), Wells (2005), Kasetsart University (2006), Duangdee et al. (2007, 2009, 2011, 2013b), ONEP
(2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018),
BEDO (2017b), GBIF (2019a).
Rhinoclavis fasciata (Bruguière, 1792). Also reported as: Rhinoclavis pharos (Hinds). Gulf of Thailand, Andaman
Sea. Reported by: Nateewathana et al. (1981), Duangdee (2004), Wells (2005), Nabhitabhata (2009), Duangdee
et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013).
Rhinoclavis kochi (Philippi, 1848). Also reported as: Cerithium kochi (Philippi, 1848); and Rhinoclavis (Provlavis)
kochi (Philippi, 1848). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Houbrick (1978), Subba Rao & Dey
(2000), Gemert (2003), Wells (2005), Nabhitabhata (2009), GBIF (2019b).
Rhinoclavis sinensis (Gmelin, 1791). Also reported as: Rhinoclavis (Rhinoclavis) sinensis (Gmelin, 1791); Vertagus
obeliscus (Bruguière, 1792); and Rhinoclavis obeliscus. Gulf of Thailand, Andaman Sea. Reported by: Morlet
(1889), Houbrick (1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Subba Rao & Dey (2000),
Gemert (2003), Duangdee (2004, 2007), Wells (2005), Kasetsart University (2006), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011, 2013b), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2013), Hylleberg & Aungtonya (2013), BEDO (2017b), GBIF (2019a, 2019b).
Rhinoclavis sordidula (Gould, 1849). Reported as: Cerithium sordidula (Gould, 1849); and Rhinoclavis (Provlavis)
sordidula (Gould, 1849). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Swennen
et al. (2001), Gemert (2003), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018),
BEDO (2017b).
Rhinoclavis vertagus (Linnaeus, 1767). Also reported as: Rhinoclavis (Rhinoclavis) vertagus (Linnaeus, 1767);
and Rhinoclavis vergatus (Linnaeus). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Houbrick
(1978), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998),
Subba Rao & Dey (2000), Gemert (2003), Kasetsart University (2006), ONEP (2007), Nabhitabhata (2009),
Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Family Dialidae
Diala albugo (R. B. Watson, 1886). Gulf of Thailand. Reported by: Gemert (2003).
Diala semistriata (Philippi, 1849). Gulf of Thailand, Andaman Sea. Reported by: Ponder & de Keyzer (1992),
Gemert (2003), Sawatna et al. (2012), BEDO (2017e), GBIF (2019a).
Diala sulcifera (A. Adams, 1862). Gulf of Thailand. Reported by: Gemert (2003).
Family Litiopidae
Alaba hungerfordi G. B. Sowerby III, 1894. Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004).
Litiopa nipponica Kuroda & Kawamoto, 1956. Gulf of Thailand. Reported by: Gemert (2003).
Styliferina goniochila A. Adams, 1860. Gulf of Thailand. Reported by: Gemert (2003).
Family Modulidae
Indomodulus tectum (Gmelin, 1791). Also reported as: Modulus tectum (Gmelin, 1792); and Modulus candidus
(Petit, 1853). Gulf of Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Gemert (2003),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), Kostina et al. (2016).
Family Pachychilidae
Faunus ater (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al. (2001), Sri-aroon et
al. (2005, 2006), Nabhitabhata (2009), GBIF (2019b).
Family Pickworthiidae
Mareleptopoma iredalei (Bavay, 1922). Reported as: Sansonia cf. iredalei (Bavay, 1922). Andaman Sea. Reported
by: BEDO (2017e).
Sansoniella minuta (Hornung & Mermod, 1927). Andaman Sea. Reported by: GBIF (2019a).
Family Planaxidae
Angiola fasciata (Pease, 1868). Andaman Sea. Reported by: GBIF (2019a).
Angiola lineata (da Costa, 1778). Reported as: Planaxis lineatus (da Costa, 1776). Gulf of Thailand, Andaman Sea.
Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009), BEDO (2017b), GBIF
(2019b).
Fossarus trochlearis A. Adams, 1853. Gulf of Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), BEDO
(2017e).
Hinea inepta (Gould, 1861). Gulf of Thailand, Andaman Sea. Reported by: Sawatna et al. (2012), BEDO (2017e).
Planaxis encausticus E. A. Smith, 1872. Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Nabhitabhata
(2009).
Planaxis sulcatus (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Suvatti (1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995),
Kilburn & Hylleberg (1998), Kasetsart University (2000, 2006), Swennen et al. (2001), Gemert (2003), Duangdee
(2004, 2007), ONR & EPP (2006), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Sanpanich
& Duangdee (2009, 2013), Samakraman et al. (2010), Sørensen (2012), Tan & Low (2014), Kostina et al. (2016),
BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Supplanaxis niger (Quoy & Gaimard, 1833). Also reported as: Planaxis nigra (Quoy & Gaimard, 1834); and Planaxis
niger. Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1995), Kilburn & Hylleberg (1998), Kasetsart University (2000), Gemert (2003), Nabhitabhata et al. (2009a),
Sørensen (2012), Kostina et al. (2016), BEDO (2017b), GBIF (2019a).
Family Potamididae
Cerithidea balteata A. Adams, 1855. Reported as: Cerithium balteatum Philippi, 1848. Gulf of Thailand, Andaman
Sea. Reported by: Houbrick (1992), Subba Rao & Dey (2000), Robba et al. (2004), Negri et al. (2014).
Cerithidea charbonnieri (Petit de la Saussaye, 1851). Also reported as: Aphanistylus charbonnieri (Petit de la
Saussaye, 1851). Gulf of Thailand. Reported by: Morlet (1889), Sri-aroon et al. (2005), Nabhitabhata (2009),
Reid (2014).
Cerithidea obtusa (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Suvatti (1937, 1938), Nateewathana et al. (1981), Amornjaruchit (1988), Nateewathana (1995), Carpenter &
Niem (1998), Kasetsart University (2000), Robba et al. (2004), Sri-aroon et al. (2005, 2006), Printrakoon et al.
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The Marine and Estuarine Molluscs of Thailand
(2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Reid (2014), BEDO (2017b), GBIF (2019a,
2019b), J. D. Taylor et al. (unpublished data).
Cerithidea quoyii (Hombron & Jacquinot, 1848). Also reported as: Cerithideopsilla quoyii (Hombron & Jacquinot,
1848); and Cerithidea quadrata G. B. Sowerby II, 1866. Gulf of Thailand, Andaman Sea. Reported by: THNHM,
Morlet (1889), Amornjaruchit (1988), Carpenter & Niem (1998), Swennen et al. (2001), Sri-aroon et al. (2005,
2006), Printrakoon et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Dechruksa et al.
(2014), Reid (2014), Ozawa et al. (2015), Reid & Ozawa (2016), BEDO (2017b), GBIF (2019b).
Cerithidea rhizoporarum (A. Adams, 1855). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Nateewathana (1995), Kasetsart University (2000), Nabhitabhata (2009).
Cerithidea weyersi Dautzenberg, 1899. Gulf of Thailand. Reported by: THNHM, Nabhitabhata (2009), Reid (2014).
Cerithideopsis malayensis D. Reid & Claremont, 2014. Reported by: Reid & Claremont (2014).
Pirenella alata (Philippi, 1849). Also reported as: Cerithideopsilla alata (Philippi, 1849); Cerithidea alata (Philippi,
1849); and Cerithidea djadjariensis (Martin, 1849). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart
University (2000), Sri-aroon et al. (2005, 2006), Meyer et al. (2008), Nabhitabhata (2009), Dechruksa et al.
(2014), Reid & Claremont (2014), Ozawa et al. (2015), Reid & Ozawa (2016), BEDO (2017b), GBIF (2019b).
Pirenella asiatica Ozawa & Reid, 2016. Gulf of Thailand. Reported by: Reid & Ozawa (2016).
Pirenella cingulata (Gmelin, 1791). Reported as: Cerithideopsilla cingulata (Gmelin, 1791); Tympanotonos
euryptera G. B. Sowerby II, 1866; Tympanotonos fluviatilis (Potiez & Michaud, 1838); Cerithidea cingulata
(Gmelin, 1791); and Cerithidea (Cerithideopsilla) cingulata (Gmelin, 1791). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Morlet (1889), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem
(1998), Kasetsart University (2000), Swennen et al. (2001), Gemert (2003), Duangdee (2004), Robba et al.
(2004), Sri-aroon et al. (2005, 2006), Duangdee et al. (2007, 2009, 2011), Meyer et al. (2008), Printrakoon et al.
(2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Dechruksa
et al. (2014), Negri et al. (2014), Reid & Claremont (2014), Ozawa et al. (2015), Premcharoen et al. (2016), Reid
& Ozawa (2016), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Pirenella incisa (Hombron & Jacquinot, 1848). Also reported as: Cerithideopsilla incisa (Hombron & Jacquinot,
1848). Gulf of Thailand, Andaman Sea. Reported by: Reid & Claremont (2014), Ozawa et al. (2015), Reid &
Ozawa (2016), BEDO (2017b).
Pirenella microptera (Kiener, 1841). Also reported as: Cerithidea microptera (Kiener, 1841). Gulf of Thailand.
Reported by: Gemert (2003), Nabhitabhata (2009), Reid & Ozawa (2016).
Telescopium telescopium (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Nateewathana
et al. (1981), Houbrick (1991), Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Kasetsart
University (2000), Subba Rao & Dey (2000), Swennen et al. (2001), Duangdee et al. (2007, 2011), Printrakoon
et al. (2008), Thamrongnawasawat et al. (2008), Sanpanich & Duangdee (2013), Reid & Claremont (2014),
BEDO (2017b), GBIF (2019b).
Terebralia palustris (Linnaeus, 1767). Gulf of Thailand, Andaman Sea. Reported by: Houbrick (1991),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee
(2013), Dechruksa et al. (2014), BEDO (2017b), GBIF (2019b).
Terebralia sulcata (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Family Scaliolidae
Finella pupoides Adams, 1860. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003). Robba et al. (2004),
Sawatna et al. (2012), BEDO (2017e), GBIF (2019a).
Finella purpureoapicata Preston, 1905. Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Robba
et al. (2004), BEDO (2017e).
Finella rufocincta (A. Adams, 1861). Gulf of Thailand. Reported by: BEDO (2017e).
Scaliola arenosa A. Adams, 1862. Gulf of Thailand. Reported by: Gemert (2003), BEDO (2017e).
Scaliola cf. bella A. Adams, 1860. Gulf of Thailand. Reported by: BEDO (2017e).
Scaliola glareosa A. Adams, 1862. Gulf of Thailand. Reported by: Robba et al. (2007), Negri et al. (2014), BEDO
(2017e).
Family Siliquariidae
Tenagodus anguinus (Linnaeus, 1758). Reported as: Siliquaria anguina (Linnaeus, 1758) and Tenagodus
(Tenagodus) anguinus (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2007).
Tenagodus cumingii Mörch, 1861. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009),
Nabhitabhata (2009), Duangdee et al. (2011), BEDO (2017b), GBIF (2019a).
Tenagodus ponderosus (Mörch, 1861). Also reported as: Siliquaria ponderosa (Mörch, 1860). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Duangdee (2004, 2007), ONEP (2007), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Tenagodus senegalensis (G. B. Sowerby II, 1876). Andaman Sea. Reported by: BEDO (2017b).
Tenagodus trochlearis Mörch, 1861. Andaman sea. Reported by: BEDO (2017b).
Tenagodus weldii Tenison Woods, 1876. Reported as: Siliquaria weldii (Tenison Woods, 1875). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011).
Family Thiaridae
Sermyla riqueti (Grateloup, 1840). Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004), Negri et al.
(2014).
Family Turritellidae
Turritella cingulifera G. B. Sowerby I, 1825. Reported as: Haustator (Kurosioia) cingulifer (Sowerby, 1825);
and Turritella fuscocinetata. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Robba et al. (2007),
Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019b).
Turritella duplicata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Nabhitabhata (2009), Taylor (2009), BEDO
(2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Turritella fastigiata Adams & Reeve in Reeve, 1849. Andaman Sea. Reported by: BEDO (2017b).
Turritella maculata Reeve, 1849. Reported as: Turritella vittulata (Adams & Reeve, 1848). Andaman Sea.
Reported by: Kilburn & Hylleberg (1998), Nabhitabhata (2009).
Turritella terebra (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Suvatti (1938), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University
(2000, 2006), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), Robba et al. (2004), ONR & EPP
(2006), Duangdee et al. (2007, 2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a, 2019b).
Superfamily Epitonioidea
Family Epitoniidae
Acrilla acuminata (G. B. Sowerby II, 1844). Reported as: Amaea (Acrilla) acuminata (Sowerby, 1844). Gulf of
Thailand. Reported by: Robba et al. (2004), GBIF (2019b).
Amaea grossicingulata (de Boury, 1913). Reported as: Amaea (Filiscala) grossicingulata (de Boury, 1912). Gulf
of Thailand. Reported by: Robba et al. (2004).
Amaea sulcata (G. B. Sowerby II, 1844). Reported as: Amaea (Amaea) sulcata (G. B. Sowerby II, 1844). Gulf of
Thailand. Reported by: Gemert (2003).
Cycloscala hyalina (G. B. Sowerby II, 1844). Gulf of Thailand. Reported by: Gemert (2003).
Eglisia brunnea Habe & Kosuge, 1966. No Thai locality given. Reported by: Weil et al. (1999).
Epidendrium aureum A. Gittenberger & E. Gittenberger, 2005. Andaman Sea. Reported by: A. Gittenberger &
E. Gittenberger (2005).
Epidendrium billeeanum (DuShane & Bratcher, 1965). Also reported as: Alora billeenae (DuShane & Bratcher,
1965); and Epitonium billeeanum (DuShane & Bratcher, 1965). Gulf of Thailand, Andaman Sea. Reported by:
ONR & EPP (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich
(2013), Sanpanich & Duangdee (2013, 2018).
Epifungium adgranulosa Gittenberger & Gittenberger, 2005. Andaman Sea. Reported by: Gittenberger &
Gittenberger (2005), Brown & Neville (2015).
Epifungium hartogi (A. Gittenberger, 2003). Also reported as Epitonium hartogi A. Gittenberger, 2003. Andaman
Sea. Reported by: Gittenberger (2003), Gittenberger & Gittenberger (2005), Brown & Neville (2015).
Epifungium lochi (A. Gittenberger & Goud, 2000). Andaman Sea. Reported by: Gittenberger & Gittenberger
(2005), Brown & Neville (2015).
Epifungium nielsi Gittenberger & Gittenberger, 2005. Andaman Sea. Reported by: Gittenberger & Gittenberger
(2005), Brown & Neville (2015).
Epifungium twilae (A. Gittenberger & Goud, 2000). Andaman Sea. Reported by: Gittenberger & Gittenberger
(2005), Brown & Neville (2015).
Epifungium ulu (Pilsbry, 1921). Andaman Sea. Reported by: Gittenberger & Gittenberger (2005), Brown & Neville
(2015).
Epitonium alizonae (Melvill, 1912). Reported as: Epitonium (Parviscala) alizonae (Melvill, 1912). Gulf of Thailand.
Reported by: Gemert (2003).
Epitonium angustum (Dunker, 1861). Reported as: Epitonium (Nitidiscala) angustum (Dunker, 1861). Gulf of
Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
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Epitonium catanuense (G. B. Sowerby, 1844). Reported as: Epitonium (Papyriscala) catanuense (G. B. Sowerby,
1844). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium columba Kilburn, 1985. Reported as: Epitonium (Parviscala) columba Kilburn, 1985. Gulf of Thailand.
Reported by: Robba et al. (2007).
Epitonium cookeanum Dall, 1917. Reported as: Epitonium? (Labeoscala) cookeanum Dall, 1917. Gulf of Thailand.
Reported by: Gemert (2003).
Epitonium fucatum (Pease, 1861). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), BEDO (2017b).
Epitonium glabratum (Hinds, 1843). Andaman Sea. Reported by: BEDO (2017b).
Epitonium goldsmithi (DuShane, 1988). Reported as: Asperiscala goldsmithi DuShane, 1988. Andaman Sea.
Reported by: DuShane (1988b), Brown & Neville (2015).
Epitonium gracile (G. B. Sowerby II, 1844). Reported as: Epitonium (Laeviscala) gracile (G. B. Sowerby II, 1844).
Gulf of Thailand. Reported by: Gemert (2003).
Epitonium hemmesi (DuShane, 1988). Andaman Sea. Reported by: BEDO (2017e).
Epitonium histricosum (Jousssaume, 1912). Reported as: Epitonium (Parviscala) histricosum (Jousseaume, 1912).
Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Epitonium immaculatum (G. B. Sowerby II, 1844). Reported as: Epitonium (Sodaliscala) immaculatum (G. B.
Sowerby II, 1844). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium imperiale (G. B. Sowerby, 1844). Reported as: Epitonium (Papyriscala) imperiale (Sowerby, 1844).
Gulf of Thailand. Reported by: THNHM, Robba et al. (2004), Sawatna et al. (2012).
Epitonium inexpertum L. G. Brown & Weil, 1999. Reported as: Epitonium (Sodaliscala) inexpertum L. G. Brown
& Weil, 1999. Gulf of Thailand. Reported by: Gemert (2003).
Epitonium irregulare (Sowerby II, 1844). Andaman Sea. Reported by: BEDO (2017b).
Epitonium japonicum (Dunker, 1861). Reported as: Epitonium (Hirtoscala) cf. japonicum (Dunker, 1861). Gulf
of Thailand. Reported by: Gemert (2003).
Epitonium linctum (de Boury & Monterosato, 1890). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Epitonium lineolatum (G. B. Sowerby, 1844). Reported as: Scalaria lineolata G. B. Sowerby, 1844. Gulf of Thailand.
Reported by: Morlet (1889).
Epitonium malhaensis (Jousseaume, 1894). Reported as: Epitonium (Papyriscala) malhaensis (Jousseaume, 1894).
Gulf of Thailand. Reported by: Gemert (2003).
Epitonium melior (Melvill & Standen, 1903). Reported as: Epitonium (Hyaloscala) melior (Melvill & Standen,
1903). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium cf. millecostatum (Pease, 1861). Andaman Sea. Reported by: BEDO (2017e).
Epitonium multicostatum (Sowerby, 1844). Reported as: Epitonium (Sodaliscala) multicostata (Sowerby, 1844).
Gulf of Thailand. Reported by: Gemert (2003).
Epitonium cf. obliquum (Sowerby, 1844). Reported as: Epitonium (Laeviscala) cf. obliquum (Sowerby, 1847). Gulf
of Thailand. Reported by: Gemert (2003).
Epitonium octagonum (G. B. Sowerby, 1905). Reported as: Epitonium (Epitonium) octagonum (G. B. Sowerby,
1905). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium pallasi (Kiener, 1838). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Thamrongnawasawat
et al. (2008), Nabhitabhata (2009).
Epitonium paumotense (Pease, 1868). Reported as: Epitonium (Sodaliscala) paumotense (Pease, 1868). Gulf of
Thailand. Reported by: Gemert (2003), Brown & Neville (2015).
Epitonium philippinarum (G. B. Sowerby II, 1844). Reported as: Epitonium cf. (Lamelliscala) philippinarum (G. B.
Sowerby II, 1844). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium replicatum (G. B. Sowerby II, 1844). Gulf of Thailand, Andaman Sea. Reported by: Robba et al. (2004).
Epitonium robillardi (G. B. Sowerby III, 1894). Reported as: Epitonium (Papyriscala) robillardi (Sowerby, 1894).
Gulf of Thailand. Reported by: Gemert (2003), Robba et al. (2004).
Epitonium sandwichense (Nyst, 1871). Reported as: Epitonium (Laeviscala) sandwichense (Nyst, 1871). Gulf of
Thailand. Reported by: Gemert (2003).
Epitonium scalare (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Swennen et al. (2001), Nabhitabhata (2009), BEDO (2017b).
Epitonium syoichiroi Masahito & Habe, 1976. Andaman Sea. Reported by: BEDO (2017b).
Epitonium symmetricum (Pease, 1868). Reported as: Epitonium (Sodaliscala) symmetricum (Pease, 1868). Gulf
of Thailand. Reported by: Gemert (2003).
Epitonium tenuicostatum (G. B. Sowerby, 1844). Reported as: Epitonium (Hirtoscala) tenuicostatum (G. B.
Sowerby, 1844). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium tenuiliratum (G. B. Sowerby II, 1874). Gulf of Thailand. Reported by: Gemert (2003).
Epitonium terebriforme (Thiele, 1925). Reported as: Epitonium (Hirtoscala) terebriformis (Thiele, 1925). Gulf of
Thailand. Reported by: Gemert (2003).
Epitonium townsendi (Melvill & Standen, 1903). Reported as: Epitonium (Parviscala) townsendi (Melvill &
Standen, 1903). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Gregorioiscala unilateralis (Martens, 1902). Gulf of Thailand. Reported by: Gemert (2003).
Janthina globosa Swainson, 1822. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Gemert (2003), Duangdee (2004), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019b).
Opalia hidryma (Melvill, 1899). Reported as: Opalia (Pliciscala) hidryma (Melvill, 1899). Gulf of Thailand.
Reported by: Robba et al. (2007).
Surrepifungium costulatum (Kiener, 1839). Also reported as: Epitonium costulatum (Kiener, 1839). Andaman
Sea. Reported by: DuShane (1988a), Gittenberger & Gittenberger (2005), Brown & Neville (2015), Galli (2015).
Surrepifungium oliverioi (Bonfitto & Sabelli, 2001). Reported as: Epitonium oliverioi Bonfitto & Sabelli, 2001.
Andaman Sea. Reported by: Gittenberger & Gittenberger (2005), Galli (2015).
Surrepifungium patamakanthini A. Gittenberger & E. Gittenberger, 2005. Reported as: Epitonium
(Surrepifungium) patamakanthini (A. Gittenberger & E. Gittenberger, 2005). Andaman Sea. Reported by: A.
Gittenberger & E. Gittenberger (2005), Thamrongnawasawat et al. (2008), Brown & Neville (2015), Galli
(2015), GBIF (2019a).
Superfamily Triphoroidea
Family Cerithiopsidae
Cerithiopsis barleei Jeffreys, 1867. Gulf of Thailand. Reported by: Sawatna et al. (2012).
Cerithiopsis subreticulata (Dunker, 1860). Reported as: Clathropsis subreticulata (Dunker, 1860). Gulf of
Thailand. Reported by: Gemert (2003).
Cerithiopsis tubercularis (Montagu, 1803). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Costulopsis albocincta (Melvill & Standen, 1896). Andaman Sea. Reported as: Joculator albocinctum (Melvill &
Standen, 1896). Reported by: BEDO (2017b).
Horologica bipartita Laseron, 1956. Reported as: ? Horologica bipartita Laseron, 1956. Gulf of Thailand.
Reported by: Gemert (2003).
Horologica cf. fraudulenta Cecalupo & Perugia, 2013. Andaman Sea. Reported by: BEDO (2017e).
Horologica semipicta (Gould, 1861). Andaman Sea. Reported by: BEDO (2017e).
Joculator cf. baraggiai Cecalupo & Perugia, 2013. Andaman Sea. Reported by: BEDO (2017e).
Joculator granulosus Cecalupo & Perugia, 2013. Andaman Sea. Reported by: BEDO (2017e).
Joculator marileutes (Melvill & Standen, 1896). Gulf of Thailand. Reported by: Gemert (2003), Robba et al.
(2004), Sawatna et al. (2012). Recorded by WoRMS Editorial Board (2019) as a taxon inquirendum.
Joculator ovatus Laseron, 1956. Andaman Sea. Reported by: BEDO (2017e).
Paraseila ampulla Laseron, 1956. Andaman Sea. Reported by: BEDO (2017e).
Seila japonica Habe, 1970. Reported as: Paraseila japonica Habe, 1970. Andaman Sea. Reported by: BEDO (2017e).
Seila morishimai (Habe, 1970). Andaman Sea. Reported by: BEDO (2017e).
Family Triphoridae
Aclophora xystica (Jousseaume, 1884). No Thai locality provided. Reported by: BEDO (2017e).
Bouchetriphora pallida (Pease, 1870). Gulf of Thailand. Reported by: Robba et al. (2004).
Cautotriphora hervieri (Kosuge, 1962). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Cheirodonta pallescens (Jeffreys, 1867). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Coriophora cnodax (Jousseaume, 1884). Also reported as: Mesophora cnodax (Jousseaume, 1884). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Dumrongrojwattana et al. (2016), BEDO (2017e).
Coriophora fusca (Dunker, 1860). Reported as: Mesophora hungerfordi (G. B. Sowerby III, 1914). Gulf of
Thailand, Andaman Sea. Reported by: Gemert (2003), Dumrongrojwattana et al. (2016), BEDO (2017e).
Coriophora monilifera (Hinds, 1843). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Costatophora iniqua (Jousseaume, 1898). Reported as: Tetraphora iniqua (Jousseaume, 1898). Gulf of Thailand.
Reported by: Gemert (2003), BEDO (2017e).
Costatophora pallida (Kosuge, 1962). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Euthymella bilix (Hinds, 1843). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Euthymella concors (Hinds, 1843). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Euthymella elegans (Hinds, 1843). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Inella asperrima (Hinds, 1843). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Iniforis formosula (Hervier, 1898). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Iniforis fusiformis (Kosuge, 1961). Gulf of Thailand, Andaman Sea. Reported by: Dumrongrojwattana et al. (2016),
BEDO (2017e).
Iniforis ikukoae (Kosuge, 1963). Also reported as: Mastoniaeforis ikukoae (Kosuge, 1963). Andaman Sea.
Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
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Iniforis progressa (Laseron, 1958). Gulf of Thailand. Reported by: Gemert (2003).
Marshallora adversa (Montagu, 1803). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Mastonia cingulifera (Pease, 1861). Andaman Sea. Reported by: BEDO (2017e).
Mastonia cf. clavata Pease, 1861. Gulf of Thailand. Reported by: BEDO (2017e).
Mastonia lamberti (Hervier, 1897). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Mastonia cf. millepunctata (Kosuge, 1962). Gulf of Thailand. Reported by: Gemert (2003).
Mastonia peanites (Jousseaume, 1898). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO
(2017e).
Mastonia rubra (Hinds, 1843). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Sawatna et al. (2012),
Kostina et al. (2016), BEDO (2017e).
Mastonia vulpina Hinds, 1843. Also reported as: Viriola vulpina (Hinds, 1843). Andaman Sea. Reported by:
Dumrongrojwattana et al. (2016), BEDO (2017e).
Mastoniaeforis jousseaumei (Hervier, 1898). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016),
BEDO (2017e).
Mastoniaeforis lifunana (Hervier, 1898). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO
(2017e).
Monophorus atratus (Kosuge, 1962). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Monophorus lucidulus (Hervier, 1898). Reported as: Obesula lucidula (Hervier, 1897). Gulf of Thailand. Reported
by: Robba et al. (2007).
Monophorus perversus (Linnaeus, 1758). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Monophorus tessellatus (Kosuge, 1963). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Nanaphora pygmaeus (Kosuge, 1963). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Nanaphora tricolor (Laseron, 1958). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Nanaphora triticea (Pease, 1861). Gulf of Thailand, Andaman Sea. Reported by: Dumrongrojwattana et al.
(2016), BEDO (2017e).
Obesula lucidula (Hervier, 1897). Gulf of Thailand. Reported by: Robba et al. (2007). This species canno t be
confirmed through WoRMS Editorial Board (2019), but is listed by Robba et al. (2007).
Obesula turricula (Hervier, 1898). Reported as: Obesula turricata (Hervier, 1898). Gulf of Thailand, Andaman
Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Opimaphora coralina Laseron, 1958. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Opimaphora sarcira Laseron, 1958. Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Similiphora similior (Bouchet & Guillemot, 1978). Gulf of Thailand. Reported by: Sawatna et al. (2012).
Similiphora triclotae Bouchet, 1997. Gulf of Thailand. Reported by: Sawatna et al. (2012).
Subulophora rutilans (Hervier, 1898). Also reported as: ? Monophorus rutilans (Hervier, 1898). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Triphora taeniolata Hervier, 1897. Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Viriola cancellata (Hinds, 1843). Gulf of Thailand, Andaman Sea. Reported by: Dumrongrojwattana et al. (2016),
BEDO (2017e).
Viriola corrugata (Hinds, 1843). Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007),
Sawatna et al. (2012).
Viriola incisa (Pease, 1861). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016).
Viriola intergranosa (Hervier, 1898). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO
(2017e).
Viriola cf. pagodus (Hinds, 1843). Reported as: Euthymella cf. pagoda (Hinds, 1843). Andaman Sea. Reported by:
BEDO (2017e).
Viriola tricincta (Dunker, 1882). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e).
Viriolopsis fallax (Kay, 1979). Andaman Sea. Reported by: Dumrongrojwattana et al. (2016), BEDO (2017e).
Subclass Heterobranchia
Infraclass “Lower Heterobranchia”
Superfamily Architectonicoidea
Family Architectonicidae
Adelphotectonica reevei (Hanley, 1862). Gulf of Thailand. Reported by: Sanpanich & Duangdee (2013).
Architectonica laevigata (Lamarck, 1816). Gulf of Thailand. Reported by: Bieler (1993).
Architectonica maculata (Link, 1807). Also reported as: Architectonica picta (Philippi, 1849). Andaman Sea.
Reported by: Bieler (1993), Swennen et al. (2001), GBIF (2019a).
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Architectonica maxima (Philippi, 1849). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Kasetsart
University (2000), Duangdee et al. (2007), Thamrongnawasawat et al. (2008).
Architectonica modesta (Philippi, 1849). Andaman Sea. Reported by: Bieler (1993).
Architectonica perdix (Hinds, 1844). Also reported as: Solarium perdix Hinds, 1844. Gulf of Thailand, Andaman
Sea. Reported by: Morlet (1889), Bieler (1993), Swennen et al. (2001), Robba et al. (2004), BEDO (2017b), GBIF
(2019b).
Architectonica perspectiva (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Suvatti
(1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Bieler (1993), Bussarawit (1995), Kasetsart
University (2000, 2006), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011), ONEP (2007), Thamrongnawasawat et al. (2008), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a, 2019b).
Architectonica purpurata (Hinds, 1844). Also reported as: Solarium purpurata Hinds, 1844. Gulf of Thailand.
Reported by: Morlet (1889), Bieler (1993).
Architectonica stellata (Philippi, 1849). Andaman Sea. Reported by: BEDO (2017b).
Architectonica taylori (Hanley, 1862). Andaman Sea. Reported by: GBIF (2019a).
Architectonica trochlearis (Hinds, 1844). Gulf of Thailand. Reported by: GBIF (2019b).
Heliacus areola (Gmelin, 1791). Andaman Sea. Reported by: Kilburn & Hylleberg (1998).
Heliacus costatus (Schepman, 1909). Reported as: Heliacus (Torinista) costatus (Schepman, 1909). Gulf of
Thailand. Reported by: Robba et al. (2007).
Heliacus fenestratus (Hinds, 1844). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017e).
Heliacus implexus (Mighels, 1845). Reported as: Heliacus (Torinista) implexus (Mighels, 1845). Andaman Sea.
Reported by: Bieler (1993), BEDO (2017e).
Heliacus infundibulifeormis (Gmelin, 1791). Andaman Sea. Reported by: Kilburn & Hylleberg (1998).
Heliacus sterkii (Pilsbry & Vanatta, 1908). Andaman Sea. Reported by: BEDO (2017b).
Heliacus stramineus (Gmelin, 1791). Reported as: Heliacus (Grandeliacus) stramineus (Gmelin, 1791). Andaman
Sea. Reported by: Bieler (1993).
Heliacus variegatus (Gmelin, 1791). Reported as: Heliacus (Heliacus) variegatus (Gmelin, 1791). Andaman Sea.
Reported by: Bieler (1993), Kilburn & Hylleberg (1998), BEDO (2017e).
Pseudotorinia delectabilis (Melvill, 1893). Gulf of Thailand. Reported by: Gemert (2003).
Pseudotorinia gemmulata (Thiele, 1925). Gulf of Thailand, Andaman Sea. Reported by: Bieler (1993), Robba
et al. (2004).
Solarium stramineum (Gmelin, 1791). Andaman Sea. Reported by: GBIF (2019a).
Superfamily Cimoidea
Family Cimidae
Graphis cf. tenuissima (Hedley, 1909). Gulf of Thailand. Reported by: Gemert (2003).
Superfamily Mathildoidea
Family Mathildidae
Mathilda carystia Melvill & Standen, 1903. Gulf of Thailand. Reported by: Robba et al. (2007).
Superfamily Murchisonelloidea
Family Murchisonellidae
Murchisonella densistriata (Nomura, 1936). Gulf of Thailand. Reported by: Gemert (2003).
Murchisonella hatienensis (Saurin, 1962). Reported as: Pandorella hatienensis Saurin, 1962. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Superfamily Orbitestelloidea
Family Orbitestellidae
Boschitestella donaldi Moolenbeek, 1994. Andaman Sea. Reported by: Moolenbeek (2007).
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Infraclass Euthyneura
Subterclass Acteonimorpha
Superfamily Acteonoidea
Family Acteonidae
Punctacteon variegatus (Bruguière, 1789). Also reported as: Acteon variegatus (Bruguière, 1789). Andaman
Sea. Reported by: Nateewathana et al. (1981), Jensen (1998, 2007), BEDO (2017b).
Punctacteon yamamurae Habe, 1976. Gulf of Thailand. Reported by: Robba et al. (2004).
Pupa nitidula (Lamarck, 1816). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich
& Duangdee (2013).
Pupa solidula (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Jensen
(1998, 2007), Robba et al. (2004), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a).
Pupa strigosa (Gould, 1859). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Jensen (1998), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Pupa sulcata (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Jensen
(1998), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO
(2017b).
Pupa tessellata (Reeve, 1842). Gulf of Thailand. Reported by: Gemert (2003).
Family Aplustridae
Bullina lineata (Gray, 1825). Reported as: Bulla lineate (Bruguière, 1792). Gulf of Thailand, Andaman Sea.
Reported by: Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Hydatina albocincta (Van der Hoeven, 1893). Gulf of Thailand, Andaman Sea. Reported by: THNHM,
Nateewathana et al. (1981), Jensen (1998, 2007), Nabhitabhata (2009).
Hydatina zonata Lightfoot, 1786. Also reported as: Hydatina velum Gmelin, 1791. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Nateewathana et al. (1981), Jensen (1998, 2007), Nabhitabhata (2009), Taylor
(2009), BEDO (2017b), GBIF (2019b).
Micromelo undatus (Bruguière, 1792). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017c).
Subterclass Ringipleura
Superorder Nudipleura
Order Nudibranchia
Suborder Cladobranchia
Superfamily Aeolidoidea
Family Aeolidiidae
Baeolidia harrietae (Rudman, 1982). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Baeolidia japonica Baba, 1933. Gulf of Thailand. Reported by: Chavanich et al. (2010b, 2013).
Cerberilla ambonensis Bergh, 1905. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Cerberilla asamusiensis Baba, 1940. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Family Facelinidae
Antonietta janthina Baba & Hamatani, 1977. Andaman Sea. Reported by: BEDO (2017c).
Caloria indica (Bergh, 1896). Also reported as: Phidiana indica (Bergh, 1896). Gulf of Thailand, Andaman Sea.
Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013),
Duangdee et al. (2013b), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018).
Hermissenda crassicornis (Eschscholtz, 1831). Gulf of Thailand. Reported by: GBIF (2019b).
Moridilla brocki Bergh, 1888. Gulf of Thailand. Reported by: Nabhitabhata (2009).
Phidiana lynceus Bergh, 1867. Gulf of Thailand. Reported by: GBIF (2019b).
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Phidiana militaris (Alder & Hancock, 1864). Also reported as: Caloria militaris (Alder & Hancock, 1864). Gulf of
Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Rudman (2010), Chavanich et al. (2013), BEDO (2017c), Mehrotra & Scott (2017).
Pteraeolidia ianthina (Angas, 1864). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Gosliner et al.
(2008), Thamrongnawasawat et al. (2008), Sanpanich & Duangdee (2009, 2018), Chavanich et al. (2010b, 2013),
Duangdee et al. (2011, 2013a, 2013b), BEDO (2017c).
Pteraeolidia semperi (Bergh, 1870). Andaman Sea. Reported by: GBIF (2019a).
Family Myrrhinidae
Phyllodesmium magnum Rudman, 1991. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010),
Duangdee et al. (2011, 2013b), BEDO (2017c), Mehrotra & Scott (2017).
Superfamily Arminoidea
Family Arminidae
Armina comta (Bergh, 1880). Gulf of Thailand. Reported by: Rudman (2010).
Armina cf. japonica (Eliot, 1913). No Thai locality provided. Reported by: Chavanich (2010b).
Armina occulta Mehrotra, Caballer & Chavanich, 2017. Gulf of Thailand. Reported by: Mehrotra et al. (2017),
Yonow (2017), GBIF (2019b).
Armina scotti Mehrotra, Caballer & Chavanich, 2017. Also reported as: Armina semperi (Bergh, 1866). Gulf of
Thailand. Reported by: Thamrongnawasawat et al. (2008), Rudman (2010), Chavanich et al. (2013), Mehrotra
& Scott (2016), Mehrotra et al. (2017).
Armina semperi (Bergh, 1866). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b, 2013),
BEDO (2017c).
Dermatobranchus albus (Eliot, 1904). Gulf of Thailand. Reported by: BEDO (2017c).
Dermatobranchus caeruleomaculatus Gosliner & Fahey, 2011. Gulf of Thailand, Andaman Sea. Reported by:
BEDO (2017c).
Dermatobranchus dendronephthyphagus Gosliner & Fahey, 2011. Gulf of Thailand, Andaman Sea. Reported
by: BEDO (2017c).
Dermatobranchus fortunatus (Bergh, 1888). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra et al.
(2017).
Dermatobranchus gonatophorus van Hasselt, 1824. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al.
(2008), Nabhitabhata (2009), Rudman (2010), Gosliner & Fahey (2011), Chavanich et al. (2013), BEDO (2017c),
GBIF (2019a).
Dermatobranchus oculus Gosliner & Fahey, 2011. Andaman Sea. Reported by: BEDO (2017c).
Dermatobranchus ornatus (Bergh, 1874). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010),
Gosliner & Fahey (2011), BEDO (2017c), GBIF (2019a).
Dermatobranchus striatus van Hasselt, 1824. Gulf of Thailand. Reported by: Rudman (2010), BEDO (2017c),
Mehrotra et al. (2017).
Family Doridomorphidae
Doridomorpha gardineri Eliot, 1903. Reported as: Doridomorpha gardineri Eliot, 1906. Andaman Sea. Reported
by: Willan & Poppe (2010).
Superfamily Dendronotoidea
Family Bornellidae
Bornella anguilla S. Johnson, 1984. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013), BEDO (2017c).
Bornella excepta Bergh, 1884. Gulf of Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007),
Nabhitabhata (2009), Chavanich et al. (2013).
Bornella hermanni Angas, 1864. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Bornella stellifer (A. Adams & Reeve [in A. Adams], 1848). Also reported as: Bornella digitata (A. Adams &
Reeve [in A. Adams], 1848). Gulf of Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007),
Duangdee et al. (2009), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Chavanich et al. (2010b, 2013),
Hylleberg & Aungtonya (2013), BEDO (2017c).
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Family Lomanotidae
Lomanotus vermiformis Eliot, 1908. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c), Mehrotra et al. (2017).
Family Scyllaeidae
Scyllaea pelagica (Linnaeus, 1758). Andaman Sea. Reported by: Wells (2005).
Notobryon wardi Odhner, 1936. Andaman Sea. Reported by: GBIF (2019a).
Family Tethydidae
Melibe bucephala Bergh, 1902. Gulf of Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007),
Gosliner et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013).
Melibe engeli Risbec, 1937. Gulf of Thailand. Reported by: BEDO (2017c).
Melibe viridis (Kelaart, 1858). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Superfamily Fionoidea
Family Flabellinidae
Coryphellina exoptata (Gosliner & Willan, 1991). Reported as: Flabellina exoptata Gosliner & Willan, 1991. Gulf
of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich
et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017).
Coryphellina indica (Bergh, 1902). Reported as: Nossis indica Bergh, 1902. Gulf of Thailand. Reported by: Bergh
(1902), Jensen (1998, 2007).
Coryphellina rubrolineata O’Donoghue, 1929. Reported as: Flabellina rubrolineata (O’Donoghue, 1929). Gulf of
Thailand, Andaman Sea. Reported by: Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), Duangdee et al. (2011, 2013a, 2013b), BEDO (2017c), Mehrotra & Scott (2017),
GBIF (2019a, 2019b).
Family Samlidae
Luisella babai (Schmekel, 1972). Reported as: Flabellina babai Schmekel, 1972. Andaman Sea. Reported by:
BEDO (2017c).
Samla bicolor (Kelaart, 1858). Also reported as: Flabellina bicolor (Kelaart, 1858). Gulf of Thailand, Andaman
Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013),
BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018).
Family Trinchesiidae
Phestilla lugubris (Bergh, 1870). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b, 2013),
BEDO (2017c), Mehrotra & Scott (2017).
Phestilla melanobrachia Bergh, 1874. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017),
Sanpanich & Duangdee (2018).
Phestilla minor Rudman, 1981. Gulf of Thailand. Reported by: Chavanich et al. (2010b, 2013), BEDO (2017c),
Sanpanich & Duangdee (2018).
Trinchesia ornata Baba, 1937. Cuthona ornata Baba, 1937. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra
& Scott (2017).
Trinchesia sibogae (Bergh, 1905). Reported as: Cuthona sibogae Bergh, 1905. Gulf of Thailand, Andaman Sea.
Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman
(2010), BEDO (2017c).
Trinchesia yamasui (Hamatani, 1993). Reported as: Cuthona yamasui (Hamatani, 1993). Andaman Sea. Reported
by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), BEDO (2017c), Mehrotra & Scott
(2017).
Family Unidentiidae
Unidentia angelvaldesi Millen & Hermosillo, 2012. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
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Superfamily Proctonotoidea
Family Janolidae
Janolus savinkini Martynov & Korshunova, 2012. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
Superfamily Tritonoidea
Family Tritoniidae
Marionia chloanthes Bergh, 1902. Gulf of Thailand, Andaman Sea. Reported by: Jensen (1998, 2007),
Nabhitabhata (2009), Chavanich et al. (2013).
Tritoniopsis elegans (Audouin, 1826). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Duangdee et al. (2009, 2013b), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c).
Suborder Doridina
Infraorder Doridoidei
Superfamily Chromodoroidea
Family Actinocyclidae
Hallaxa fuscescens (Pease, 1871). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b, 2013),
BEDO (2017c).
Hallaxa iju Gosliner & S. Johnson, 1994. Andaman Sea. Reported by: BEDO (2017c).
Hallaxa indecora (Bergh, 1905). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017c).
Family Cadlinidae
Aldisa erwinkoehleri Perrone, 2001. Gulf of Thailand, Andaman Sea. Reported by: Perrone (2001), Gosliner et al.
(2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013),
BEDO (2017c), GBIF (2019a).
Family Chromodorididae
Ardeadoris angustolutea (Rudman, 1990). Reported as: Noumea angustolutea Rudman, 1990. Gulf of Thailand.
Reported by: Gosliner et al. (2008), Nabhitabhata (2009), BEDO (2017c).
Ardeadoris averni Rudman, 1985. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Ardeadoris electra (Rudman, 1990). Gulf of Thailand. Reported by: BEDO (2017c).
Ardeadoris poliahu (Bertsch & Gosliner, 1989). Reported as: Glossodoris poliahu Bertsch & Gosliner, 1989.
Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017c).
Cadlinella ornatissima (Risbec, 1928). Reported as: Cadlina ornamatissima (Risbec, 1928). Gulf of Thailand,
Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2018), Chavanich et al. (2010b, 2013), Rudman (2010), Duangdee et al. (2013b),
BEDO (2017c), Mehrotra & Scott (2017).
Ceratosoma gracillimum Semper in Bergh, 1876. Andaman Sea. Reported by: BEDO (2017c).
Ceratosoma tenue Abraham, 1876. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017c).
Ceratosoma trilobatum (J. E. Gray, 1827). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c),
Sanpanich & Duangdee (2018).
Chromodoris annae Bergh, 1877. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c), Sanpanich & Duangdee (2018).
Chromodoris aspersa (Gould, 1852). Andaman Sea. Reported by: BEDO (2017c).
Chromodoris elisabethina Bergh, 1877. Gulf of Thailand, Andaman Sea. Reported by: Wells (2005),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013).
Chromodoris joshi Gosliner & Behrens, 1998. No Thai locality provided. Reported by: Nabhitabhata (2009),
BEDO (2017c).
Chromodoris lineolata (van Hasselt, 1824). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Chavanich et al. (2010b, 2013), BEDO (2017c).
Chromodoris lochi Rudman, 1982. Gulf of Thailand. Reported by: GBIF (2019b).
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The Marine and Estuarine Molluscs of Thailand
Chromodoris mandapamensis Valdés, Mollo & Ortea, 1999. Gulf of Thailand. Reported by: Chavanich et al.
(2013), BEDO (2017c).
Chromodoris strigata Rudman, 1982. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), BEDO (2017c).
Diversidoris crocea (Rudman, 1986). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Diversidoris flava (Eliot, 1904). Reported as Noumea flava Eliot, 1904. Gulf of Thailand, Andaman Sea. Reported
by: Nabhitabhata (2009), Rudman (2010).
Doriprismatica atromarginata (Cuvier, 1804). Reported as: Glossodoris atromarginata (Cuvier, 1804); Casella
atromarginata (Cuvier, 1804); and Doriprismatica atromarginata pallida Bergh, 1905. Gulf of Thailand, Andaman
Sea. Reported by: Bergh (1902), Jensen (1998, 2007), Thamrongnawasawat et al. (2008), Duangdee et al.
(2009, 2011), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra
& Scott (2017), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Felimida rodomaculata (Ortea & Valdés, 1992). Reported as: Chromodoris rufomaculata Ortea & Valdés, 1992.
Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Sanpanich & Duangdee (2018).
Glossodoris cincta (Bergh, 1888). Also reported as: Chromodoris rufomaculata Pease, 1871. Gulf of Thailand,
Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich &
Duangdee (2018), GBIF (2019a).
Glossodoris hikuerensis (Pruvot-Fol, 1954). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c), Sanpanich & Duangdee
(2018), GBIF (2019a).
Glossodoris pallida (Rüppell & Leuckart, 1828). Gulf of Thailand, Andaman Sea. Reported by:
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c), Sanpanich &
Duangdee (2018).
Glossodoris rufomarginata (Bergh, 1890). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c).
Goniobranchus albonares (Rudman, 1990). Reported as: Chromodoris albonares Rudman, 1990. Gulf of Thailand,
Andaman Sea. Reported by: Chavanich et al. (2013), BEDO (2017c).
Goniobranchus albopunctatus Garrett, 1879. Also reported as: Chromodoris albopunctata (Garrett, 1879). Gulf
of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009), Nabhitabhata (2009), Rudman (2010).
Goniobranchus annulatus (Eliot, 1904). Reported as: Chromodoris annulata Eliot, 1904. Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee
et al. (2009), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Sanpanich
& Duangdee (2018), GBIF (2019a).
Goniobranchus aureomarginata (Cheeseman, 1881). Also reported as: Chromodoris aureomarginata Cheeseman,
1881. No Thai locality provided. Reported by: Chavanich et al. (2010b).
Goniobranchus aureopurpureus (Collingwood, 1881). Reported as: Chromodoris aureopurpurea Collingwood,
1881. Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra &
Scott (2017), Sanpanich & Duangdee (2018).
Goniobranchus bombayanus (Winckworth, 1946). Reported as: Chromodoris naiki Valdés, Mollo & Ortea, 1999.
Andaman Sea. Reported by: Gosliner et al. (2008).
Goniobranchus cavae (Eliot, 1904). Reported as: Chromodoris cavae Eliot, 1904. Andaman Sea. Reported by:
Yonow (2012), BEDO (2017c).
Goniobranchus coi (Risbec, 1956). Reported as: Chromodoris coi (Risbec, 1956). Gulf of Thailand, Andaman Sea.
Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c),
Sanpanich & Duangdee (2018).
Goniobranchus conchyliatus (Yonow, 1984). Reported as: Chromodoris conchyliata Yonow, 1984. Gulf of
Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Yonow (2012).
Goniobranchus decorus (Pease, 1860). Reported as: Chromodoris decora (Pease, 1860). Andaman Sea. Reported
by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013).
Goniobranchus fidelis (Kelaart, 1858). Reported as: Chromodoris fidelis (Kelaart, 1858). Gulf of Thailand, Andaman
Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013),
Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF (2019a).
Goniobranchus geminus (Rudman, 1987). Reported as: Chromodoris geminus Rudman, 1987. Gulf of Thailand,
Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009,
2013), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013), BEDO (2017c), Sanpanich & Duangdee
(2018).
Goniobranchus geometricus (Risbec, 1928). Also reported as: Chromodoris geometrica Risbec, 1928. Gulf of
Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al.
(2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF
(2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Goniobranchus gleniei (Kelaart, 1858). Reported as: Chromodoris gleniei (Kelaart, 1858). Gulf of Thailand,
Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich
et al. (2013), BEDO (2017c), Sanpanich & Duangdee (2018).
Goniobranchus hintuanensis (Gosliner & Behrens, 1998). Reported as: Chromodoris hintuanensis Gosliner &
Behrens, 1998. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c).
Goniobranchus kuniei (Pruvot-Fol, 1930). Reported as: Chromodoris kuniei Pruvot-Fol, 1930. Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c).
Goniobranchus leopardus (Rudman, 1987). Also reported as: Chromodoris leopardus Rudman, 1987. Andaman
Sea. Reported by: Chavanich et al. (2010b), BEDO (2017c).
Goniobranchus mandapamensis (Valdés, Mollo & Ortea, 1999). Reported as: Chromodoris mandapamensis
Valdés, Mollo & Ortea, 1999. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Nabhitabhata
(2009), Chavanich et al. (2013), Mehrotra & Scott (2017).
Goniobranchus naiki (Valdés, Mollo & Ortea, 1999). Andaman Sea. Reported by: BEDO (2017c).
Goniobranchus preciosus (Kelaart, 1858). Reported as: Chromodoris preciosa (Kelaart, 1858). Gulf of Thailand,
Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013a), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017).
Goniobranchus reticulatus (Quoy & Gaimard, 1832). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
Goniobranchus roboi (Gosliner & Behrens, 1998). Reported as: Chromodoris roboi Gosliner & Behrens, 1998.
Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich
et al. (2013), BEDO (2017c).
Goniobranchus sinensis (Rudman, 1985). Reported as: Chromodoris sinensis Rudman, 1985. Gulf of Thailand,
Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Chavanich et al. (2010b, 2013), BEDO (2017c).
Goniobranchus tinctorius (Rüppell & Leuckart, 1830). Reported as: Chromodoris tinctoria (Rüppell & Leuckart,
1828). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), BEDO (2017c).
Goniobranchus trimarginatus (Winckworth, 1946). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
Goniobranchus tumuliferus (Collingwood, 1881). Reported as: Chromodoris tumulifera Collingwood, 1881. Gulf
of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c),
Mehrotra & Scott (2017).
Goniobranchus verrieri (Crosse, 1875). No Thai locality provided. Reported by: BEDO (2017c).
Hypselodoris apolegma (Yonow, 2001). Andaman Sea. Reported by: BEDO (2017c).
Hypselodoris bollandi Gosliner & R. F. Johnson, 1999. Gulf of Thailand, Andaman Sea. Reported by:
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c).
Hypselodoris bullockii (Collingwood, 1881). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra
& Scott (2017), GBIF (2019a).
Hypselodoris carnea Bergh, 1889. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009).
Hypselodoris emma Rudman, 1977. Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Chavanich
et al. (2013), BEDO (2017c).
Hypselodoris ghardaqana (Gohar & Aboul-Ela, 1957). Reported as: Risbecia ghardaqana Gohar & Aboul-Ela,
1957. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
BEDO (2017c), Gosliner et al. (2018).
Hypselodoris iacula Gosliner & R. F. Johnson, 1999. Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c),
Sanpanich & Duangdee (2018), GBIF (2019a).
Hypselodoris infucata (Rüppell & Leuckart, 1830). Gulf of Thailand, Andaman Sea. Reported by:
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c),
Mehrotra & Scott (2017), Sanpanich & Duangdee (2018).
Hypselodoris kanga Rudman, 1977. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Chavanich et al.
(2010b, 2013), Rudman (2010), BEDO (2017c), Sanpanich & Duangdee (2018).
Hypselodoris krakatoa Gosliner & Johnson, 1999. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al.
(2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2018), Rudman (2010), Chavanich et al. (2013), BEDO (2017c).
Hypselodoris maculosa (Pease, 1871). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013b), Nabhitabhata (2009), Chavanich et al.
(2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF
(2019b).
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The Marine and Estuarine Molluscs of Thailand
Hypselodoris maridadilus Rudman, 1977. Andaman Sea. Reported by: Sanpanich & Duangdee (2018).
Hypselodoris maritima (Baba, 1949). Gulf of Thailand. Reported by: Gosliner et al. (2008), BEDO (2017c),
Mehrotra & Scott (2017).
Hypselodoris nigrostriata (Eliot, 1904). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Rudman (2010), Chavanich et al. (2013), BEDO (2017c), Sanpanich & Duangdee (2018).
Hypselodoris obscura Stimpson, 1855. Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b,
2013), BEDO (2017c).
Hypselodoris pulchella (Rüppell & Leuckart, 1830). Also reported as: Risbecia pulchella (Rüppell & Leuckart,
1828). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008),
Duangdee et al. (2009, 2011), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO
(2017c), Sanpanich & Duangdee (2018), GBIF (2019a).
Hypselodoris saintvincentia Burn, 1962. Andaman Sea. Reported by: BEDO (2017c).
Hypselodoris tryoni (Garrett, 1873). Reported as: Risbecia tryoni (Garrett, 1873). Gulf of Thailand, Andaman Sea.
Reported by: Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018).
Hypselodoris whitei (A. Adams & Reeve, 1850). Andaman Sea. Reported by: BEDO (2017c).
Hypselodoris zebrina Alder & Hancock, 1864. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata
(2009), Rudman (2010), BEDO (2017c).
Hypselodoris zephyra Gosliner & R. F. Johnson, 1999. No Thai locality reported. Reported by: Gosliner et al. (2008).
Mexichromis lemniscata (Quoy & Gaimard, 1832). Reported as: Durvilledoris lemniscata (Quoy & Gaimard, 1832).
Andaman Sea. Reported by: Jensen (1998, 2007), Gosliner et al. (2008), Nabhitabhata (2009), Chavanich et al.
(2013), BEDO (2017c).
Mexichromis multituberculata (Baba, 1953). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra
& Scott (2017), Sanpanich & Duangdee (2018).
Mexichromis pusilla (Bergh, 1874). Reported as: Durvilledoris pusilla (Bergh, 1874). Andaman Sea. Reported by:
Rudman (2010), BEDO (2017c).
Mexichromis similaris (Rudman, 1986). Reported as: Durvilledoris similaris Rudman, 1986. Gulf of Thailand,
Andaman Sea. Reported by: Gosliner et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c),
Sanpanich & Duangdee (2018).
Mexichromis trilineata (A. Adams & Reeve, 1850). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
Miamira sinuata (van Hasselt, 1824). Reported as: Ceratosoma sinuatum (van Hasselt, 1824). Gulf of Thailand,
Andaman Sea. Reported by: Jensen (2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c).
Thorunna africana Rudman, 1984. Andaman Sea. Reported by: BEDO (2017c).
Thorunna australis (Risbec, 1928). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Chavanich et al. (2013).
Thorunna florens Baba, 1949. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009).
Thorunna horologia Rudman, 1984. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017c).
Verconia norba (Er. Marcus & Ev. Marcus, 1970). Andaman Sea. Reported by: BEDO (2017c).
Verconia romeri (Risbec, 1928). Andaman Sea. Reported by: BEDO (2017c).
Family Hexabranchidae
Hexabranchus sanguineus (Rüppell & Leuckart, 1830). Gulf of Thailand, Andaman Sea. Reported by: Chavanich
et al. (2010b, 2013), BEDO (2017c), GBIF (2019b).
Superfamily Doridoidea
Family Discodorididae
Asteronotus cespitosus (van Hasselt, 1824). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al.
(2010b, 2013), BEDO (2017c), GBIF (2019a).
Carminodoris grandiflora (Pease, 1860). Reported as: Hoplodoris grandiflora (Pease, 1860). Gulf of Thailand.
Reported by: Rudman (2010).
Carminodoris nodulosa (Angas, 1864). Reported as: Hoplodoris nodulosa (Angas, 1864). Gulf of Thailand,
Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b,
2013), BEDO (2017c).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Discodoris boholiensis Bergh, 1877. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009), Chavanich et al. (2010b, 2013), Rudman (2010),
Duangdee et al. (2013b), BEDO (2017c), Mehrotra & Scott (2017).
Halgerda bacalusia Fahey & Gosliner, 1999. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017),
GBIF (2019a).
Halgerda formosa Bergh, 1880. Andaman Sea. Reported by: Gosliner et al. (2008).
Halgerda punctata Farran, 1905. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Nabhitabhata
(2009), Rudman (2010), Yonow (2012), BEDO (2017c), GBIF (2019a).
Halgerda stricklandi Fahey & Gosliner, 1999. Also reported as: Hypselodoris stricklandi Fahey & Gosliner, 1999.
Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Sanpanich & Duangdee (2018), GBIF (2019a).
Halgerda tessellata (Bergh, 1880). Also reported as: Hypselodoris tessellata (Bergh, 1880). Gulf of Thailand,
Andaman Sea. Reported by: Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich
et al. (2010b, 2013), Rudman (2010), Duangdee et al. (2013b), BEDO (2017c), Sanpanich & Duangdee (2018).
Halgerda willeyi Eliot, 1904. Reported as: Hypselodoris willeyi Eliot, 1904. Gulf of Thailand, Andaman Sea. Reported
by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2013).
Jorunna funebris (Kelaart, 1859). Gulf of Thailand, Andaman Sea. Reported by: Camacho-García & Gosliner (2008),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich &
Duangdee (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017),
GBIF (2019a, 2019b).
Jorunna rubescens Bergh, 1876. Also reported as: Kentodoris rubescens Bergh, 1876. Gulf of Thailand, Andaman
Sea. Reported by: Jensen (1998, 2007), Camacho-García & Gosliner (2008), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c).
Paradoris erythraeensis (Vayssière, 1912). Gulf of Thailand. Reported by: Dayrat (2006, 2010).
Paradoris liturata (Bergh, 1905). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Nabhitabhata
(2009).
Peltodoris murrea (Abraham, 1877). Andaman Sea. Reported by: Dayrat (2010).
Peltodoris rubra (Bergh, 1905). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), Dayrat (2010), Rudman (2010).
Platydoris annulata Dorgan, Valdés & Gosliner, 2002. Gulf of Thailand, Andaman Sea. Reported by: Jensen
(2007), Chavanich et al. (2013), BEDO (2017c).
Platydoris cruenta (Quoy & Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019a, 2019b).
Platydoris dierythros Fahey & Valdés, 2003. Gulf of Thailand. Reported by: Chavanich et al. (2010b, 2013),
Rudman (2010), BEDO (2017c).
Platydoris ellioti (Alder & Hancock, 1864). Gulf of Thailand. Reported by: Rudman (2010), BEDO (2017c).
Platydoris pulchra Eliot, 1904. Andaman Sea. Reported by: Rudman (2010), BEDO (2017c).
Platydoris scabra (Cuvier, 1804). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c).
Rostanga orientalis Rudman & Avern, 1989. Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al.
(2010b, 2013), BEDO (2017c).
Sebadoris fragilis (Alder & Hancock, 1864). Gulf of Thailand, Andaman Sea. Reported by: Dayrat (2010),
Chavanich et al. (2013).
Tayuva lilacina (Gould, 1852). Andaman Sea. Reported by: Chavanich et al. (2010b), BEDO (2017c).
Thordisa villosa (Alder & Hancock, 1864). Also reported as: Thordisa maculigera Bergh, 1877. Gulf of Thailand,
Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007), Gosliner et al. (2008), Nabhitabhata (2009),
Chavanich et al. (2013).
Family Dorididae
Doriopsis pecten (Collingwood, 1881). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2010b,
2013).
Doriopsis granulosa Pease, 1860. Gulf of Thailand. Reported by: BEDO (2017c).
Superfamily Onchidoridoidea
Family Aegiridae
Aegires villosus Farran, 1905. Andaman Sea. Reported by: Gosliner et al. (2008), BEDO (2017c).
Notodoris minor Eliot, 1904. Reported as: Aegires minor (Eliot, 1904). Gulf of Thailand. Reported by: BEDO (2017c).
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The Marine and Estuarine Molluscs of Thailand
Family Goniodorididae
Murphydoris singaporensis Sigurdson, 1991. Gulf of Thailand. Reported by: Swennen & Buatip (2012).
Okenia plebeia Bergh, 1902. Also reported as: Idalia plebeia Bergh, 1902. Gulf of Thailand, Andaman Sea.
Reported by: Bergh (1902), Jensen (1998, 2007), Chavanich et al. (2013).
Trapania reticulata Rudman, 1987. Andaman Sea. Reported by: Rudman (2010).
Trapania miltabrancha Gosliner & Fahey, 2008. No Thai locality provided. Reported by: BEDO (2017c).
Superfamily Phyllidioidea
Family Dendrodorididae
Dendrodoris carbunculosa (Kelaart, 1858). Andaman Sea. Reported by: BEDO (2017c), GBIF (2019a).
Dendrodoris fumata (Rüppell & Leuckart, 1830). Also reported as: Doriopsis rubra (Kelaart, 1858?). Gulf of
Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007), Nabhitabhata (2009), Chavanich
et al. (2010b, 2013), BEDO (2017c).
Dendrodoris krusensternii (Gray, 1850). Also reported as: Dendrodoris denisoni (Angas, 1864). Gulf of Thailand,
Andaman Sea. Reported by: Gosliner et al. (2008), Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra &
Scott (2017).
Dendrodoris nigra (Stimpson, 1855). Also reported as: Doriopsis nigra (Stimpson, 1855). Gulf of Thailand,
Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007), Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017).
Dendrodoris rainfordi Allan, 1932. Gulf of Thailand. Reported by: Gosliner et al. (2008), Rudman (2010), BEDO
(2017c).
Dendrodoris tuberculosa (Quoy & Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al.
(2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), Chavanich et al. (2013), BEDO
(2017c), Mehrotra & Scott (2017), GBIF (2019b).
Doriopsilla carneola (Angus, 1864). Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat et al.
(2008), Chavanich et al. (2010b, 2013), BEDO (2017c).
Doriopsilla pallida Bergh, 1902. Gulf of Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen (1998, 2007),
Nabhitabhata (2009), Chavanich et al. (2013).
Family Phyllidiidae
Ceratophyllidia africana Eliot, 1903. No Thai locality provided. Reported by: Gosliner et al. (2008).
Phyllidia alyta Yonow, 1996. Andaman Sea. Reported by: Duangdee et al. (2011, 2013a), Yonow (2012), BEDO
(2017c), Sanpanich & Duangdee (2018).
Phyllidia coelestis Bergh, 1905. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen (1998,
2007), Wells (2005), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013a,
2013b), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Yonow (2011), BEDO (2017c), Mehrotra & Scott
(2017), GBIF (2019a).
Phyllidia elegans Bergh, 1869. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen (1998,
2007), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF
(2019a).
Phyllidia exquisita Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen
(1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Yonow (2012),
Chavanich et al. (2013), BEDO (2017c), Sanpanich & Duangdee (2018).
Phyllidia marindica (Yonow & Hayward, 1991). Reported as: Fryeria marindica (Yonow & Hayward, 1991);
and Phyllidia (Fryeria) marindica (Yonow & Hayward, 1991). Gulf of Thailand, Andaman Sea. Reported by:
Brunckhorst (1993), Jensen (1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee
et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010),
Yonow (2012), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF (2019a).
Phyllidia ocellata Cuvier, 1804. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Wells (2005),
Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013a, 2013b), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich &
Duangdee (2018), GBIF (2019a, 2019b).
Phyllidia picta Pruvot-Fol, 1957. Also reported as: Fryeria picta Pruvot-Fol, 1957. Gulf of Thailand, Andaman Sea.
Reported by: Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013a, 2013b),
Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO (2017c), Mehrotra & Scott (2017), Sanpanich &
Duangdee (2018).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Phyllidia polkadotsa Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Nabhitabhata (2009), Rudman (2010), BEDO (2017c), GBIF (2019a, 2019b).
Phyllidia varicosa Lamarck, 1801. Also reported as: Phyllidia arabica Ehrenberg, 1831. Gulf of Thailand, Andaman
Sea. Reported by: Brunckhorst (1993), Yonow (1996), Jensen (1998, 2007), Wells (2005), Thamrongnawasawat
et al. (2008), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Chavanich et al. (2010b, 2013),
BEDO (2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018), GBIF (2019a, 2019b).
Phyllidiella annulata Gray, 1853. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2018).
Phyllidiella backeljaui Dominguez, Quintas & Troncoso, 2007. Gulf of Thailand, Andaman Sea. Reported by:
BEDO (2017c).
Phyllidiella lizae Brunckhorst, 1993. Andaman Sea. Reported by: BEDO (2017c).
Phyllidiella nigra (Hasselt, 1824). Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen
(1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013a,
2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2018), Chavanich et al. (2010b, 2013), Rudman
(2010), BEDO (2017c), Mehrotra & Scott (2017), GBIF (2019a).
Phyllidiella pustulosa (Cuvier, 1804). Also reported as: Phyllidiella nobilis Bergh, 1869. Gulf of Thailand, Andaman
Sea. Reported by: Bergh (1902), Brunckhorst (1993), Jensen (1998, 2007), Wells (2005), Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009), Sanpanich &
Duangdee (2009, 2018), Chavanich (2010b), Rudman (2010), BEDO (2017c), Mehrotra & Scott (2017), GBIF
(2019a).
Phyllidiella rudmani Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen
(1998, 2007), Wells (2005), Gosliner et al. (2008), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata
(2009), Chavanich et al. (2010b, 2013), Yonow (2012), BEDO (2017c), Sanpanich & Duangdee (2018), GBIF
(2019a).
Phyllidiella striata (Bergh, 1889). Reported as: Phyllidiopsis striata Bergh, 1889. Gulf of Thailand, Andaman Sea.
Reported by: Brunckhorst (1993), Jensen (1998, 2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Chavanich et al. (2010b), Duangdee et al. (2013a, 2013b), BEDO (2017c), Sanpanich & Duangdee (2018), GBIF
(2019a).
Phyllidiella zeylanica (Kelaart, 1859). Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Yonow
(1996), Jensen (1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009,
2013a, 2013b), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Yonow (2012), Hylleberg & Aungtonya
(2013), BEDO (2017c), Sanpanich & Duangdee (2018), GBIF (2019a).
Phyllidiopsis annae Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993),
Jensen (1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2013a,
2013b), Nabhitabhata (2009), Chavanich et al. (2013), BEDO (2017c), Sanpanich & Duangdee (2018), GBIF
(2019a).
Phyllidiopsis burni Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2018).
Phyllidiopsis cardinalis Bergh, 1875. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO
(2017c), GBIF (2019a).
Phyllidiopsis fissurata Brunckhorst, 1993. Andaman Sea. Reported by: BEDO (2017c).
Phyllidiopsis gemmata Pruvot-Fol, 1957. Andaman Sea. Reported by: Brunckhorst (1993), Jensen (1998, 2007),
Gosliner et al. (2008), Duangdee et al. (2009, 2013a), Nabhitabhata (2009), Yonow (2012), Chavanich et al.
(2013), BEDO (2017c).
Phyllidiopsis krempfi Pruvot-Fol, 1957. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993), Jensen
(1998, 2007), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b,
2013), Yonow (2012), Duangdee et al. (2013a), BEDO (2017c), GBIF (2019a).
Phyllidiopsis loricata (Bergh, 1873). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott (2017).
Phyllidiopsis phiphiensis Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Brunckhorst (1993),
Jensen (1998, 2007), Wells (2005), Gosliner et al. (2008), Thamrongnawasawat et al. (2008), Duangdee et al.
(2009, 2013a), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Rudman (2010), BEDO (2017c), Sanpanich
& Duangdee (2018), GBIF (2019a).
Phyllidiopsis pipeki Brunckhorst, 1993. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Chavanich et al. (2010b, 2013), BEDO (2017c), Sanpanich & Duangdee (2018).
Phyllidiopsis shireenae Brunckhorst, 1990. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Duangdee et al. (2009, 2013a), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), BEDO
(2017c), Sanpanich & Duangdee (2018).
Phyllidiopsis xishaensis (Lin, 1983). Gulf of Thailand, Andaman Sea. Reported by: Yonow (2012), Chavanich et al.
(2013), BEDO (2017c), GBIF (2019a). According to Yonow (2012), the record of Phyllidiella striata (Bergh,
1889) by Brunckhorst (1993) is actually Phyllidiopsis xishaensis.
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The Marine and Estuarine Molluscs of Thailand
Reticulidia suzanneae Valdes & Behrens, 2002. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al.
(2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Chavanich et al. (2010b, 2013), Duangdee
et al. (2013a, 2013b), BEDO (2017c), GBIF (2019a).
Superfamily Polyceroidea
Family Polyceridae
Gymnodoris alba (Bergh, 1877). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009),
Chavanich et al. (2010b; 2013), BEDO (2017c).
Gymnodoris bicolor (Alder & Hancock, 1864). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott
(2017).
Gymnodoris ceylonica (Kelaart, 1858). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2013),
BEDO (2017c), Mehrotra & Scott (2017).
Gymnodoris citrina (Bergh, 1877). Also reported as: Gymnodoris citrine. Gulf of Thailand, Andaman Sea. Reported
by: Chavanich et al. (2010b, 2013), BEDO (2017c).
Gymnodoris impudica (Rüppell & Leuckart, 1828). Reported as: Gymnodoris rubropapulosa (Bergh, 1905). Gulf
of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009), Chavanich et al. (2010b, 2013), BEDO
(2017c), Mehrotra & Scott (2017).
Gymnodoris okinawae Baba, 1936. Andaman Sea. Reported by: BEDO (2017c).
Gymnodoris pattani Swennen, 1996. Gulf of Thailand. Reported by: Swennen (1996), Jensen (1998), Chavanich
et al. (2013), GBIF (2019b).
Gymnodoris striata (Eliot, 1908). Reported as: Gymnodoris (Anlogium) striata (Eliot, 1908). Andaman Sea.
Reported by: Jensen (1998), Chavanich et al. (2013), BEDO (2017c).
Gymnodoris tuberculosa Knutson & Gosliner, 2014. Andaman Sea. Reported by: BEDO (2017c).
Kalinga ornata Alder & Hancock, 1864. Andaman Sea. Reported by: Chavanich et al. (2013).
Nembrotha purpureolineata O’Donoghue, 1924. Andaman Sea. Reported by: BEDO (2017c).
Plocamopherus margaretae Vallès & Gosliner, 2006. Reported as: Plocamopherus margaritae Vallès & Gosliner,
2006. Andaman Sea. Reported by: Gosliner et al. (2008).
Roboastra arika Burn, 1967. Reported as: Roboastra arika. No Thai locality provided. Reported by: Chavanich
et al. (2010b). Considered a taxon inquirendum by WoRMS Editorial Board (2019).
Roboastra gracilis (Bergh, 1877). Andaman Sea. Reported by: Gosliner et al. (2008), Chavanich et al. (2010b,
2013), Duangdee et al. (2013b), BEDO (2017c).
Tambja affinis (Eliot, 1904). Andaman Sea. Reported by: Gosliner et al. (2008), Chavanich et al. (2013), BEDO
(2017c).
Tambja amakusana Baba, 1987. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017c), Mehrotra & Scott
(2017).
Tambja gabrielae Pola, Cervera & Gosliner, 2005. Andaman Sea. Reported by: Gosliner et al. (2008).
Tambja morosa (Bergh, 1877). Andaman Sea. Reported by: Gosliner et al. (2008), Chavanich et al. (2010b, 2013),
BEDO (2017c).
Tambja victoriae Pola, Cervera & Gosliner, 2005. Andaman Sea. Reported by: Chavanich et al. (2013), BEDO
(2017c).
Thecacera pennigera (Montagu, 1815). Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2013),
BEDO (2017c), Mehrotra & Scott (2017).
Tyrannodoris luteolineata (Baba, 1936). Also reported as: Roboastra luteolineata (Baba, 1936). Andaman Sea.
Reported by: Chavanich et al. (2010b, 2013), BEDO (2017c).
Order Pleurobranchida
Superfamily Pleurobranchoidea
Family Pleurobranchaeidae
Euselenops luniceps (Cuvier, 1816). Andaman Sea. Reported by: Jensen (2007).
Pleurobranchaea brockii Bergh, 1897. Andaman Sea. Reported by: Jensen (2007).
Family Pleurobranchidae
Berthella martensi Pilsbry, 1896. Gulf of Thailand, Andaman Sea. Reported by: Gosliner et al. (2008),
Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Rudman (2010), BEDO (2017c).
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Pleurobranchus forskalii Rüppell & Leuckart, 1828. Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra &
Scott (2017).
Pleurobranchus grandis Pease, 1868. Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat
et al. (2008), Nabhitabhata (2009), BEDO (2017c).
Subterclass Ringipleura
Superorder Ringiculimorpha
Superfamily Ringiculoidea
Family Ringiculidae
Ringicula gouldi Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004).
Ringicula propinquans Hinds, 1844. Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014).
Subterclass Tectipleura
Superorder Acochlidiimorpha
Superfamily Acochlidioidea
Family Aitengidae
Aiteng ater Swennen & Buatip, 2009. Gulf of Thailand. Reported by: Swennen & Buatip (2009).
Order Aplysiida
Superfamily Akeroidea
Family Akeridae
Akera constricta Kuroda, 1947. Andaman Sea. Reported by: BEDO (2017b). This species cannot be confirmed
through WoRMS Editorial Board (2019), but is shown in DiscoverLife (2020).
Akera soluta (Gmelin, 1791). Andaman Sea. Reported by: Nateewathana et al. (1981), Jensen (1998, 2007),
Valdés & Barwick (2005).
Superfamily Aplysioidea
Family Aplysiidae
Aplysia dactylomela Rang, 1828. Gulf of Thailand. Reported by: BEDO (2017c).
Aplysia euchlora (Adams in M. E. Gray, 1850). Andaman Sea. Reported by: Jensen (1998), Rudman (2010).
Aplysia kurodai Baba, 1937. Gulf of Thailand. Reported by: BEDO (2017c).
Aplysia parvula Mörch, 1863. Gulf of Thailand. Reported by: Thamrongnawasawat et al. (2008), Duangdee et al.
(2011, 2013b).
Bursatella leachii Blainville, 1817. Also reported as: Aclesia ocelligera Bergh, 1902. Gulf of Thailand, Andaman
Sea. Reported by: Bergh (1902), Jensen (1998, 2007), Rudman (2010), BEDO (2017c).
Dolabella auricularia (Lightfoot, 1786). Gulf of Thailand. Reported by: Rudman (2010).
Petalifera petalifera (Rang, 1828). Also reported as: Aplysiella unguifera (Rang, 1828). Andaman Sea. Reported
by: Bergh (1902), Jensen (1998, 2007), Rudman (2010).
Phyllaplysia incerta (Bergh, 1902). Reported as: Aplysiella incerta Bergh, 1902. Andaman Sea. Reported by:
Bergh (1902), Jensen (1998, 2007), Rudman (2010).
Stylocheilus longicauda (Quoy & Gaimard, 1825). Gulf of Thailand, Andaman Sea. Reported by: Jensen (1998),
Hylleberg & Aungtonya (2013), BEDO (2017c), Mehrotra & Scott (2017).
Stylocheilus striatus (Quoy & Gaimard, 1832). Gulf of Thailand. Reported by: BEDO (2017c), Mehrotra & Scott
(2017), GBIF (2019b).
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The Marine and Estuarine Molluscs of Thailand
Order Cephalaspidea
Superfamily Bulloidea
Family Bullidae
Bulla ampulla Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889), Nateewathana
et al. (1981), Wells (2005), Malaquias & Reid (2008), Duangdee et al. (2009), Sanpanich & Duangdee (2013),
BEDO (2017b), GBIF (2019a).
Bulla striata Bruguière, 1792. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Duangdee et al. (2009, 2013b),
Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Bulla vernicosa Gould, 1859. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009, 2011, 2013b),
Nabhitabhata (2009).
Family Retusidae
Pyrunculus pyriformis (A. Adams, 1850). Reported as: Retusa (Pyriformis) pyriformis (A. Adams, 1855). Gulf of
Thailand. Reported by: Gemert (2003).
Retusa concentrica (A. Adams, 1850). Also reported as: Retusa (Coleophysis) concentrica (A. Adams, 1850). Gulf
of Thailand. Reported by: Gemert (2003), Robba et al. (2004).
Retusa minima Yamakawa, 1911. Gulf of Thailand. Reported by: Robba et al. (2004).
Retusa succincta (A. Adams, 1862). Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014).
Family Tornatinidae
Truncacteocina coarctata (A. Adams, 1850). Reported as: Acteocina coarctata (A. Adams, 1850). Gulf of Thailand.
Reported by: Gemert (2003).
Acteocina decoratoides Habe, 1955. Reported as: Didontoglossa decoratoides Habe, 1955. Gulf of Thailand.
Reported by: Robba et al. (2004).
Acteocina gordonis (Yokoyama, 1927). Reported as: Tornatina gordonis (Yokoyama, 1927). Gulf of Thailand.
Reported by: Robba et al. (2004).
Acteocina inconspicua Olsson & McGinty, 1958. Reported as: Acteocina inconspicua (Adams, 1872). Gulf of
Thailand. Reported by: Robba et al. (2004).
Superfamily Cylichnoidea
Family Cylichnidae
Cylichna involuta (A. Adams, 1850). Reported as: Eocylichna involuta (A. Adams, 1850). Gulf of Thailand. Reported
by: Gemert (2003).
Cylichna modesta (Thiele, 1925). Also reported as: Adamnestia modesta (Thiele, 1925). Gulf of Thailand.
Reported by: Swennen et al. (2001), Robba et al. (2004), Negri et al. (2014).
Decorifer longispiratus (Yamakawa, 1911). Gulf of Thailand. Reported by: Robba et al. (2004). This species
cannot be confirmed through WoRMS Editorial Board (2019), but is listed by Robba et al. (2004).
Superfamily Haminoeoidea
Family Haminoeidae
Aliculastrum cylindricum (Helbling, 1779). Reported as: Atys cylindricus Helbling, 1779. Gulf of Thailand.
Reported by: Morlet (1889), Gemert (2003), Nabhitabhata (2009), Sanpanich & Duangdee (2013), GBIF (2019b).
Aliculastrum debile (Pease, 1860). Gulf of Thailand. Reported by: Mehrotra & Scott (2017).
Atys naucum (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Bergh
(1902), Nateewathana et al. (1981), Kurozumi et al. (1989), Jensen (1998, 2007), Gemert (2003), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), BEDO (2017b).
Bakawan rotundata (A. Adams, 1850). Also reported as: Haloa cf. fusca (A. Adams, 1850). Gulf of Thailand.
Reported by: Oskars et al. (2019), Oskars & Malaquias (2019).
Diniatys monodonta (A. Adams, 1850). Gulf of Thailand. Reported by: Gemert (2003).
Haloa natalensis (Krauss, 1848). No Thai locality located. “Kaempam” was cited in the original description, but
this locality could not be found. Reported by: Oskars & Malaquias (2019).
Haminoea cymbalum Quoy & Gaimard, 1833. Gulf of Thailand, Andaman Sea. Reported by: KUMF,
Thamrongnawasawat et al. (2008), Nabhitabhata (2009).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Haminoea japonica Pilsbry, 1895. Andaman Sea. Reported by: GBIF (2019a).
Haminoea perrieri Morlet, 1889. Gulf of Thailand. Reported by: Morlet (1889).
Liloa mongii (Audouin, 1826). Reported as: Liloa curta (A. Adams, 1850). Gulf of Thailand. Reported by: Gemert
(2003).
Liloa porcellana (Gould, 1859). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et
al. (2011), Sanpanich & Duangdee (2013).
Limulatys constrictus Habe, 1952. Gulf of Thailand. Reported by: Robba et al. (2004).
Nipponatys volvulina (A. Adams, 1862). Gulf of Thailand. Reported by: Robba et al. (2004).
Smaragdinella calyculata (Broderip & G. B. Sowerby I, 1829). Andaman Sea. Reported by: Sørensen (2012).
Superfamily Philinoidea
Family Aglajidae
Chelidonura castanea Yonow, 1994. Andaman Sea. Reported by: Gosliner et al. (2008).
Chelidonura pallida Risbec, 1951. Andaman Sea. Reported by: Nabhitabhata (2009), Rudman (2010), Duangdee
et al. (2013a).
Chelidonura punctata Eliot, 1903. Andaman Sea. Reported by: Gosliner et al. (2008), Thamrongnawasawat et al.
(2008), Duangdee et al. (2009, 2013a), Nabhitabhata (2009), Rudman (2010), BEDO (2017c), Tibiriçá &
Malaquias (2017), Sanpanich & Duangdee (2018), GBIF (2019a).
Melanochlamys papillata Gosliner, 1990. Andaman Sea. Reported by: Jensen (1998, 2007).
Philinopsis speciosa Pease, 1860. Also reported as: Philinopsis cyanea Martens, 1879. Gulf of Thailand, Andaman
Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata (2009), BEDO (2017c), Mehrotra & Scott
(2017).
Tubulophilinopsis gardineri (Eliot, 1903). Reported as: Philinopsis gardineri (Eliot, 1903). Andaman Sea. Reported
by: Duangdee et al. (2013b), Sanpanich & Duangdee (2018).
Tubulophilinopsis lineolata (H. Adams & A. Adams, 1854). Also reported as: Dorideum lineolatum (H. Adams &
A. Adams, 1854); and Philinopsis lineolata (H. Adams & A. Adams, 1854). Gulf of Thailand, Andaman Sea. Reported
by: Bergh (1902), Jensen (1998, 2007), BEDO (2017c).
Tubulophilinopsis pilsbryi (Eliot, 1900). Also reported as: Philinopsis pilsbryi (Eliot, 1900). Gulf of Thailand,
Andaman Sea. Reported by: Duangdee et al. (2009), Nabhitabhata (2009), BEDO (2017c), Mehrotra & Scott
(2017), Sanpanich & Duangdee (2018).
Family Philinidae
Philine orca Gosliner, 1988. Gulf of Thailand. Reported by: Gemert (2003).
Philine orientalis A. Adams, 1855. Reported as: Philine argentata Gould, 1859. Andaman Sea. Reported by: KUMF,
Jensen (2007), Nabhitabhata (2009).
Philine quadripartita Ascanius, 1772. Gulf of Thailand. Reported by: Jensen (1998).
Philine striolata A. Adams, 1862. Gulf of Thailand. Reported by: Gemert (2003).
Superorder Eupulmonata
Order Ellobiida
Superfamily Ellobioidea
Family Ellobiidae
Allochroa layardi (H. Adams & A. Adams, 1855). Reported as: Allochroa pfeifferiana (Dunker). Andaman Sea.
Reported by: Nateewathana et al. (1981).
Auriculastra semiplicata (H. Adams & A. Adams, 1854). Reported as: Cylindrotis quadrasi Möllendorff, 1895.
No Thai locality provided. Reported by: Higo et al. (1999).
Auriculastra siamensis (Brandt, 1974). Gulf of Thailand. Reported by: Raven & Vermeulen (2007).
Auriculastra subula (Quoy & Gaimard, 1832). Andaman Sea. Reported by: Higo et al. (1999), Raven & Vermeulen
(2007), Romero et al. (2016), J. D. Taylor et al. (unpublished data).
Cassidula aurisfelis (Bruguière, 1789). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Sri-aroon et al. (2005), Sanpanich & Duangdee (2013), Dechruksa et al. (2014), Romero et al. (2016), BEDO
(2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Cassidula multiplicata (Martens, 1865). Gulf of Thailand, Andaman Sea. Reported by: Sri-aroon et al. (2005),
Dechruksa et al. (2014), J. D. Taylor et al. (unpublished data).
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Cassidula nucleus (Gmelin, 1791). Also reported as: Auricula mustelina Deshayes, 1830; and Cassidula mustelina
(Deshayes, 1830). Andaman Sea. Reported by: Morlet (1889), Higo et al. (1999), Sri-aroon et al. (2005, 2006),
Dechruksa et al. (2014), BEDO (2017b).
Ellobium aurisjudae (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Subba Rao & Dey (2000), Gemert (2003), Sri-aroon et al. (2005), Duangdee et al. (2009), Hylleberg & Aungtonya
(2013), Sanpanich & Duangdee (2013), Dechruksa et al. (2014), Romero et al. (2016), BEDO (2017b), GBIF
(2019a, 2019b), J. D. Taylor et al. (unpublished data).
Ellobium aurismidae (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, Gemert (2003),
Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b).
Laemodonta punctigera (H. Adams & A. Adams, 1854). Gulf of Thailand, Andaman Sea. Sri-aroon et al. (2005),
GBIF (2019b).
Laemodonta siamensis (Morlet, 1875). Gulf of Thailand, Andaman Sea. Reported by: Higo et al. (1999), Raven &
Vermeulen (2007), Sawatna et al. (2012), Dechruksa et al. (2014).
Laemodonta typica (H. Adams & A. Adams, 1854). Gulf of Thailand. Reported by: Gemert (2003), Sawatna et al.
(2012).
Melampus adamsianus L. Pfeiffer, 1855. Gulf of Thailand. Reported by: Raven & Vermeulen (2007).
Melampus fasciatus (Deshayes, 1830). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Dechruksa et al. (2014), BEDO (2017b), GBIF (2019a).
Melampus flavus (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017b).
Melampus granifer (Mousson, 1849). Gulf of Thailand. Reported by: Premcharoen et al. (2016).
Melampus nucleolus Martens, 1865. Reported as: Micromelampus (Micromelampus) nucleolus Martens, 1865. No
Thai locality provided. Reported by: Higo et al. (1999).
Melampus cf. nuxeastaneus Kuroda, 1949. Gulf of Thailand. Reported by: Gemert (2003).
Melampus pulchellus (Petit de la Saussaye, 1843). Andaman Sea. Reported by: Romero et al. (2016).
Melampus sincaporensis L. Pfeiffer, 1855. Also reported as: Melampus siamensis E. von Martens, 1865. Gulf of
Thailand, Andaman Sea. Reported by: Gemert (2003), Sri-aroon et al. (2005), Raven & Vermeulen (2007).
Melampus zonatus (Anton, 1847). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
BEDO (2017b). This species cannot be confirmed through WoRMS Editorial Board (2019), but is recorded by
Galli (2015) and DiscoverLife (2020).
Pythia plicata (Gray, 1825). No Thai locality provided. Reported by: Subba Rao & Dey (2000), Gemert (2003).
Pythia reeveana Pfeiffer, 1853. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2009), Sanpanich
& Duangdee (2013), Dechruksa et al. (2014).
Order Systellommatophora
Superfamily Onchidioidea
Family Onchidiidae
Morrisonietta gracilis Brandt, 1968. Gulf of Thailand. Reported by: GBIF (2019b).
Onchidium stuxbergi (Westerlund, 1883). Reported as: Labbella ajuthiae (Labbé, 1935). Gulf of Thailand. Reported
by: Labbé (1935), Dayrat et al. (2016, 2017, 2019).
Onchidium typhae Buchannan, 1800. Andaman Sea. Reported by: Dayrat et al. (2016) as possibly occurring on
the west coast of Thailand.
Platevindex martensi (Semper, 1885). No Thai locality provided. Reported by: Dayrat et al. (2017).
Superorder Hygrophila
Order Pteropoda
Suborder Euthecosomata
Superfamily Limacinoidea
Family Limacinidae
Heliconoides inflatus (d’Orbigny, 1835). Reported as: Limacina inflata (d’Orbigny, 1834). Gulf of Thailand,
Andaman Sea. Reported by: Rottman (1976), Vongpanich (2001).
Limacina bulimoides (d’Orbigny, 1835). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976),
Vongpanich (2001).
Limacina trochiformis (d’Orbigny, 1835). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976),
Vongpanich (2001).
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Superfamily Cavolinioidea
Family Cavoliniidae
Cavolinia gibbosa (d’Orbigny, 1835). Andaman Sea. Reported by: Vongpanich (2001), BEDO (2017b).
Cavolinia globulosa Gray, 1850. Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich (2001),
BEDO (2017b).
Cavolinia inflexa (Lesueur, 1813). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001), BEDO (2017b).
Cavolinia tridentata (Forsskål in Niebuhr, 1775). Andaman Sea. Reported by: Vongpanich (2001), BEDO (2017b).
Cavolinia uncinata (d’Orbigny, 1835). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001), BEDO (2017b).
Cuvierina columnella (Rang, 1827). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001).
Diacavolinia flexipes van der Spoel, Bleeker & Kobayashi, 1993. Gulf of Thailand. Reported by: Gemert (2003),
Robba et al. (2007).
Diacavolinia longirostris (Lesueur, 1821). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976),
Vongpanich (2001), BEDO (2017b, 2017e).
Diacria quadridentata (Lesueur, 1821). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001), BEDO (2017b, 2017e).
Diacria trispinosa (Lesueur, 1821). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001), BEDO (2017b).
Family Cliidae
Clio cuspidata (Bosc, 1801). Gulf of Thailand. Reported by: Rottman (1976).
Clio pyramidata Linnaeus, 1767. Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich
(2001), BEDO (2017b).
Clio recurva (Children, 1823). Gulf of Thailand. Reported by: Rottman (1976).
Family Creseidae
Creseis acicula (Rang, 1828). Gulf of Thailand, Andaman Sea. Reported by: Vongpanich (2001), Gemert (2003),
BEDO (2017b).
Creseis chierchiae (Boas, 1886). Reported as: Creseis bulgia Sakthivel, 1974. Gulf of Thailand, Andaman Sea.
Reported by: Rottman (1976), Vongpanich (2001).
Creseis conica Eschscholtz, 1829. Reported as: Creseis virgula conica Eschscholtz, 1829. Gulf of Thailand, Andaman
Sea. Reported by: Vongpanich (2001).
Creseis virgula (Rang, 1828). Reported as: Creseis virgula virgula (Rang, 1828). Gulf of Thailand, Andaman Sea.
Reported by: Rottman (1976), Vongpanich (2001).
Hyalocylis striata (Rang, 1828). Gulf of Thailand, Andaman Sea. Reported by: Rottman (1976), Vongpanich (2001),
BEDO (2017b).
Styliola subula (Quoy & Gaimard, 1827). Gulf of Thailand. Reported by: Rottman (1976).
Superorder Pylopulmonata
Order Runcinida
Superfamily Amphiboloidea
Family Amphibolidae
Salinator burmana (Blanford, 1867). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
GBIF (2019b). This species is listed by WoRMS Editorial Board (2019) as a taxon inquirendum.
Superfamily Pyramidelloidea
Family Amathinidae
Amathina bicarinata Pease, 1861. Reported as: Capulus bicarinatus Pease, 1861. Gulf of Thailand. Reported by:
Gemert (2003).
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Amathina oyamai (Masuda & Noda, 1976). Reported as: Amathina oyamai oyamai (Masuda & Noda, 1976). Gulf
of Thailand. Reported by: Negri et al. (2014).
Amathina tricarinata (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), BEDO (2017b).
Leucotina speciosa (A. Adams, 1853). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Monotygma amoena (A. Adams, 1853). Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al.
(2007), Sawatna et al. (2012).
Family Pyramidellidae
Asmunda metula (A. Adams, 1860). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Asmunda secta (Saurin, 1958). Reported as: Turbonilla (Asmunda) secta Saurin, 1958. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Asmunda tenuicostata (Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004). Gulf of Thailand. Reported
by: Robba et al. (2004).
Asmunda tribulationis (Hedley, 1909). Reported as: Zaphella tribulationis (Hedley, 1909). Gulf of Thailand.
Reported by: Robba et al. (2007).
Babella hastula Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Babella funiculata Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Babella khmeriana (Saurin, 1962). Reported as: Numaegilina khmeriana Saurin, 1962. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Babella obliquissima (Saurin, 1959). Reported as: Numaegilina obliquissima Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Besla canaensis Saurin, 1959. Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Besla cossmanni (Hornung & Mermod, 1924). Gulf of Thailand. Reported by: Robba et al. (2004).
Besla dheeradiloki Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Chaiwathee et al. (2007), Sawatna et al. (2012), Negri et al. (2014).
Besla henriettae Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Besla joae Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Besla ranae Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Besla reamensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Brachystomia minutiovum (Nomura, 1936). Reported as: Ondina minutiovum (Nomura, 1936). Gulf of Thailand.
Reported by: Robba et al. (2004).
Careliopsis flexuosa Saurin, 1962. Reported as: Careliopsis flexuosus Saurin, 1962. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Chrysallida chetelati Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Chrysallida culaoniana Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Chrysallida foveata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Sawatna et al. (2012), Galli (2015).
Chrysallida genouillyi Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Chrysallida phanthietina Saurin, 1958. Reported as: Chrysallida (Chrysallida) phanthietina Saurin, 1958. Gulf of
Thailand. Reported by: Saurin (1962), Robba et al. (2004), Gemert (2016).
Chrysallida pura (Saurin, 1962). Reported as: Pyrgulina (Partulida) pura Saurin, 1962. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Chrysallida saurini Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Negri et al. (2014), Galli (2015).
Chrysallida stupa Hori & Fukuda, 1999. Reported as: Miralda stupa (Hori & Fukuda, 1999). Gulf of Thailand.
Reported by: Robba et al. (2007), Galli (2015).
Chrysallida tribulationis (C. Hedley, 1909). Gulf of Thailand. Reported by: Galli (2015).
Chrysallida trifuniculata (Saurin, 1962). Reported as: Tragula trifuniculata Saurin, 1962. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Cingulina archimedea Melvill, 1896. Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007),
Negri et al. (2014).
Cingulina dussaulti Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Cingulina inaequalis Saurin, 1958. Gulf of Thailand. Reported by: Saurin (1962), Robba et al. (2004), Sawatna et
al. (2012), Gemert (2016).
Cossmannica bancoensis Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Cossmannica behainei Saurin, 1959. Gulf of Thailand. Reported by: Robba et al. (2004).
Cossmannica catinati Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Costosyrnola cuabeina Saurin, 1959. Gulf of Thailand. Reported by: Robba et al. (2004).
Costosyrnola culaopagi Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Robba et al. (2004), Gemert
(2016).
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Costosyrnola gradata Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Costosyrnola thailandica Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004), Negri et al. (2014), Galli (2015).
Costosyrnola timida Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Ebala torquata (Saurin, 1962). Reported as: Anisocycla torquatum Saurin, 1962. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Egila curtisensis Laseron, 1959. Gulf of Thailand. Reported by: Robba et al. (2004).
Egilina babellina Saurin, 1958. Reported as: Egilina (Egilina) babellina Saurin, 1958. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Egilina callista (Melvill, 1893). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Egilina tenuis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Elodiamea excavata Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016). Listed by WoRMS
Editorial Board (2019) as a taxon inquirendum.
Eulimastoma eutropia (Melvill, 1899). Also reported as: Odostomia eutropia Melvill, 1899. Gulf of Thailand.
Reported by: Robba et al. (2004), Negri et al. (2014).
Eulimella disciformis Peñas & Rolán, 2016. Reported as: Ebalina angusta Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Eulimella pyrgoidella Saurin, 1959. Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014).
Eulimella siamensis Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004).
Eulimella thalensis Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004).
Exesilla dextra (Saurin, 1959). Gulf of Thailand. Reported by: Robba et al. (2007).
Exesilla laseroni Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Sawatna et al. (2012).
Herviera gliriella (Melvill & Standen, 1896). Reported as: Odostomella purpurea Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Hinemoa laxefuniculata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004), Sawatna et al. (2012), Negri et al. (2014).
Kleinella delicatula Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Leucotina ampulla Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Linopyrga primitractus (Saurin, 1959). Reported as: Pyrgulina (Linopyrga) primitractus Saurin, 1959. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Longchaeus acus (Gmelin, 1791). Andaman Sea. Reported by: Duangdee et al. (2011), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a).
Longchaeus insularum (Pilsbry, 1922). Reported as: Longchaeus canaliculatus (Sowerby, 1873). Gulf of Thailand.
Reported by: Sanpanich & Duangdee (2013).
Marginodostomia prava (Saurin, 1962). Reported as: Odostomia (Cyclodostomia) prava Saurin, 1962. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Marginodostomia striatissima Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand.
Reported by: Robba et al. (2004), Chaiwathee et al. (2007), Galli (2015).
Megastomia ampliata (Saurin, 1962). Reported as: Odostomia (Megastomia) ampliata Saurin, 1962. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Megastomia binhdinhensis (Saurin, 1958). Reported as: Odostomia (Megastomia) binhdinhensis Saurin, 1958.
Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Megastomia martinae (Saurin, 1959). Reported as: Odostomia (Megastomia) martinae Saurin, 1959. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Megastomia metayei (Saurin, 1959). Reported as: Odostomia (Megastomia) metayei Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Megastomia saurini (Corgan & Van Aartsen, 1998). Reported as: Odostomia (Megastomia) elata Saurin, 1958.
Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Megastomia serenei (Saurin, 1959). Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007).
Megastomia supramarginata (Saurin, 1959). Reported as: Odostomia (Megastomia) supramarginata Saurin,
1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Megastomia unifilosa (Saurin, 1962). Reported as: Odostomia (Megastomia) unifilosa Saurin, 1962. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Milda ventricosa (Guérin, 1831). Andaman Sea. Also reported as: Pyramidella ventricosa (Guérin). Reported by:
Nateewathana et al. (1981), Chaiwathee et al. (2007), Hylleberg & Aungtonya (2013), BEDO (2017b).
Miralda diadema (A. Adams, 1860). Gulf of Thailand. Reported by: Robba et al. (2007).
Miralda franciscae Saurin, 1958. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Miralda pretiosior Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Miralda oscillaeformis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
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The Marine and Estuarine Molluscs of Thailand
Miralda primaliciae Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Miralda scopulorum (R. B. Watson, 1886). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al. (2012).
Miralda senex (Hedley, 1902). Gulf of Thailand. Reported by: Robba et al. (2007).
Moerchia morleti P. Fischer, 1877. Reported as: Moerchiella morleti (P. Fischer, 1877). Gulf of Thailand. Reported
by: Gemert (2003).
Morrisonietta spiralis Brandt, 1968. Gulf of Thailand. Reported by: Negri et al. (2014).
Mumiola tessellata A. Adams, 1863. Also reported as: Mumiola superba Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Robba et al. (2004), Chaiwathee et al. (2007), Sawatna et al. (2012).
Nisipyrgiscus zetemia (Melvill, 1910). Reported as: Pyrgiscilla zetemia (Melvill, 1910). Gulf of Thailand.
Reported by: Robba et al. (2007).
Nisiturris obliqua (Saurin, 1959). Reported as: Chemnitzia obliqua Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Noemiamea edentata (Saurin, 1959). Reported as: Odostomia (Oda) edentata Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Odetta hians Saurin, 1961. Reported as: Menestho (Odetta) hians (Saurin, 1961). Reported by: Saurin (1962),
Gemert (2016). This species cannot be confirmed through WoRMS Editorial Board (2019), but is listed on OBIS
(2020).
Odostomella germaini (Dautzenberg & Fischer, 1907). Reported as: Salassia germaini (Dautzenberg & Fischer,
1906). Gulf of Thailand. Reported by: Robba et al. (2004).
Odostomia biformis Saurin, 1962. Reported as: Odostomia (Brachystomia) biformis Saurin, 1962. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Odostomia bullula Gould, 1861. Gulf of Thailand. Reported by: Robba et al. (2004).
Odostomia carinata H. Adams, 1873. Gulf of Thailand. Reported by: Robba et al. (2004).
Odostomia caudana (Saurin, 1959). Reported as: Odostomia (Megastomia) caudana (Saurin, 1959). Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Odostomia decouxi Saurin, 1959. Reported as: Odostomia (Odostomia) decouxi Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Odostomia garcerii Saurin, 1959. Reported as: Odostomia (Odostomia) garcerii Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Odostomia honis Saurin, 1959. Reported as: Odostomia (Odostomia) honis Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Odostomia khanhoana Saurin, 1959. Reported as: Odostomia (Odostomia) khanhoana Saurin, 1959. Gulf of
Thailand. Reported by: Saurin (1962), Gemert (2016).
Odostomia magnumos Saurin, 1962. Reported as: Odostomia (Heida) magnumos Saurin, 1962. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Odostomia cf. sperabilis (Hedley, 1909). Andaman Sea. Reported by: BEDO (2017e).
Odostomia sublimpida Yokoyama, 1920. Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al.
(2007), Sawatna et al. (2012), Negri et al. (2014), Gemert (2016).
Oscilla jocosa Melvill, 1904. Gulf of Thailand. Reported by: Robba et al. (2007).
Otopleura auriscati (Holten, 1802). Gulf of Thailand, Andaman Sea. Reported by: Cernohorsky (1972),
Nateewathana, et al. (1981), Duangdee et al. (2011), Hylleberg & Aungtonya (2013), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a).
Oscilla obtusantis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Oscilla ongcopensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Oscilla voorwindei (Laseron, 1959). Reported as: Toscilla voorwindei (Laseron, 1959). Gulf of Thailand, Andaman
Sea. Reported by: BEDO (2017e).
Parodostomia treina (Saurin, 1959). Reported as: Brachystomia treina (Saurin, 1959). Gulf of Thailand. Reported
by: Robba et al. (2004).
Polemicella aartseni Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004), Sawatna et al. (2012), Negri et al. (2014), Galli (2015).
Polemicella piscatorum Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Robba et al. (2004), Sawatna
et al. (2012), Gemert (2016).
Polemicella saurini Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Sawatna et al. (2012).
Puposyrnola basistriata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004).
Puposyrnola tracta Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Puposyrnola vienae Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyramidella dolabrata (Linnaeus, 1758). Also reported as: Pyramidella acus (Gmelin, 1791); and Pyramidella
terebellum (Müller). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana, et al. (1981), Nabhitabhata
(2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019b).
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Pyramidella maculosa Lamarck, 1822. Also reported as: Pyramidella sulcata (A. Adams, 1864). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, Duangdee et al. (2011), Sanpanich & Duangdee (2013, 2018).
Pyrgiscus basicinctus (Saurin, 1959). Reported as: Pyrgiscilla basicincta Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Pyrgiscus erica (Thiele, 1925). Gulf of Thailand. Reported by: Robba et al. (2004), Negri et al. (2014). This species
cannot be confirmed through WoRMS Editorial Board (2019), but is listed on OBIS (2020).
Pyrgiscus renatae (Saurin, 1959). Reported as: Pyrgiscilla renatae Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Pyrgiscus semistriata (Saurin, 1962). Reported as: Pyrgiscilla semistriata Saurin, 1962. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Pyrgiscus siamensis (Saurin, 1962). Reported as: Pyrgiscilla siamensis Saurin, 1962. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Pyrgiscus tabulae (Saurin, 1962). Reported as: Pyrgiscilla tabulae Saurin, 1962. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Pyrgiscus thuanae (Saurin, 1959). Reported as: Pyrgiscilla thuanae Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Pyrgulina amabilis Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962).
Pyrgulina buriti Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina cambodgiensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina cheveyi Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina epentromidea Melvill, 1899. Gulf of Thailand. Reported by: Saurin (1958), Gemert (2016).
Pyrgulina kampotensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina kepensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina krempfi Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina melvilli Dautzenberg & Fischer, 1907. Gulf of Thailand. Reported by: Sawatna et al. (2012).
Pyrgulina noviportus Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Pyrgulina phohaiensis Saurin, 1958. Also reported as: Pyrgulina (Pyrgulina) phohaiensis Saurin, 1958. Gulf of
Thailand. Reported by: Saurin (1962), Galli (2015), Gemert (2016).
Pyrgulina pupaeformis (Souverbie, 1865). Reported as: Pyrgulina montbruni Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Pyrgulina tenerrima (Melvill, 1906). Reported as: Pyrgulina nigraerupis Saurin, 1959; and Chrysallida phohaiensis
(Saurin, 1958). Gulf of Thailand. Reported by: Robba et al. (2007).
Pyrgulina trochiformis Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Quirella humilis (Preston, 1905). Reported as: Chemnitzia humilis (Yokoyama, 1924); and Turbonilla humilis
(Yokoyama, 1924). Gulf of Thailand. Reported by: Robba et al. (2004, 2007), Sawatna et al. (2012), Negri et al.
(2014).
Quirella lyngei Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Sawatna et al. (2012).
Salassia bicarinata Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported by:
Robba et al. (2004), Negri et al. (2014).
Sinuatodostomia labunensis Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand.
Reported by: Robba et al. (2004), Galli (2015).
Sinuatodostomia somsaki Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand.
Reported by: Robba et al. (2004), Galli (2015).
Spiralinella muinaiensis Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Styloptygma fromageti Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Syrnola baygiongina Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Syrnola elegans (A. Adams, 1854). Reported as: Syrnola vietnamica Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Robba et al. (2004), Gemert (2016).
Syrnola macella Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Syrnola praecostulata Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Syrnola quaelibet Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Syrnola strigulata (A. Adams, 1863). Gulf of Thailand. Reported by: Robba et al. (2007).
Tiberia cathaysiae Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Tiberia grimaudi Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Robba et al. (2004), Chaiwathee
et al. (2007), Gemert (2016).
Tiberia ngani Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Tiberia semicostata Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Tiberia thaii Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Tibersyrnola cinnamomea (A. Adams, 1863). Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee
et al. (2007).
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The Marine and Estuarine Molluscs of Thailand
Turbolidium uniliratum (Saurin, 1959). Reported as: Tragula unilirata Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Turbonilla abbotti (Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004). Gulf of Thailand. Reported
by: Robba et al. (2004).
Turbonilla ailhaudi (Saurin, 1959). Reported as: Chemnitzia ailhaudi (Saurin, 1959). Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Turbonilla ambigua (Saurin, 1962). Reported as: Chemnitzia ambigua Saurin, 1962. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Turbonilla annamitica (Saurin, 1959). Reported as: Chemnitzia annamitica Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Turbonilla biangulata (Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004). Gulf of Thailand. Reported
by: Robba et al. (2004), Sawatna et al. (2012).
Turbonilla bonellii (Saurin, 1959). Reported as: Chemnitzia bonellii Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Turbonilla crassa Nomura, 1936. Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007),
Sawatna et al. (2012).
Turbonilla cummingi Hori & Okutani, 1997. Reported as: Chemnitzia cummingi (Hori & Okutani, 1997). Gulf of
Thailand. Reported by: Robba et al. (2007).
Turbonilla dunkeriformis (Saurin, 1959). Reported as: Chemnitzia dunkeriformis Saurin, 1959. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Turbonilla funiculata de Folin, 1868. Gulf of Thailand. Reported by: Robba et al. (2004).
Turbonilla geraudiei (Saurin, 1959). Reported as: Chemnitzia geraudiei Saurin, 1959. Gulf of Thailand. Reported
by: Saurin (1962), Gemert (2016).
Turbonilla holocenica Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004. Gulf of Thailand. Reported
by: Robba et al. (2004).
Turbonilla integra (Saurin, 1959). Reported as: Chemnitzia integra Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Turbonilla langae (Saurin, 1959). Reported as: Chemnitzia langae Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Robba et al. (2004), Sawatna et al. (2012), Negri et al. (2014), Gemert (2016).
Turbonilla lerichei (Saurin, 1959). Reported as: Chemnitzia lerichei Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Turbonilla marnayae Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Turbonilla melvilli Dautzenberg, 1912. Reported as: Chrysallida melvilli (Dautzenberg & Fischer, 1906). Gulf of
Thailand. Reported by: Robba et al. (2004).
Turbonilla mumia (A. Adams, 1861). Reported as: Pyrgiscus mumia (Adams, 1861). Gulf of Thailand. Reported
by: Robba et al. (2007).
Turbonilla nodai (Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004). Gulf of Thailand. Reported by:
Robba et al. (2004), Negri et al. (2014).
Turbonilla plana (Robba, Di Geronimo, Chaimanee, Negri & Sanfilippo, 2004). Gulf of Thailand. Reported by:
Robba et al. (2004), Sawatna et al. (2012).
Turbonilla punctiperipherata Nomura, 1936. Reported as: Chemnitzia punctiperipherata (Nomura, 1936). Gulf
of Thailand. Reported by: Robba et al. (2007).
Turbonilla regis (Saurin, 1959). Reported as: Chemnitzia regis Saurin, 1959. Gulf of Thailand. Reported by:
Saurin (1962), Gemert (2016).
Turbonilla rolingiana Saurin, 1962. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Turbonilla romeasiana Saurin, 1962. Reported as: Turbonilla romeasaina Saurin, 1962. Gulf of Thailand.
Reported by: Saurin (1962), Gemert (2016).
Turbonilla sandoi Nomura, 1938. Gulf of Thailand. Reported by: Robba et al. (2004), Chaiwathee et al. (2007).
Turbonilla vinhi Saurin, 1959. Gulf of Thailand. Reported by: Saurin (1962), Gemert (2016).
Superorder Sacoglossa
Superfamily Oxynooidea
Family Cylindrobullidae
Cylindrobulla phuketi Jensen, 1989. Andaman Sea. Reported by: Jensen (1990, 1998, 2007), Thamrongnawasawat
et al. (2008), Aungtonya & Sungjeen (2014).
Family Juliidae
Julia japonica (Kuroda & Habe, 1951). Andaman Sea. Reported by: BEDO (2017e).
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Superfamily Plakobranchoidea
Family Costasiellidae
Costasiella coronata Swennen, 2007. Gulf of Thailand. Reported by: Swennen (2007).
Costasiella kuroshimae Ichikawa, 1993. Gulf of Thailand. Reported by: Mehrotra & Scott (2016), BEDO (2017c).
Costasiella usagi Ichikawa, 1993. Gulf of Thailand. Reported by: Mehrotra & Scott (2016).
Family Hermaeidae
Cyerce bourbonica Yonow, 2012. Andaman Sea. Reported by: Yonow (2012).
Family Limapontiidae
Ercolania halophilae K. R. Jensen, Kohnert, Bendell & Schrödl, 2014. Andaman Sea. Reported by: Jensen et al.
(2014), GBIF (2019a).
Ercolania translucens Jensen, 1993. Andaman Sea. Reported by: Jensen (1998, 2007).
Family Plakobranchidae
Elysia asbecki Wägele, Stemmer, Burghardt & Händeler, 2010. Gulf of Thailand. Reported by: Mehrotra & Scott
(2016), BEDO (2017c).
Elysia bangtawaensis Swennen, 1998. Andaman Sea. Reported by: Jensen (1998, 2007), Swennen (1998), Rudman
(2010), Swennen (2011), BEDO (2017c).
Elysia bennettae Thompson, 1973. Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata
(2009).
Elysia chilkensis Eliot, 1916. Andaman Sea. Reported by: Nabhitabhata (2009).
Elysia hirasei Baba, 1955. Andaman Sea. Reported by: Rudman (2010), BEDO (2017c), Sanpanich & Duangdee
(2018).
Elysia leucolegnote Jensen, 1990. Gulf of Thailand, Andaman Sea. Recorded by: Swennen (2011).
Elysia marginata (Pease, 1871). Andaman Sea. Reported by: Sanpanich & Duangdee (2018).
Elysia ornata (Swainson, 1840). Andaman Sea. Reported by: Jensen (1992, 1996), Nabhitabhata (2009).
Elysia pusilla Bergh, 1872. Also reported as: Elysiella pusilla Bergh, 1872; and Elysia siamensis Swennen, 1997.
Andaman Sea. Reported by: Jensen (1992, 1996, 1998, 2007), Swennen (1997).
Gascoignella jabae Swennen, 2001. Gulf of Thailand. Recorded by: Swennen (2001).
Gascoignella nukuli Swennen, 2001. Gulf of Thailand. Recorded by: Swennen (2001).
Plakobranchus ocellatus van Hasselt, 1824. Also reported as: Placobranchus ocellatus van Hasselt, 1824; and
Plakobranchus ianthobaptus (Gould, 1852). Gulf of Thailand, Andaman Sea. Reported by: Bergh (1902), Jensen
(1992, 1996, 1998, 2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee
(2009), Duangdee et al. (2011), Yonow (2012), BEDO (2017c), Mehrotra & Scott (2017), GBIF (2019a).
Thuridilla albopustulosa Gosliner, 1995. Andaman Sea. Reported by: Duangdee et al. (2013b).
Thuridilla gracilis (Risbec, 1928). Also reported as: Thuridilla ratna (Marcus, 1965). Gulf of Thailand, Andaman
Sea. Reported by: Jensen (1992, 1996, 1998, 2007), Gosliner et al. (2008), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Rudman (2010), Duangdee et al. (2011, 2013b), Hylleberg & Aungtonya (2013), BEDO
(2017c), Mehrotra & Scott (2017), Sanpanich & Duangdee (2018).
Thuridilla moebii (Bergh, 1888). Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata
(2009), Duangdee et al. (2013b).
Thuridilla virgata (Bergh, 1888). Gulf of Thailand. Reported by: BEDO (2017c).
Superorder Siphonarimorpha
Order Siphonariida
Superfamily Siphonarioidea
Family Siphonariidae
Siphonaria atra Quoy & Gaimard, 1833. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Way & Purchon (1981), Kurozumi et al. (1989), Duangdee et al. (2009, 2011, 2013), Hylleberg &
Aungtonya (2013), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a).
Siphonaria guamensis Quoy & Gaimard, 1833. Andaman Sea. Reported by: J. D. Taylor et al. (unpublished data).
Siphonaria japonica (Donovan, 1824). Gulf of Thailand. Reported by: Samakraman et al. (2010), BEDO (2017b).
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The Marine and Estuarine Molluscs of Thailand
Siphonaria javanica (Lamarck, 1819). Andaman Sea. Reported by: BEDO (2017b).
Siphonaria kurracheensis Reeve, 1856. Andaman Sea. Reported by: Nateewathana et al. (1981), Way & Purchon
(1981), Hylleberg & Aungtonya (2013), BEDO (2017b).
Siphonaria laciniosa (Linnaeus, 1758). Andaman Sea. Reported by: Duangdee et al. (2009, 2013b), Samakraman
et al. (2010), Sanpanich & Duangdee (2013), BEDO (2017b).
Siphonaria normalis Gould, 1846. Andaman Sea. Reported by: Nateewathana et al. (1981), Sørensen (2012),
BEDO (2017b), GBIF (2019a).
Siphonaria sirius Pilsbry, 1894. Also reported as: Anthosiphonaria sirius (Pilsbry, 1894). Andaman Sea. Reported
by: Nateewathana et al. (1981), Hylleberg & Aungtonya (2013), BEDO (2017b).
Siphonaria subatra Pilsby, 1904. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2013a,
2013b).
Order Umbraculida
Superfamily Umbraculoidea
Family Umbraculidae
Umbraculum umbraculum Lightfoot, 1786. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009).
Subclass Neritimorpha
Order Cycloneritida
Superfamily Neritoidea
Family Neritidae
Clithon avellana (Récluz, 1842). Reported as: Nerita avellana Récluz, 1842. Gulf of Thailand. Reported by: Morlet
(1889).
Clithon corona (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Fukumori & Kano (2014), Eichhorst
(2016), BEDO (2017b).
Clithon coronatum (Leach, 1815). Also reported as: Theodoxus coronatus. Andaman Sea. Reported by: THNHM,
KUMF, Nabhitabhata (2009), BEDO (2017b).
Clithon faba (G. B. Sowerby I, 1836). Reported as: Clithon (Clithon) faba (G. B. Sowerby I, 1836). Gulf of Thailand,
Andaman Sea. Reported by: Gemert (2003), Sri-aroon et al. (2005, 2006), Eichhorst (2016), GBIF (2019b).
Clithon michaudi (Récluz, 1841). No Thai locality provided. Reported by: Galli (2015).
Clithon oualaniense (Lesson, 1831). Also reported as: Neritina oualaniense Lesson, 1831; and Theodoxus
oualaniansis. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Nateewathana et al. (1981), Swennen
et al. (2001), Gemert (2003), Sri-aroon et al. (2005), Meyer et al. (2008), Thamrongnawasawat et al. (2008),
Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013), Fukumori & Kano (2014),
Eichhorst (2016), BEDO (2017b), GBIF (2019b), J. D. Taylor et al. (unpublished data).
Clithon peguense (Blanford, 1867). Andaman Sea. Reported by: Nateewathana et al. (1981), Bussarawit (1995),
Nabhitabhata (2009), Dechruksa et al. (2014), BEDO (2017b).
Clithon pulchellum (Récluz, 1843). No Thai locality given. Reported by: Eichhorst (2016).
Clithon rarispina (Mousson, 1849). Andaman Sea. Reported by: BEDO (2017b).
Clithon retropictum (Martens, 1879). Reported by: KUMF, Nabhitabhata (2009).
Clithon sowerbianum (Récluz, 1843). Reported as: Clithon (Clithon) sowerbyana (Récluz, 1843). Andaman Sea.
Reported by: THNHM, Nateewathana et al. (1981), Duangdee et al. (2009, 2011, 2013a), BEDO (2017b), GBIF
(2019a).
Neripteron bensoni (Récluz, 1850). Reported as: Neritina guerini (Récluz, 1841). Gulf of Thailand, Andaman Sea.
Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Dekker (2000), Nabhitabhata (2009), BEDO
(2017b).
Neripteron cornucopia (Benson, 1836). Also reported as: Neritina cornucopia Benson, 1836. Gulf of Thailand.
Reported by: Morlet (1889), Eichhorst (2016), BEDO (2017b).
Neripteron siquijorense (Récluz, 1844). Reported as: Puperia siquijorensis (Récluz, 1841) and Neritina
siquijorensis (Récluz, 1843). Andaman Sea. Reported by: BEDO (2017b).
Neripteron violaceum (Gmelin, 1791). Reported as: Neripteron (Dostia) violaceum (Gmelin, 1791), Neritina
violacea (Gmelin, 1791) and Dostia violacea (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by:
THNHM, Morlet (1889), Tantanasiriwong (1978), Nateewathana et al. (1981), Kasetsart University (2000),
Subba Rao & Dey (2000), Gemert (2003), Duangdee (2004, 2007), Robba et al. (2004), Sri-aroon et al. (2005,
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
2006), Printrakoon et al. (2008), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), Dechruksa et al. (2014), Fukumori & Kano (2014), Eichhorst (2016),
BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Nerita albicilla (Linnaeus, 1758). Also reported as: Nerita (Theliostyla) albicilla (Linnaeus, 1758). Gulf of
Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti (1937, 1938), Andersen (1977),
Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Nateewathana (1995), Kurozumi
et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Subba Rao &
Dey (2000), Swennen et al. (2001), Gemert (2003), Duangdee (2004, 2007), Wells (2005), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011), ONEP (2007), Printrakoon et al. (2008), Thamrongnawasawat et al. (2008),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), Sørensen (2012), Hylleberg & Aungtonya
(2013), Kostina et al. (2016), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Nerita antiquata Récluz, 1841. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Nerita balteata Reeve, 1855. Also reported as: Nerita (Cymostyla) balteata Reeve, 1855; Nerita articulata Gould,
1847; Nerita birmanica (Philippi & Chemnitz, 1848); and Nerita lineata. Gulf of Thailand, Andaman Sea. Reported
by: THNHM, KUMF, Morlet (1889), Andersen (1977), Tantanasiriwong (1978), Nateewathana et al. (1981),
Amornjaruchit (1988), Nateewathana (1995), Kasetsart University (2000), ONEP (2007), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013), Fukumori & Kano (2014), Eichhorst (2016), Kostina
et al. (2016), BEDO (2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Nerita chamaeleon Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti
(1937, 1938), Andersen (1977), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Carpenter & Niem (1998), Subba Rao & Dey (2000), Swennen et al. (2001), Duangdee
(2004, 2007), Kasetsart University (2006), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013b),
ONEP (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013),
Sørensen (2012), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
Nerita costata (Gmelin, 1791). Also reported as: Nerita (Ritena) costata (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Nateewathana (1995), Carpenter & Niem (1998), Dekker (2000), Kasetsart University
(2000, 2006), Duangdee (2004), Wells (2005), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011, 2013b),
ONEP (2007), Thamrongnawasawat et al. (2008), Sørensen (2012), Hylleberg & Aungtonya (2013), Sanpanich
& Duangdee (2013), Eichhorst (2016), BEDO (2017b), Sanpanich & Duangdee (2018), GBIF (2019a), J. D. Taylor
et al. (unpublished data).
Nerita histrio Linnaeus, 1758. Also reported as: Nerita squamulata Le Guillou, 1841; and Nerita (Theliostyla)
squamulata Le Guillou, 1841. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Andersen (1977),
Tantanasiriwong (1978), Kurozumi et al. (1989), Bussarawit (1995), Carpenter & Niem (1998), Subba Rao &
Dey (2000), Gemert (2003), Duangdee (2004), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Nabhitabhata
(2009), Sørensen (2012), Sanpanich & Duangdee (2013), BEDO (2017b).
Nerita insculpta (Récluz, 1841). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Andersen (1977),
Tantanasiriwong (1978), Nateewathana et al. (1981), Amornjaruchit (1988), ONEP (2007), Nabhitabhata (2009),
BEDO (2017b).
Nerita litterata Gmelin, 1791. Also reported as: Nerita (Amphinerita) rumphii (Récluz). Gulf of Thailand, Andaman
Sea. Reported by: Kurozumi et al. (1989), Sørensen (2012).
Nerita maura Récluz, 1842. Andaman Sea. Reported by: Galli (2015), BEDO (2017b).
Nerita maxima (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017b).
Nerita nigrita (Röding, 1798). Also reported as: Nerita (Cymostyla) nigrita (Röding, 1798). Andaman Sea.
Reported by: Andersen (1977), Tantanasiriwong (1978), Nateewathana et al. (1981), Nabhitabhata (2009),
Eichhorst (2016), BEDO (2017b).
Nerita oryzarum (Récluz, 1844). Andaman Sea. Reported by: Suvatti (1938), Kilburn & Hylleberg (1998),
Nabhitabhata (2009), BEDO (2017b).
Nerita planospira Anton, 1839. Gulf of Thailand, Andaman Sea. Reported by: Andersen (1977), Nateewathana
et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Dekker (2000), Kasetsart University (2000),
Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2013), BEDO (2017b), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Nerita plicata Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Andersen (1977), Tantanasiriwong
(1978), Nateewathana et al. (1981), Wells (2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009),
Sørensen (2012), Duangdee et al. (2013b), BEDO (2017b).
Nerita polita Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1937, 1938),
Andersen (1977), Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit
(1995), Dekker (2000), Kasetsart University (2000, 2006), Subba Rao & Dey (2000), Duangdee (2004), Wells
(2005), ONR & EPP (2006), Sri-aroon et al. (2006), ONEP (2007), Thamrongnawasawat et al. (2008), Duangdee
et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Sørensen (2012), Galli (2015),
BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al. (unpublished data).
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The Marine and Estuarine Molluscs of Thailand
Nerita undata (Linnaeus, 1758). Also reported as: Nerita (Cymostyla) undata (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1937, 1938), Gemert (2003), Duangdee (2004), Wells
(2005), ONR & EPP (2006), ONEP (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013),
Duangdee et al. (2011), Sørensen (2012), GBIF (2019a).
Nerita undulata Gmelin, 1791. Also reported as: Nerita grayana Récluz, 1844. Andaman Sea. Reported by:
Nateewathana et al. (1981), Sørensen (2012). Nerita grayana Récluz, 1844, was synonymised with Nerita
undulata Gmelin, 1791, by Krijnen et al. (2011), but incorrectly listed as a synonym of Nerita essingtoni Récluz,
1842, based on Eichhorst (2016) by WoRMS Editorial Board (2019).
Neritina communis (Quoy & Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019b). This
species cannot be confirmed through WoRMS Editorial Board (2019), but is listed on Galli (2015).
Neritina pulligera (Linnaeus, 1767). Andaman Sea. Reported by: KUMF, Gemert (2003), Sri-aroon et al. (2005),
Nabhitabhata (2009), BEDO (2017b).
Neritina squamaepicta (Récluz, 1843). Reported as: Neritina squamipicta (Récluz, 1843). No Thai locality
provided. Reported by: Galli (2015).
Neritina turrita (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009).
Neritodryas dubia (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Sri-aroon et al. (2005), Printrakoon
et al. (2008), BEDO (2017b).
Septaria lineata (Lamarck, 1816). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017b).
Septaria tesselata (Lamarck, 1816). Reported as: Septaria tesselaria (Lamarck, 1816). Andaman Sea. Reported
by: Eichhorst (2016).
Smaragdia souverbiana (Montrouzier in Souverbie & Montrouzier, 1863). Gulf of Thailand. Reported by:
Gemert (2003).
Family Neritiliidae
Neritilia rubida (Pease, 1865). No Thai locality provided. Reported by: Higo et al. (1999), Kano & Kase (2003).
Family Phenacolepadidae
Plesiothyreus newtoni G. B. Sowerby III, 1894. Reported as: Phenacolepas newtoni (Sowerby, 1894). Gulf of
Thailand. Reported by: Robba et al. (2007).
Plesiothyreus osculans (C. B. Adams, 1852). Reported as: Phenacolepas osculans (C. B. Adams, 1852). Gulf of
Thailand. Reported by: Gemert (2003). Phenacolepas is a junior synonym of Plesiothyreus (see Fukumori et al.,
2019).
Zacalantica galathea (Lamarck, 1819). Reported as: Phenacolepas galathea (Lamarck, 1819). Gulf of Thailand.
Reported by: Gemert (2003).
Zacalantica scobinata (Gould, 1859). Reported as: Phenacolepas scobinata (Gould, 1859). Gulf of Thailand.
Reported by: Gemert (2003).
Zacalantica cf. unguiformis (Gould, 1859). Reported as: Phenacolepas cf. unguiformis (Gould, 1859). Gulf of
Thailand. Reported by: Gemert (2003).
Superfamily Neritopsoidea
Family Neritopsidae
Neritopsis radula (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), BEDO (2017b).
Subclass Patellogastropoda
Superfamily Lottioidea
Family Lottiidae
Lottia dorsuosa (Gould, 1859). Reported as: Cellana dorsuosa. No Thai locality given. Reported by: THHNM.
Lottia tenuisculptata Sasaki & Okutani, 1994. Gulf of Thailand. Reported by: GBIF (2019b).
Nipponacmea nigrans (Kira, 1961). Gulf of Thailand. Reported by: Sanpanich & Duangdee (2009, 2013), Duangdee
et al. (2011).
Notoacmea flammea (Quoy & Gaimard, 1834). Andaman Sea. Reported by: Sørensen (2012).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Patelloida mufria (Hedley, 1915). Andaman Sea. Reported by: Sørensen (2012).
Patelloida pygmaea (Dunker, 1860). Also reported as: Patelloida (Asteracmea) pygmaea (Dunker, 1860). Gulf of
Thailand, Andaman Sea. Reported by: Kurozumi et al. (1989), Gemert (2003), Sanpanich & Duangdee (2009,
2013), Duangdee et al. (2011), BEDO (2017b).
Patelloida saccharina (Linnaeus, 1758). Also reported as: Patelloida saccharina lanx. Gulf of Thailand, Andaman
Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Swennen et al. (2001), ONR
& EPP (2006), Duangdee (2007), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013), Samakraman et al. (2010), Sørensen (2012), Kostina et al. (2016), BEDO (2017b), J. D. Taylor
et al. (unpublished data). GBIF (2019a) records the subspecies Patelloida saccharina stella (Lesson, 1831) from
Thailand.
Patelloida striata (Quoy & Gaimard, 1834). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem
(1998), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Family Nacellidae
Cellana enneagona (Reeve, 1854). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), BEDO (2017b),
GBIF (2019b).
Cellana orientalis (Pilsbry, 1891). Also reported as: Cellana radiata orientalis (Pilsbry, 1891). Andaman Sea.
Reported by: Nateewathana et al. (1981), BEDO (2017b).
Cellana radiata (Born, 1778). Also reported as: Cellana radiata radiata (Born, 1778); Cellana luzonica (Reeve,
1855); and Acmaea bombayana Smith, 1911. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al.
(1981), Way & Purchon (1981), Bussarawit (1995), Kilburn & Hylleberg (1998), Gemert (2003), Nabhitabhata
(2009), Duangdee et al. (2011), Sørensen (2012), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a),
J. D. Taylor et al. (unpublished data).
Cellana rota (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), Kilburn &
Hylleberg (1998), Wells (2005), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2013), GBIF (2019b).
Cellana testudinaria (Linnaeus, 1758). Also reported as: Patella testudinaria (Linnaeus, 1758). Gulf of Thailand,
Andaman Sea. Reported by: THNHM, KUMF, Powell (1973), Nateewathana et al. (1981), Bussarawit (1995),
Carpenter & Niem (1998), Duangdee (2004, 2007), Wells (2005), Kasetsart University (2006), ONR & EPP
(2006), Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018),
Kostina et al. (2016), BEDO (2017b), GBIF (2019a).
Cellana toreuma (Reeve, 1854). Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981),
Kurozumi et al. (1989), Samakraman et al. (2010), Duangdee et al. (2011), Sanpanich & Duangdee (2013),
Kostina et al. (2016), BEDO (2017b).
Superfamily Patelloidea
Family Patellidae
Scutellastra flexuosa (Quoy & Gaimard, 1834). Reported as: Patella flexuosa (Quoy & Gaimard, 1834). Gulf of
Thailand, Andaman Sea. Reported by: Way & Purchon (1981), Bussarawit (1995), Kasetsart University (2000),
Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), BEDO (2017b), GBIF (2019a).
Subclass Vetigastropoda
Order Lepetellida
Superfamily Fissurelloidea
Family Fissurellidae
Cymbula sanguinans (Reeve, 1854). Reported as: Patella sanguinans (Reeve, 1854). Gulf of Thailand. Reported
by: Morlet (1889).
Diodora funiculata (Reeve, 1850). Andaman Sea. Reported by: BEDO (2017b).
Diodora galeata (Helbling, 1779). Gulf of Thailand. Reported by: Gemert (2003).
Diodora cf. jukesii (Reeve, 1850). Gulf of Thailand. Reported by: Gemert (2003).
Diodora mus (Reeve, 1850). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981),
Nabhitabhata (2009), Duangdee et al. (2013a), BEDO (2017b).
Diodora octagona (Reeve, 1850). Also reported as: Diodora cf. reevei Schepman, 1908. Gulf of Thailand. Reported
by: Gemert (2003), GBIF (2019b).
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The Marine and Estuarine Molluscs of Thailand
Diodora quadriradiata (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Robba et al. (2004), Nabhitabhata (2009), BEDO (2017b), GBIF (2019a).
Diodora ruppellii (G. B. Sowerby I, 1835). Gulf of Thailand, Andaman Sea. Reported by: Gemert (2003), Duangdee
et al. (2011), Sanpanich & Duangdee (2013).
Diodora sieboldii (Reeve, 1850). Andaman Sea. Reported by: BEDO (2017b).
Diodora singaporensis (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University (2006),
ONEP (2007).
Diodora subcalyculata (Schepman, 1908). Andaman Sea. Reported by: BEDO (2017b).
Diodora ticaonica (Reeve, 1850). Also reported as: Diodora tricaonica (Reeve, 1850). Gulf of Thailand. Reported
by: Gemert (2003), Duangdee (2004), Duangdee et al. (2007, 2011), Nabhitabhata (2009), Sanpanich & Duangdee
(2009, 2013).
Diodora venusta (Reeve, 1850). Reported as: Fissurella venusta (Reeve, 1850). Gulf of Thailand. Reported by: Morlet
(1889). This species cannot be confirmed through WoRMS Editorial Board (2019), but is listed on OBIS (2020).
Emarginula crassicostata (G. B. Sowerby II, 1863). Andaman Sea. Reported by: Gemert (2003), Kasetsart University
(2006), ONR & EPP (2006), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Emarginella eximia (A. Adams, 1852). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Emarginula foveolata Schepman, 1908. Andaman Sea. Reported by: BEDO (2017b).
Emarginula puncticulata A. Adams, 1852. Andaman Sea. Reported by: BEDO (2017b).
Emarginula retecosa A. Adams, 1851. Andaman Sea. Reported by: BEDO (2017b).
Emarginula sicula J. E. Gray, 1825. Andaman Sea. Reported by: BEDO (2017b).
Emarginula undulata Melvill & Standen, 1903. Gulf of Thailand. Reported by: Robba et al. (2004).
Fissurella cyathulum (Reeve, 1850). Reported as: Diodora cyathulum (Reeve, 1850). Andaman Sea. Reported by:
BEDO (2017b).
Hemitoma octoradiata (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017b).
Macroschisma sinense A. Adams, 1855. Gulf of Thailand. Reported by: GBIF (2019b).
Montfortista panhi (Quoy & Gaimard, 1834). Reported as: Hemitoma panhi (Quoy & Gaimard, 1834). Gulf of
Thailand, Andaman Sea. Reported by: Kilburn & Hylleberg (1998), Nabhitabhata (2009), BEDO (2017b).
Scutus emarginatus (Philippi, 1825). Gulf of Thailand. Reported by: Robba et al. (2004).
Scutus sinensis (Blainville, 1825). Also reported as: Scutus (Ariscutus) sinensis. Gulf of Thailand, Andaman Sea.
Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Swennen et al. (2001), Duangdee
(2007), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Scutus unguis (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee et al.
(2007, 2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Superfamily Haliotoidea
Family Haliotidae
Haliotis asinina Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Chaitiamwong et al.
(1971), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Jarayabhand & Paphavasit
(1996), Carpenter & Niem (1998), Jarayabhand et al. (1998), Geiger & Poppe (2000), Kasetsart University (2006),
ONR & EPP (2006), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Geiger & Owen (2012), Hylleberg
& Aungtonya (2013), Sanpanich & Duangdee (2013), Kostina et al. (2016), BEDO (2017b), GBIF (2019a).
Haliotis clathrata Reeve, 1846. Also reported as: Haliotis thailandis Dekker & Patamakanthin, 2001; Haliotis
tomricei Patamakanthin, 2002; and Haliotis clathrata thailandis (Patamakanthin, 2002). Gulf of Thailand,
Andaman Sea. Reported by: Patamakanthin (2001, 2002), Gemert (2003), Thamrongnawasawat et al. (2008),
Geiger & Owen (2012), Duangdee et al. (2013b), Aungtonya & Sungjeen (2014), Galli (2015), BEDO (2017b).
Haliotis diversicolor Reeve, 1846. Andaman Sea. Reported by: GBIF (2019a).
Haliotis glabra Gmelin, 1791. Andaman Sea. Reported by: Carpenter & Niem (1998), Geiger & Poppe (2000),
Nabhitabhata (2009).
Haliotis jacnensis Reeve, 1846. Gulf of Thailand. Reported by: Geiger & Poppe (2000).
Haliotis ovina (Gmelin, 1791). Also reported as: Haliotis (Ovinotis) ovina (Gmelin, 1791); Ovinotis ovina (Gmelin);
Haliotis patamakanthini Dekker, Regter & Gras, 2001; Haliotis ovina patamakanthini Dekker, Regter & Gras,
2001; and Ovinotis ovina. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Chaitiamwong et al.
(1971), Tantanasiriwong (1978), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995),
Jarayabhand & Paphavasit (1996), Carpenter & Niem (1998), Jarayabhand et al. (1998), Geiger & Poppe (2000),
Gemert (2003), Duangdee (2004), Wells (2005), Kasetsart University (2006), ONEP (2007), Thamrongnawasawat
et al. (2008), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009,
2013, 2018), Geiger & Owen (2012), Hylleberg & Aungtonya (2013), Galli (2015), Kostina et al. (2016), BEDO
(2017b), GBIF (2019a, 2019b).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Haliotis planata G. B. Sowerby II, 1882. Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), ONR & EPP (2006), Thamrongnawasawat
et al. (2008), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Geiger & Owen (2012), Sanpanich & Duangdee
(2013), BEDO (2017b).
Haliotis varia Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: Nateewathana et al. (1981), Way &
Purchon (1981), Jarayabhand & Paphavasit (1996), Jarayabhand et al. (1998), Thamrongnawasawat et al. (2008),
Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Geiger & Owen
(2012), Hylleberg & Aungtonya (2013), BEDO (2017b), GBIF (2019a).
Superfamily Scissurelloidea
Family Scissurellidae
Scissurella evaensis Bandel, 1998. Andaman Sea. Reported by: Geiger & Jansen (2004).
Scissurella mirifica (A. Adams, 1862). Reported as: Scissurella declinans Watson, 1886. Andaman Sea. Reported
by: Geiger & Jansen (2004).
Scissurella spinosa Geiger & Jansen, 2004. Andaman Sea. Reported by: Geiger & Jansen (2004).
Sinezona plicata (Hedley, 1899). Andaman Sea. Reported by: Geiger & Jansen (2004).
Order Pleurotomariida
Superfamily Pleurotomarioidea
Family Pleurotomariidae
Bayerotrochus indicus (Anseeuw, 1999). Andaman Sea. Reported by: Galli (2015).
Order Seguenziida
Superfamily Seguenzioidea
Family Chilodontaidae
Euchelus asper (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Tantanasiriwong (1978),
Nateewathana et al. (1981), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO
(2017b), GBIF (2019a).
Euchelus atratus (Gmelin, 1791). Reported as: Herpetopoma atratum (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: Sanpanich & Duangdee (2013), BEDO (2017b), GBIF (2019a, 2019b), J. D. Taylor et al.
(unpublished data).
Euchelus horridus (Philippi, 1846). Gulf of Thailand. Reported by: Gemert (2003).
Euchelus punctigerus (A. Adams, 1853). Gulf of Thailand. Reported by: Gemert (2003). This species cannot be
confirmed through WoRMS Editorial Board (2019), but is listed on GBIF (2020).
Euchelus scaber (Linnaeus, 1758). Reported as: Trochus scaber Linnaeus, 1758. Gulf of Thailand. Reported by:
Morlet (1889).
Herpetopoma exasperatum (A. Adams, 1853). Andaman Sea. Reported by: BEDO (2017e).
Herpetopoma instrictum (Gould, 1849). Reported as: Euchelus aff. instrictus (Gould, 1849). Gulf of Thailand.
Reported by: Gemert (2003).
Hybochelus cancellatus (Krauss, 1848). Also reported as: Hybochelus cancellata (Pilsbry). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Gemert (2003), Robba et al.
(2004), Wells (2005), Nabhitabhata (2009), Duangdee et al. (2011, 2013b), Sanpanich & Duangdee (2013),
Negri et al. (2014), BEDO (2017b).
Family Eucyclidae
Turcica maculata (Brazier, 1877). Gulf of Thailand. Reported by: Nabhitabhata (2009).
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The Marine and Estuarine Molluscs of Thailand
Order Trochida
Superfamily Trochoidea
Family Angariidae
Angaria delphinus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Kurozumi et al. (1989), Bussarawit (1995), Gemert (2003), Duangdee
(2004, 2007), Kasetsart University (2006), ONR & EPP (2006), ONEP (2007), Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2013, 2018), Duangdee et al. (2011, 2013a), Hylleberg & Aungtonya (2013), Kostina et al.
(2016), BEDO (2017b), GBIF (2019a, 2019b).
Angaria lilianae Monsecour & Monsecour, 2000. Andaman Sea. Reported by: Monsecour & Monsecour, 2000.
Family Calliostomatidae
Calliostoma iris Kuroda & Habe, 1961. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Calliostoma tranquebaricum (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Swennen et al.
(2001), Taylor (2009), Vilvens (2009).
Calliostoma virescens Coen, 1933. Reported as: Calliostoma tornatum (Coen, 1933). Gulf of Thailand. Reported
by: BEDO (2017b).
Tristichotrochus haliarchus (Melvill, 1889). Reported as: Calliostoma (Benthastelena) haliarchus (Melvill, 1889).
No Thai locality provided. Reported by: Vilvens (2009).
Family Colloniidae
Bothropoma pilula (Dunker, 1860). Reported as: Neocollonia pilula (Dunker, 1860). Gulf of Thailand. Reported
by: Gemert (2003).
Family Conradiidae
Conradia cingulifera A. Adams, 1860. Gulf of Thailand. Reported by: Gemert (2003).
Family Liotiidae
Bathyliotina schepmani (Habe, 1953). Andaman Sea. Reported by: BEDO (2017b).
Cyclostrema solariellum Melvill, 1893. Gulf of Thailand. Reported by: Gemert (2003).
Cyclostrema quadricarinatum Melvill & Standen, 1901. Reported as: Circulus quadricarinatus (Melvill & Standen,
1901). Gulf of Thailand. Reported by: Robba et al. (2004).
Dentarene sarcina Iredale, 1929. Reported as: Pentarene sarcina Iredale, 1929. Gulf of Thailand. Reported by:
Gemert (2003).
Liotinaria peronii (Kiener, 1839). Reported as: Liotina peronii (Kiener, 1839). Andaman Sea. Reported by: BEDO
(2017e).
Pseudoliotina discoidea (Reeve, 1843). Gulf of Thailand. Reported by: Gemert (2003).
Family Phasianellidae
Hiloa variabilis (Pease, 1861). Reported as: Tricolia variabilis (Pease, 1861). Gulf of Thailand. Reported by:
Gemert (2003).
Tricolia fordiana (Pilsbry, 1888). Andaman Sea. Reported by: Robertson (1985).
Family Skeneidae
Dillwynella aff. planorbis Hasegawa, 1997. Gulf of Thailand. Reported by: Gemert (2003).
Leucorhynchia caledonica Crosse, 1867. Andaman Sea. Reported by: BEDO (2017e).
Leucorhynchia crossei Tryon, 1888. Andaman Sea. Reported by: BEDO (2017e).
Family Solariellidae
Minolia punctata A. Adams, 1860. Gulf of Thailand. Reported by: Gemert (2003).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Family Tegulidae
Rochia conus (Gmelin, 1791). Reported as: Trochus conus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported
by: Carpenter & Niem (1998), Nabhitabhata (2009).
Rochia maxima (Koch, 1844). Andaman Sea. Reported by: BEDO (2017b).
Rochia nilotica (Linnaeus, 1767). Also reported as: Trochus niloticus Linnaeus, 1758; and Tectus niloticus Linnaeus,
1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti (1938), Tantanasiriwong (1978),
Nateewathana et al. (1981), Way & Purchon (1981), Amornjaruchit (1988), Kurozumi et al. (1989), Bussarawit
(1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Duangdee (2004, 2007), Wells (2005),
ONR & EPP (2006), ONEP (2007), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013b),
Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Tectus fenestratus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009), GBIF (2019a).
Tectus virgatus (Gmelin, 1791). Andaman Sea. Reported by: GBIF (2019a).
Tectus pyramis (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong
(1978), Nateewathana et al. (1981), Way & Purchon (1981), Bussarawit (1995), Carpenter & Niem (1998),
Duangdee (2004), Wells (2005), Kasetsart University (2006), ONR & EPP (2006), Thamrongnawasawat et al.
(2008), Duangdee et al. (2009, 2013a, 2013b), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013,
2018), Hylleberg & Aungtonya (2013), Kostina et al. (2016), BEDO (2017b), GBIF (2019b).
Family Trochidae
Calliotrochus marmoreus (Pease, 1861). Gulf of Thailand. Reported by: Gemert (2003).
Chrysostoma paradoxum (Born, 1778). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti
(1938), Kurozumi et al. (1989), Gemert (2003), Duangdee (2004), Wells (2005), Kasetsart University (2006),
ONR & EPP (2006), ONEP (2007), Duangdee et al. (2009, 2011, 2013b), Sanpanich & Duangdee (2009, 2013,
2018). Reported as Trochus nicobaricus Gmelin, 1791, by Morlet (1889). Gulf of Thailand. There is no Trochus
nicobaricus Gmelin, 1791, listed in WoRMS Editorial Board (2019), but there is a Turbo nicobaricus Gmelin,
1791 (a synonym of Chrysostoma paradoxum [Born, 1778]), which is likely to be the species reported by Morlet
(1889), GBIF (2019a).
Clanculus bicarinatus Angas, 1880. Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Clanculus bronni Dunker, 1860. Gulf of Thailand. Reported by: Gemert (2003).
Clanculus clanguloides (Wood, 1828). Andaman Sea. Reported by: Wells (2005), Sanpanich & Duangdee (2018).
Clanculus denticulatus (Gray, 1827). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al.
(1981), Bussarawit (1995), Nabhitabhata (2009), Duangdee et al. (2011), Hylleberg & Aungtonya (2013),
Sanpanich & Duangdee (2013), Kostina et al. (2016), BEDO (2017b), GBIF (2019a). The status of this species is
uncertain. There are numerous references to it occurring in Thailand by various authors, and GBIF (2020)
shows the species to be widespread in the Indo-West Pacific, but Wilson (1993) shows the range as southern
Australia.
Clanculus edentulus A. Adams, 1853. Gulf of Thailand. Reported by: Kostina et al. (2016).
Clanculus margaritarius (Philippi, 1846). Reported as: Clanculus unedo Adams, 1853. Gulf of Thailand, Andaman
Sea. Reported by: Gemert (2003), Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011), BEDO (2017b).
Clanculus microdon (A. Adams, 1853). Andaman Sea. Reported by: Suvatti (1938), Nabhitabhata (2009).
Clanculus philippii (Koch in Philippi, 1843). Andaman Sea. Reported by: Sanpanich & Duangdee (2013).
Clanculus samoensis (Hombron & Jacquinot, 1848). Also reported as: Clanculus atropurpureus (Gould, 1849).
Gulf of Thailand, Andaman Sea. Reported by: THNHM, Way & Purchon (1981), Gemert (2003), Duangdee et al.
(2011), Sanpanich & Duangdee (2013), Kostina et al. (2016).
Conotalopia musiva (Gould, 1861). Reported as: Pseudominolia musiva (Gould, 1861). Gulf of Thailand. Reported
by: Gemert (2003).
Isanda coronata A. Adams, 1854. Andaman Sea. Reported by: BEDO (2017b).
Jujubinus gilberti (Montrouzier, 1878). Reported as: Cantharidus gilberti (Montrouzier, 1878). Gulf of Thailand.
Reported by: Gemert (2003).
Jujubinus cf. polychromus (A. Adams, 1853). Andaman Sea. Reported by: BEDO (2017e).
Monilea belcheri (Philippi, 1849). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Swennen et al. (2001), Gemert (2003), Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Sanpanich & Duangdee (2013), BEDO (2017b).
Monilea callifera (Lamarck, 1822). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Swennen
et al. (2001), Wells (2005), Duangdee et al. (2007, 2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a, 2019b).
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The Marine and Estuarine Molluscs of Thailand
Monilea smithi (Wood, 1828). Andaman Sea. Reported by: Tantanasiriwong (1978), Bussarawit (1995), Duangdee
et al. (2007), Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Monodonta canalifera Lamarck, 1816. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF,
Tantanasiriwong (1978), Nateewathana et al. (1981), Duangdee (2004), Kasetsart University (2006), ONEP
(2007), Duangdee et al. (2011), Sanpanich & Duangdee (2013), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Monodonta labio (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti
(1937, 1938), Nateewathana et al. (1981), Way & Purchon (1981), Takenouchi (1985), Kurozumi et al. (1989),
Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006), Gemert (2003), Duangdee
(2004, 2007), ONR & EPP (2006), Duangdee et al. (2007, 2009, 2011), ONEP (2007), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013), Kostina et al. (2016), BEDO (2017b), GBIF
(2019a, 2019b), J. D. Taylor et al. (unpublished data).
Pagodatrochus variabilis (H. Adams, 1873). Andaman Sea. Reported as: Minolia variabilis A. Adams. Reported
by: Nateewathana et al. (1981).
Pseudostomatella decolorata (Gould, 1848). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Rossiteria nucleus (Philippi, 1849). Reported as: Monilea nucleus (Philippi, 1849). Gulf of Thailand. Reported by:
Gemert (2003).
Rubritrochus pulcherrimus (A. Adams, 1855). Reported as: Gibbula pulcherrima (Adams, 1855). Andaman Sea.
Reported by: Nateewathana et al. (1981), Bussarawit (1995), Nabhitabhata (2009).
Stomatella auricula (Lamarck, 1816). Also reported as: Stomatella varia (A. Adams, 1850). Gulf of Thailand,
Andaman Sea. Reported by: Nateewathana et al. (1981), Way & Purchon (1981), Kurozumi et al. (1989),
Bussarawit (1995), Gemert (2003), Nabhitabhata (2009), BEDO (2017b), BEDO (2017e).
Stomatella impertusa (Burrows, 1815). Also reported as: Stomatella (Gena) impertusa (Burrow, 1815). Gulf of
Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009, 2013), Duangdee et al. (2011, 2013b),
BEDO (2017e).
Stomatia phymotis Helbling, 1779. Reported as: Stomatia imperforata Gmelin, 1791. Gulf of Thailand. Reported
by: Gemert (2003).
Stomatolina crenulata (Preston, 1908). Reported as: Stomatolina crenulata (Preston, 1908). Gulf of Thailand.
Reported by: Gemert (2003).
Synaptocochlea pulchella (A. Adams, 1850). Andaman Sea. Reported by: BEDO (2017e).
Trochus chloromphalus (A. Adams, 1853). Andaman Sea. Reported by: Bussarawit (1995).
Trochus histrio (Reeve, 1842). Also reported as: Thalotia histrio (Reeve, 1842). Gulf of Thailand, Andaman Sea.
Reported by: Wells (2005), Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Trochus maculatus Linnaeus, 1758. Also reported as: Trochus (Trochus) maculatus Linnaeus, 1758. Gulf of
Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889), Suvatti (1937, 1938), Tantanasiriwong
(1978), Nateewathana et al. (1981), Way & Purchon (1981), Kurozumi et al. (1989), Bussarawit (1995), Carpenter
& Niem (1998), Kasetsart University (2000, 2006), Gemert (2003), Duangdee (2004, 2007), ONR & EPP (2006),
Duangdee et al. (2007, 2009, 2011, 2013a, 2013b), ONEP (2007), Thamrongnawasawat et al. (2008), Sanpanich
& Duangdee (2009, 2013, 2018), Sørensen (2012), Kostina et al. (2016).
Trochus nigropunctatus Reeve, 1861. Also reported as: Trochus hanleyanus (Reeve, 1842). Gulf of Thailand,
Andaman Sea. Reported by: KUMF, Carpenter & Niem (1998), Gemert (2003), Wells (2005), Nabhitabhata (2009),
Duangdee et al. (2011), Sanpanich & Duangdee (2013).
Trochus radiatus Gmelin, 1791. Gulf of Thailand, Andaman Sea. Reported by: KUMF, Morlet (1889), Suvatti
(1938), Gemert (2003), ONR & EPP (2006), Duangdee et al. (2009, 2011), Nabhitabhata (2009), Sanpanich &
Duangdee (2013), BEDO (2017b), GBIF (2019a).
Trochus rota (Dunker, 1860). Reported as: Trochus sacellum rota (Dunker, 1862). Andaman Sea. Reported by:
Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), BEDO (2017b).
Trochus stellatus Gmelin, 1791. Andaman Sea. Reported by: KUMF, Duangdee et al. (2009, 2011), Nabhitabhata
(2009), Sanpanich & Duangdee (2013).
Umbonium elegans (Kiener, 1838). Gulf of Thailand. Reported by: Morlet (1899).
Umbonium vestiarium (Linnaeus, 1758). Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Suvatti
(1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Carpenter & Niem (1998), Kasetsart University
(2000), Swennen et al. (2001), Gemert (2003), Duangdee (2004), Robba et al. (2004), ONR & EPP (2006),
Duangdee et al. (2007, 2009), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Sanpanich & Duangdee
(2013), GBIF (2019a).
Family Turbinidae
Astralium calcar (Linnaeus, 1758). Also reported as: Astraea calcar (L., 1758). Gulf of Thailand, Andaman Sea.
Reported by: KUMF, Way & Purchon (1981), Nabhitabhata (2009).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Astralium haematragum (Menke, 1829). Gulf of Thailand, Andaman Sea. Reported by: Bussarawit (1995), Wells
(2005), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013b), Nabhitabhata (2009), Sanpanich
& Duangdee (2013).
Astralium rhodostomum (Lamarck, 1822). Also reported as: Astraea tuberosa (Philippi, 1846). Gulf of Thailand,
Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Kilburn
& Hylleberg (1998), Gemert (2003), Wells (2005), Duangdee et al. (2009, 2011, 2013a, 2013b), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), BEDO (2017b), GBIF (2019a).
Astralium semicostatum (Kiener, 1850). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Duangdee (2004),
Kasetsart University (2006), ONR & EPP (2006), Thamrongnawasawat et al. (2008), Duangdee et al. (2009,
2011), Nabhitabhata (2009), Sanpanich & Duangdee (2013).
Astralium stellare (Gmelin, 1791). Andaman Sea. Reported by: Wells (2005).
Bolma guttata (A. Adams, 1863). Andaman Sea. Reported by: BEDO (2017b).
Bolma henica (Watson, 1879). Andaman Sea. Reported by: Nabhitabhata (2009), Duangdee et al. (2011),
Sanpanich & Duangdee (2013).
Lunella cinerea (Born, 1778). Also reported as: Turbo cinereus (Born, 1778). Gulf of Thailand, Andaman Sea.
Reported by: THNHM, KUMF, Tantanasiriwong (1978), Nateewathana et al. (1981), Way & Purchon (1981),
Bussarawit (1995), Carpenter & Niem (1998), Duangdee (2004), Wells (2005), Thamrongnawasawat et al.
(2008), Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich & Duangdee (2013), BEDO (2017b), GBIF
(2019a), J. D. Taylor et al. (unpublished data).
Lunella coronata (Gmelin, 1791). Also reported as: Turbo coronatus (Gmelin, 1791). Gulf of Thailand, Andaman
Sea. Reported by: Carpenter & Niem (1998), Kasetsart University (2006), Nabhitabhata (2009), Duangdee et al.
(2011), Sanpanich & Duangdee (2013).
Turbo argyrostomus Linnaeus, 1758. Also reported as: Turbo (Marmarostoma) argyrostomus argyrostomus
Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Amornjaruchit (1988), Bussarawit (1995), Nateewathana (1995), Carpenter &
Niem (1998), Kasetsart University (2000, 2006), Alf & Kreipl (2003), Gemert (2003), Duangdee (2004), Wells
(2005), Thamrongnawasawat et al. (2008), Nabhitabhata (2009), Duangdee et al. (2011, 2013a, 2013b),
Sanpanich & Duangdee (2013, 2018), BEDO (2017b), GBIF (2019a, 2019b).
Turbo bruneus (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Tantanasiriwong (1978),
Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem (1998), Kasetsart University (2000, 2006),
Swennen et al. (2001), Duangdee (2004, 2007), ONR & EPP (2006), ONEP (2007), Thamrongnawasawat et al.
(2008), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata (2009), Sanpanich & Duangdee (2009, 2013),
Kostina et al. (2016), BEDO (2017b), GBIF (2019a, 2019b).
Turbo chrysostomus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Bussarawit (1995),
Carpenter & Niem (1998), BEDO (2017b).
Turbo crassus W. Wood, 1828. Andaman Sea. Reported by: GBIF (2019a).
Turbo elegans Philippi, 1846. Gulf of Thailand. Reported by: Morlet (1889). Listed by WoRMS Editorial Board
(2019) as a nomen dubium.
Turbo histrio Reeve, 1848. Andaman Sea. Reported by: GBIF (2019a).
Turbo heterocheilus Pilsbry, 1889. Gulf of Thailand. Reported by: Alf & Kreipl (2003), Gemert (2003), GBIF
(2019b).
Turbo intercostalis Menke, 1846. Also reported as: Turbo (Marmarostoma) intercostalis Menke, 1846. Gulf of
Thailand, Andaman Sea. Reported by: Morlet (1889), Alf & Kreipl (2003), GBIF (2019a, 2019b).
Turbo marmoratus Linnaeus, 1758. Also reported as: Turbo (Lunatica) marmoratus Linnaeus, 1758. Gulf of
Thailand, Andaman Sea. Reported by: THNHM, Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit
(1995), Chantrapornsyl (1995), Carpenter & Niem (1998), Alf & Kreipl (2003), Wells (2005), Nabhitabhata
(2009), Sanpanich & Duangdee (2013), GBIF (2019a).
Turbo petholatus Linnaeus, 1758. Gulf of Thailand, Andaman Sea. Reported by: THNHM, KUMF, Morlet (1889),
Suvatti (1938), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995), Carpenter & Niem
(1998), Gemert (2003), Duangdee (2004, 2007), Wells (2005), Kasetsart University (2006), ONR & EPP
(2006), ONEP (2007), Thamrongnawasawat et al. (2008), Duangdee et al. (2009, 2011, 2013a), Nabhitabhata
(2009), Sanpanich & Duangdee (2009, 2013, 2018), Hylleberg & Aungtonya (2013 ), BEDO (2017b), GBIF
(2019b).
Turbo setosus (Gmelin, 1791). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998), GBIF
(2019b).
Turbo sparverius Gmelin, 1791. Gulf of Thailand. Reported by: GBIF (2019b).
Turbo stenogyrus P. Fischer, 1873. Andaman Sea. Reported by: BEDO (2017b).
Turbo tursicus Reeve, 1848. Also reported as: Turbo (Marmarostoma?) tursicus Reeve, 1848; and Turbo
somnueki Patamakanthin, 2001. Andaman Sea. Reported by: Patamakanthin (2001), Alf & Kreipl (2003),
Thamrongnawasawat et al. (2008), Aungtonya & Sungjeen (2014), BEDO (2017b).
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The Marine and Estuarine Molluscs of Thailand
Class Polyplacophora
Subclass Neoloricata
Order Chitonida
Suborder Acanthochitonina
Superfamily Cryptoplacoidea
Family Acanthochitonidae
Craspedochiton laqueatus (G. B. Sowerby II, 1842). Reported as: Craspedochiton tetrica (Carpenter in Pilsbry,
1893). Andaman Sea. Reported by: Schwabe (2006).
Suborder Chitonina
Superfamily Chitonoidea
Family Chitonidae
Acanthopleura gemmata (Blainville, 1825). Gulf of Thailand, Andaman Sea. Reported by: Wells (2005), Duangdee
et al. (2009), GBIF (2019a), J. D. Taylor et al. (unpublished data).
Liolophura japonica (Lischke, 1873). Reported as: Nuttallina allantophora Dall, 1919. Gulf of Thailand, Andaman
Sea. Reported by: Kurozumi et al. (1989), Higo et al. (1999), GBIF (2019a, 2019b).
Onithochiton lyellii (Sowerby in Broderip & Sowerby, 1832). Andaman Sea. Reported by: Way & Purchon (1981).
Rhyssoplax pulcherrima (G. B. Sowerby II, 1842). Andaman Sea. Reported by: Kaas et al. (2006), GBIF (2019a).
Family Ischnochitonidae
Ischnochiton bouryi Dupuis, 1917. Andaman Sea. Reported by: GBIF (2019a).
Order Lepidopleurida
Suborder Lepidopleurina
Family Leptochitonidae
Leptochiton vietnamensis Sirenko, 1998. Andaman Sea. Reported by: Schwabe (2006).
Class Scaphopoda
Order Dentaliida
Family Dentaliidae
Dentalium aprinum Linnaeus, 1767. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Swennen
et al. (2001), GBIF (2019a).
Dentalium variabile Deshayes, 1825. Gulf of Thailand. Reported by: Robba et al. (2003, 2004), Chaiwathee et al.
(2007), Negri et al. (2014).
Graptacme eborea (Conrad, 1846). Reported as: Dentalium (Rhabdus) eburneum Linnaeus, 1767. Gulf of Thailand.
Reported by: Suvatti (1937, 1938), Nabhitabhata (2009).
Paradentalium hexagonum (Gould, 1859). Gulf of Thailand. Reported by: Robba et al. (2004).
Family Laevidentaliidae
Laevidentalium eburneum (Linnaeus, 1767). Andaman Sea. Reported by: Scarabino (1995, 2008), GBIF (2019a).
Family Gadilinidae
Episiphon virgula (Hedley, 1903). Reported as: Episiphon subrectum. Gulf of Thailand. Reported by: Robba et al.
(2003, 2004).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Order Gadilida
Suborder Gadilimorpha
Family Gadilidae
Dischides prionotus (R. B. Watson, 1879). Gulf of Thailand. Reported by: Robba et al. (2004), Sawatna et al.
(2012).
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The Marine and Estuarine Molluscs of Thailand
LITERATURE CITED
Abbott RT (1960) The genus Strombus in the Indo-Pacific. Indo-Pacific Mollusca, 1: 33–44.
Abbott RT (1961) The genus Lambis in the Indo-Pacific. Indo-Pacific Mollusca, 1: 147–174.
Abbott RT (1968) The helmet shells of the world (Cassidae). Part 1. Indo-Pacific Mollusca, 2: 7–202.
Adamkewicz L (1989) Differences in the frequencies of several shell characters in the clam Donax variabilis around
Cape Hatteras, North Carolina. The Veliger, 32: 21–28.
Adulyanukosol K & Poovachiranon S (2007) Dugong (Dugong dugon) and seagrass in Thailand: Present status and
future challenges. In: Arai N (ed.) Proceedings of the 3rd International Symposium on SEASTAR2000 and Asian
Bio–logging Science (The 7th SEASTAR2000 Workshop). Graduate School of Informatics, Kyoto University,
Japan, pp. 41–50.
ALA (2019–2020) Atlas of Living Australia. https://ala.org.au (Accessed 23 August 2020).
Alf A & Kreipl K (2003) The Family Turbinidae, Subfamily Turbininae, Genus Turbo. A Conchological Iconography.
ConchBooks, Hackenheim, 68 pp.
Altena CO van R & Gittenberger E (1981) The genus Babylonia (Prosobranchia, Buccinidae). Zoologische
Verhandelingen, 188: 1–57.
Amornjaruchit S (1988) Economically important molluscan shellfish of Thailand. In: McCoy EW & Chongpeepien
T (eds.) Bivalve Mollusc Culture Research in Thailand. ICLARM Technical Reports 19. Department of Fisheries,
Bangkok, Thailand; International Center for Living Aquatic Resources Management, Manila, Philippines; and
Deutsche Gesellschaft filr Technische Zusarnmenarbeit (GTZ), GmbH, Eschborn, Federal Republic of Germany,
pp. 1–18.
AMS (2020) Discover and Learn. Australian Museum, Sydney. https://australianmuseum.net.au/learn/ (Accessed
6 April 2020).
Aungtonya C, Bussabong K, Wongkamhaeng K, Eedkerd I & Ubonsurwan S (2006) Type specimens in the reference
collection of Phuket Marine Biological Center. Reference Collection and Aquarium Contributions, 4: 1–45. [In
Thai].
Aungtonya C & Hylleberg J (1998) Check list of sublittoral molluscs, with nine new records for the Andaman Sea.
Phuket Marine Biological Center Special Publication, 18: 317–322.
Aungtonya C, Nateewathana A & Tendal OS (2014) A catalogue of the samples of the class Cephalopoda (Phylum
Mollusca) in the reference collection of Phuket Marine Biological Center. Phuket Marine Biological Center
Special Publication, 32: 119–128.
Aungtonya C, Nateewathana A, Tendal OS & Nabhitabhata J (2011) New records of the bobtail squid, Euprymna
hyllebergi Nateewathana, 1997 with designation of a neotype. Phuket Marine Biological Center Research
Bulletin, 70: 15–27.
Aungtonya C & Sungjeen S (2014) Type specimens in the reference collection of Phuket Marine Biological Center,
Thailand. Phuket Marine Biological Center Special Publication, 32: 129–150.
Aungtonya C, Thaiklang N & Srisawan D (1999) Recent records of Bivalvia in the reference collection of Phuket
Marine Biological Center, Thailand. Phuket Marine Biological Center Special Publication, 19: 371–383.
Aungtonya C, Thaipatl S & Tendal O (2000) A preliminary report on the Thai–Danish Bioshelf surveys (1996–2000)
of the west coast of Thailand, Andaman Sea. Phuket Marine Biological Center Research Bulletin, 63: 53–76.
Azuma M (1976) Systematic studies on the recent Japanese family Ovulidae (Gastropoda)-V. Venus, 35: 185–206.
Banchongmanee S & Upanoi T (2000) Giant clams in the Andaman Sea, Thailand. Part 1: Distribution and abundance
at Rawi, Butang and Kata Islands. Phuket Marine Biological Center Special Publication, 21: 69–73.
Bech M (1996) Field release of cultured muricid gastropods (Chicoreus ramosus) at artificial reefs in the Andaman
Sea, Thailand. Phuket Marine Biological Center Special Publication, 16: 53–59.
Bech M (1998) Imposex and population characteristics of Thais distinguenda as an indicator of organotin
contamination along the south east coast of Phuket Island, Thailand. Phuket Marine Biological Center Special
Publication, 18: 129–138.
BEDO (2017a) Checklist of Marine Mollusks in Thailand (Bivalvia). Biodiversity -Based Economy Development
Office, Bangkok, 296 pp. [In Thai].
BEDO (2017b) Checklist of Marine Mollusks in Thailand (Gastropoda). Biodiversity-Based Economy Development
Office, Bangkok, 380 pp. [In Thai].
BEDO (2017c) Checklist of Marine Mollusks in Thailand (Nudibranchia). Biodiversity-Based Economy Development
Office, Bangkok, 148 pp. [In Thai].
BEDO (2017d) Checklist of Marine Mollusks in Thailand (Cephalopoda). Biodiversity-Based Economy Development
Office, Bangkok, 96 pp. [In Thai].
BEDO (2017e) Checklist of Marine Mollusks in Thailand (Microsnails). Biodiversity-Based Economy Development
Office, Bangkok, 96 pp. [In Thai].
Bergh R (1902) The Danish Expedition to Siam 1899–1900. Det Kongelige Danske Videnskabernes Selskab
Skrifter. 6 række. Naturvidenskab og Mathematik Afdeling, 12: 153–218.
166
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Bernard FR, Cai YY & Morton B (1993) Catalogue of the Living Marine Bivalve Molluscs of China. Hong Kong
University Press, Hong Kong, 146 pp.
Bernard PA (ed.) (1984) Coquillages du Gabon [Shells of Gabon]. Pierre A. Bernard, Libreville, Gabon, 140 pp.
Bertin V (1881) Révision des Donacidées du Muséum d´Histoire naturelle (Paris). Nouvelles archives du Muséum
d’Histoire naturelle, série 2, 4: 57–121.
Beu AG (1987) Taxonomy of gastropods of the families Ranellidae (= Cymatiidae) and Bursidae. Part 2. Descriptions
of 14 new Indo-West Pacific taxa, with revisions of related taxa. New Zealand Journal of Zoology, 13: 273–355.
Beu AG (1998) Indo-West Pacific Ranellidae, Bursidae and Personidae (Mollusca: Gastropoda): A monograph of
the New Caledonian fauna and revisions of related taxa. Mémoires du Muséum national d’Histoire naturelle,
178: 1–255.
Beu AG (2005) Neogene fossil tonnoidean gastropods of Indonesia. Scripta Geologica, 130: 1–186.
Beu AG & Cernohorsky WO (1986) Taxonomy of gastropods of the families Ranellidae (= Cymatiidae) an d
Bursidae. Part 1. Adoption of Ranellidae, and revision of Linatella Gray, 1857. New Zealand Journal of Zoology,
13: 241–266.
Beu AG & Kay EA (1988) Taxonomy of gastropods of the families Ranellidae (= Cymatiidae) and Bursidae. Part IV.
The Cymatium pileare complex. Journal of the Royal Society of New Zealand, 18: 185–223.
Bieler R (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des
Naturwissenschaftlichen Vereins in Hamburg (NF), 30: 1–376.
Bieler R, Chalermwat K, Mikkelsen P & Wells FE (2008a) International Marine Bivalve Workshop 2005: Introduction
and summary. In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand:
Proceedings of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005,
with contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 1–6.
Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) (2008b) Molluscs of Eastern Thailand: Proceedings
of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005, with
contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 1 –264.
Bouchet P (1975) Nudibranches nouveaux des côtes du Sénégal. Vie et Milieu, 25: 119–132.
Bouchet P & Perrine D (1996) More gastropods feeding at night on parrotfishes. Bulletin of Marine Science, 59:
224–228.
Brandt RAM (1974) The non–marine aquatic Mollusca of Thailand. Archiv für Molluskenkunde, 105: 1–423.
Bratcher T & Cernohorsky WO (1987) Living Terebras of the World. American Malacologists, Melbourne, Florida,
240 pp.
Brown LG & Neville BD (2015) Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca:
Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa, 3907: 1–188.
Bruce N, Berggren M & Bussarawit S (eds.) (2002) Proceedings of the International Workshop on the Crustacea of
the Andaman Sea, part 1 and 2. Phuket Marine Biological Center Special Publication, 23: 1–532.
Brunckhorst D (1993) The systematics and phylogeny of phyllidiid nudibranchs (Doridoidea). Records of the
Australian Museum, Supplement 16: 1–107.
Burch JQ & Burch RL (1967) The family Olividae. Pacific Science, 21: 503–522.
Bussarawit S (1991) The family Muricidae in the PMBC reference collection: Application of a database system.
Phuket Marine Biological Center Special Publication, 9: 29–34.
Bussarawit S (1992) Field surveys on muricids in Thai waters. Phuket Marine Biological Center Special Publication,
10: 143–149.
Bussarawit S (1995) Molluscs from the marine national parks: Surin and Le–Pae Island, Andaman Sea, Thailand.
Phuket Marine Biological Center Special Publication, 15: 119–126.
Bussarawit S & Cedhagen T (2010) Field Guide to the Oyster Fauna of Thailand. Kyoto University Press, Japan,
47 pp.
Bussarawit S, Gravlund P, Glenner H & Rasmussen AR (2006) Phylogenetic analysis of Thai oyster (Ostreidae)
based on partial sequences of the mitochondrial 165tdna gene. Phuket Marine Biological Center Research
Bulletin, 67: 1–9.
Cai L-Z, Hwang J-S, Dahms H-U, Fu S-J & Guo T (2014) Effect of the invasive bivalve Mytilopsis sallei on the
macrofaunal fouling community and the environment of Yundang Lagoon, Xiamen, China. Hydrobiologia, 741:
101–111.
Callomon P & Snyder MA (2008) A new species of Fusinus from Thailand and the Andaman Islands with three
distinct growth phases. Venus, 66: 119–126.
Camacho-García YE & Gosliner TM (2008) Systematic revision of Jorunna Bergh, 1876 (Nudibranchia:
Discodorididae) with a morphological phylogenetic analysis. Journal of Molluscan Studies, 74: 143 –181.
Carlton JT (1992) Introduced marine and estuarine mollusks of North Am erica: An end-of-the-20th-century
perspective. Journal of Shellfish Research, 11: 489–505.
Carpenter KE & Niem VH (eds.) (1998) FAO species identification guide for fishery purpose. The living marine
resources of the Western Central Pacific. Volume 1. Seaweeds, corals, bivalves and gastropods. Food and
Agricultural Organization, Rome, 686 pp.
Cate CN (1973) A systematic revision of the recent cypraeid family Ovulidae. The Veliger, Supplement 15: 1–116.
167
The Marine and Estuarine Molluscs of Thailand
Cernohorsky WO (1972) Marine Shells of the Pacific, Volume II. Pacific Publications, Sydney, 411 pp.
Cernohorsky WO (1976a) The Mitridae of the world. Part I. The subfamily Mitrinae. Indo-Pacific Mollusca, 3:
373–528.
Cernohorsky WO (1976b) The taxonomy of some Indo-Pacific Mollusca. Part 4. With descriptions of new taxa and
remarks on Nassarius coppingeri (Smith). Records of the Auckland Institute and Museum, 13: 111–129.
Cernohorsky WO (1981) The family Buccinidae. Part 1: The genera Nassaria, Trajana and Neoteron. Monographs
of Marine Mollusca, 2: 1–52.
Cernohorsky WO (1984) Systematics of the family Nassariidae (Mollusca: Gastropoda). Bulletin of the Auckland
Institute and Museum, 14: 1–356.
Cernohorsky WO (1991) The Mitridae of the world. Part 2. The subfamily Mitrinae concluded and subfamilies
Imbricariinae and Cylindromitrinae. Monographs of Marine Mollusca, 4: 1–164.
Chaitiamwong S (1993) Identification of cephalopods found in the Gulf of Thailand. Bangkok Marine Fisheries
Development Center, Marine Fisheries Division, Department Fisheries, Technical Report, 23: 1–78.
Chaitiamwong S, Devahudi T & Waritswat A (1971) Review of taxonomic nomenclature of some commercially
important shellfish in Thai waters. In: Second Symposium on Marine Fisheries , April 19–20, 1971. Marine
Fisheries Laboratory, Bangkok, pp. 1–28.
Chaiwathee S, Dumrongrojwattana P, Matchacheep S & Chalermwat C (2007) Pre-list of some marine microsnails
from Bangsaen beach, Chonburi province (Mollusca: Scaphopoda and Gastropoda). In: Proceedings of 45th
Kasetsart University Annual Conference: Science. January 30–February 2, 2007. Kasetsart University, Bangkok,
pp. 573–576.
Chalermwat K, Szuster BW & Flaherty M (2003) Shellfish aquaculture in Thailand. Aquaculture Economics &
Management, 7: 249–261.
Chang KM (1985) Newly introduced false mussel to Taiwan (Bivalvia: Dreissenidae). Bulletin of Malacology, ROC,
11: 61–67.
Chantawong P & Suksawat C (1997) Cephalopod distribution and abundance in the northern part of Phang–nga
Province, Thailand. Phuket Marine Biological Center Special Publication, 17: 181–191.
Chantrapornsyl S (1995) Green turbo (Turbo mamoratus Linnaeus), an endangered species in Thailand. Phuket
Marine Biological Center Special Publication, 15: 39–40.
Chantrapornsyl S, Kittiwattanawong K & Adulyanukosol K (1996) Distribution and abundance of giant clam
around Lee–Pae Island. The Andaman Sea, Thailand. Phuket Marine Biological Center Special Publication, 16:
195–200.
Chavanich S, Tan LT & Vallejo B (eds.) (2010a) Report on the Current Status of Marine Non–Indigenous Species in
the Western Pacific Region. Intergovernmental Oceanographic Commission Subcommission for the Western
Pacific (IOC/WESTPAC), Bangkok, Thailand, 61 pp.
Chavanich S, Harris LG & Voranop V (2010b) Nudibranchs of Thailand. Biodiversity Research and Training
Program, Bangkok, 104 pp.
Chavanich S, Viyakarn V, Sanpanich K & Harris LG (2013) Diversity and occurrence of nudibranchs in Thailand.
Marine Biodiversity, 43: 31–36.
Chotiyaputta C, Nootmorn P & Jirapunpipat K (2002) Review of cephalopod fishery production and long term
changes in fish communities in the Gulf of Thailand. Bulletin of Marine Science, 71: 223–238.
Chotiyaputta C, Okutani T & Chaitiamvong S (1991) A new pygmy cuttlefish from the Gulf of Thailand, Idiosepius
thailandicus n. sp. (Cephalopoda: Idiosepiidae). Venus, 50: 165–174.
Chotiyaputta C, Okutani T & Chaitiamvong S (1992) Systematic Study of Cephalopods in Thailand. Final Report of
JSPS–NRCT. National Research Council Thailand, Bangkok, 100 pp.
Cilia DP & Deidun A (2012) Branching out: Mapping the spatial expansion of the Lessepsian invader mytilid
Brachidontes pharaonis around the Maltese Islands. Marine Biodiversity Records, Marine Biological Association
of the United Kingdom, 5: 1–8.
Coan EV, Lutaenko KA, Zhang J & Sun Q (2015) The molluscan taxa of A. W. Grabau & S. G. King (1928) and their
types. Malacologia, 58: 179–224.
Coan EV & Valentich-Scott P (2012) Bivalve Seashells of Tropical West America. Marine Bivalve Mollusks from
Baja California to Northern Peru. Santa Barbara Museum of Natural History, Santa Barbara, California, 1258 pp.
Coomans HE, Moolenbeek RG & Wils E (1980) Alphabetical revision of the (sub)species in recent Conidae. 3. albus
to antillarum with the description of Conus algoensis agulhasi, nov. subspecies. Basteria, 44: 17–49.
Coomans HE, Moolenbeek RG & Wils E (1986) Alphabetical revision of the (sub)species in recent Conidae.
9. ebraeus to extraordinarius with the description of Conus elegans ramalhoi, nov. subspecies. Basteria, 50:
93–150.
Coppejans E, Prathep A, Leliaert F, Lewmanomont K & De Clerck O (2010) Seaweeds of Mu Ko Tha Lae Tai (SE
Thailand). Methodologies and Field Guide to the Dominant Species. Biodiversity Research and Training
Program (BRT), Bangkok, 274 pp.
Coppejans E, Prathep A, Lewmanomont K, Hayashizaki K, De Clerck O, Leliaert F, Terada R (2017) Seaweeds and
Seagrasses of the Southern Andaman Sea Coast of Thailand. Kagoshima University Museum, Kagoshima City,
245 pp.
168
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Cosel R von (2002) Seven new species of Solen (Bivalvia: Solenidae) from the tropical Western Pacific, with remarks
on other species. In: Falkner M, Groh K & Speight MCD (eds.) Collectanea Malagologica: Festschrift für Gerhard
Falkner. ConchBooks, Hackenheim, Germany; Verlag der Friedrich–Held–Gesellschaft, München, Germany, pp.
301–341.
Cossignani T (1994) Bursidae of the World. Mostra Mondiale Malacologia, Ancona, Italy, 119 pp.
Cossignani T (2006) Marginellidae and Cystiscidae of the World. L’Informatore Piceno, Ancona, Italy, 408 pp.
Cotton BC (1961) South Australian Mollusca Pelecypoda. W. L. Hawkes, Government Printer, Adelaide, 363 pp.
Crowe TP & Frid CLJ (eds.) (2015) Marine Ecosystems: Human Impacts on Biodiversity, Functioning and Services.
Cambridge University Press, Cambridge, 409 pp.
D’Asaro CN (1992) Gunnar Thorson’s world-wide collection of prosobranch egg capsules: Muricidae. Ophelia, 35:
1–101.
D’Asaro CN (1997) Gunnar Thorson’s world-wide collection of prosobranch egg capsules: Melongenidae. Ophelia,
46: 83–125.
da Motta AJ (1981) A new Andaman Sea olive. La Conchiglia, 13: 20–21.
da Motta AJ & Lenevat P (1979) Conus shells of Thailand. da Motta AJ & Lenevat P, Bangkok, 20 pp.
Dance SP & Poppe GT (1999) Family Harpidae. A Conchological Iconography. ConchBooks, Hackenheim, Germany,
68 pp.
Dayrat B (2006) A taxonomic revision of Paradoris sea slugs (Mollusca, Gastropoda, Nudibranchia, Doridina).
Zoological Journal of the Linnean Society, 147: 125–238.
Dayrat B (2010) A monographic revision of basal discodorid sea slugs (Mollusca: Gastropoda: Nudibranchia:
Doridina). Proceedings of the California Academy of Sciences, Series 4, 61, Supplement I: 1–403.
Dayrat B (2011) A warning for ecologists and conservation biologists using species checklists: How the European
marine fauna ‘lost’ all of its 16 Discodoris species (Mollusca: Gastropoda). Organisms, Diversity and Evolution,
11: 75–82.
Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Quang NX, Tan SK & Tan SH (2016) Integrative taxonomy
of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys, 636: 1–40.
Dayrat B, Goulding TC, Apte D, Bhave V & Xuân QN (2017) A new genus and four new species of onchidiid
slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae). Journal of Natural History, 51:
1851–1897.
Dayrat B, Goulding TC, Khalil M, Apte D & Tan SH (2019) A new species and new records of Onchidium slugs
(Gastropoda, Euthyneura, Pulmonata, Onchidiidae) in South-East Asia. Zookeys, 892: 27–58.
Dechruksa W, Sritongtae S, Namchoate S & Krailis D (2014) Species diversity of molluscs obtained from mangrove
forest of estuary and river in the eastern coast of Thailand. In: Tokyo International Conference on Life Science
and Biological Engineering 2014. Pp. 931–937.
Dekker H (1996) Strombus orchroglottis betuleti Kronenberg, 1991 in Thailand (Gastropoda: Strombidae). Vita
Marina, 44: 19.
Dekker H (2000) The Neritidae (Gastropoda) from the circumarabian seas, with the description of two new species,
a new subgenus and a new genus. Vita Marina, 47: 29–64.
Dekker H & Kool HH (1999) A re-evaluation of the Nassarius albescens (Dunker, 1846) species-group (Gastropoda:
Nassariidae). Vita Marina, 46: 53–60.
Derrick B, Noranarttragoon P, Zeller D, Teh LCL & Pauly D (2017) Thailand’s missing marine fisheries catch (1950–
2014). Frontiers in Marine Science, 4: 1–11.
Dharma B (1988) Siput dan Kerang Indonesia (Indonesian Shells). PT Sarana Graha, Jakarta, 111 pp.
Dijkstra HH (2008) A new species of recent scallop of the genus Serratovola (Bivalvia, Pectinidae) from the tropical
Indo-West Pacific. Basteria, 72: 57–63.
Dijkstra HH (2013) Pectinoidea (Bivalvia: Propeamussiidae and Pectinidae) from the Panglao region, Philippine
Islands. Vita Malacologica, 10: 1–108.
Dijkstra HH & Beu AG (2018) Living scallops of Australia and adjacent waters (Mollusca: Bivalvia: Pectinoidea:
Propeamussiidae, Cyclochlamydidae and Pectinidae). Records of the Australian Museum, 70: 113–330.
Dijkstra HH & Kilburn RN (2001) The family Pectinidae in South Africa and Mozambique (Mollusca: Bivalvia:
Pectinoidea). African Invertebrates, 42: 263–321.
Dijkstra HH & Knudsen J (1998) Some Pectinoidea (Mollusca: Bivalvia: Propeamussiidae, Pectinidae) of the Red
Sea. Molluscan Research, 19: 43–103.
Dijkstra HH & Marshall BA (1997) Pectinoidea (Mollusca: Bivalvia: Propeamussiidae: Pectinidae) of Lord Howe
Island, Norfolk Island and the Kermadec Islands. Molluscan Research, 18: 73–114.
DiscoverLife (2020) University of Georgia, USA; University of New South Wales, Australia.
https://www.discoverlife.org/ (Accessed 13 May 2020).
Dogan A, Önen M & Öztürk B (2007) A new record of the invasive Red Sea mussel Brachidontes pharaonis (Fischer
P., 1870) (Bivalvia: Mytilidae) from the Turkish coasts. Aquatic Invasions, 2: 461–463.
Dornellas APS & Simone LRL (2011) Annotated list of type specimens of mollusks deposited in Museu de Zoologia
da Universidade de São Paulo, Brazil. Arquivos de Zoologia, 42: 1–81.
169
The Marine and Estuarine Molluscs of Thailand
Duangdee T (2004) Biodiversity of mollusks of Chang Islands, Trat Province. In: Meksumpun S (ed.) Final Report.
Research Program on Aquatic Resources and Environment Potentials for Development of Eco –tourism of
Chang Islands. Department of Marine Science, Kasetsart University, Bangkok, Thailand, pp. 1–39.
Duangdee T (2007) Species diversity and distribution of molluscs in coral reefs and adjacent areas at Ang Thong
Islands, Surat Thani province. In: Proceedings of 45th Kasetsart University Annual Conference: Fisheries .
January 30–February 2, 2007. Kasetsart University, Bangkok, pp. 373–381.
Duangdee T, Sanpanich K & Jitchoom P (2007) Marine Mollusks in Kram Island and Adjacent Area. Plant Genetic
Conservation Project Under the Royal Initiative of Her Royal Highness Princess Maha Chakri Sirindhorn,
Bangkok, 76 pp.
Duangdee T, Sanpanich K & Thongpinyochai S (2009) The species diversity of marine mollusks in coral reefs and
adjacent areas at Tarutoa National Park, Satun Province. In: Proceeding of 4th RSPG Researchers Club
Conference “Thai Resources: Turn to the New Way in Thai Way (Thai Innovation)”. 20th–23rd October 2009,
Khao Kheow Open Zoo, Si Racha, Chon Buri Province. Pp. 251–266.
Duangdee T, Sanpanich K & Thongpinyochai S (2011) The species diversity of marine mollusks in coral reefs at
Ang Thong Islands, Surat Thani Province. In: Proceeding of 5th RSPG Researchers Club Conference “Step to
The World Wide with Confidence”. 3rd–5th November 2011, Nong Rawieng Training Center, Rajamangala
University of Technology Isan, Nakhon Ratchasima Province. Pp. 235–250.
Duangdee T, Sanpanich K & Tongpinyochai S (2013a) The species diversity of marine molluscs in Surin Islands,
Phangnga Province. In: The 6th Conference of RSPG Botanical and Biological Club. 20–26 December 2013,
Srinagarind Dam, Electricity Generating Authority of Thailand, Sri Sawat District, Kanchanaburi Province .
Pp. 294–310.
Duangdee T, Sanpanich K, Kuisorn V & Tongpinyochai S (2013b) Marine Molluscs Along Similan Islands. Work
Square Co. Ltd., Bangkok, 104 pp.
Dumrongrojwattana P, Tanamaib S, Wongkamhaeng K & Nabhitabhata J (2016) Triphorid snails family from
Similan Islands (Gastropoda: Triphoroidea: Triphoridae), Southern Thailand. In: The 5th Burapha University
International Conference, July 28–29, 2016, Pattaya Thailand. Burapha University, Bang Saen, Thailand, pp.
283–287.
DuShane H (1988a) Geographical distribution of some Epitoniidae (Mollusca: Gastropoda) associated with fungiid
corals. The Nautilus, 102: 30–35.
DuShane H (1988b) New Hawaiian species of Epitoniidae (Mollusca: Gastropoda). The Veliger, 31: 267–271.
Eibye-Jacobsen D (ed.) (2002) Proceedings of the international workshop on the polychaetes of the Andaman Sea.
Phuket Marine Biological Center Special Publication, 24: 1–424.
Eichhorst TE (2016) Neritidae of the World. ConchBooks, Hackenheim, 694 pp.
Emerson WK (1986) A new species of Morum from the Andaman Sea (Gastropoda: Volutacea). The Nautilus, 100:
96–98.
Emerson WK & Cernohorsky WO (1973) The genus Drupa in the Indo-Pacific. Indo-Pacific Mollusca, 3: 1–40.
Engl W (1997) New species of the family Eulimidae from the Canary Islands. Part I: Description of Sticteulima
richteri n. sp. La Conchiglia, 283: 44–47.
FAO (2019) Food and Agriculture Organization of the United Nations. Fishery and Aquaculture Country Profiles.
The Kingdom of Thailand. http://www.fao.org/fishery/facp/THA/en (Accessed 24 May 2019).
Forestry Research Center (2006) Biodiversity of marine & coastal ecosystems. In: Biodiversity Survey & Information
System Project. Volume 3. Final report presented to the Office of Natural Resources and Environmental Policy
and Planning. Faculty of Forestry, Kasetsart University, Bangkok, pp. 11–117.
Fossilworks (2020) Fossilworks. http://fossilworks.org (Accessed 23 August 2020).
Fraussen K & Stratmann D (2013) The Family Babyloniidae. A Conchological Iconography. ConchBooks, Hackenheim,
146 pp.
Fukumori H & Kano Y (2014) Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods.
Marine Biology, 161: 213–227.
Fukumori H, Yahagi T, Warén A & Kano Y (2019) Amended generic classification of the marine gastropod family
Phenacolepadidae: Transitions from snails to limpets and shallow-water to deep-sea hydrothermal vents and
cold seeps. Zoological Journal of the Linnean Society, 185: 636–655.
Furuse K & Hasegawa K (1984) Mytilopsis sallei found in Tokyo Bay. Chiribotan, 15: 18.
Galil BS (2000) A sea under siege—alien species in the Mediterranean. Biological Invasions, 2: 177–186.
Galindo LA, Kool HH & Dekker H (2017) Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae):
Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of
Taxonomy, 275: 1–43.
Galli C (2015) WMSDB – Worldwide Mollusc Species Data Base. Bagni Liggia, Genova, Italy. Last updated 7 April
2016. https://www.bagniliggia.it/WMSD/WMSDhome.htm (Accessed 23 August 2020).
Ganmanee M, Narita T, Iida S & Sekiguchi H (2003) Feeding habits of asteroids, Luidia quinaria and Astropecten
scoparius, in Ise Bay, Central Japan. Fisheries Science, 69: 1121–1134.
García AA & Oliver G (2008) Species discrimination in seven species of Barbatia (Bivalvia: Arcoidea) from Thailand
with a redescription of B. grayana (Dunker, 1858). In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE
170
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
(eds.) Molluscs of Eastern Thailand. Proceedings of the International Marine Bivalve Workshop, Chanthaburi,
Thailand, August–September 2005, with contributions on other molluscan groups. Raffles Bulletin of Zoology,
Supplement 18: 7–23.
Garcia EF (2001) A new species of Latirus (Gastropoda: Fasciolariidae) from the southern Philippines. Novapex, 2:
149–151.
GBIF (2019a) GBIF.org (14 April 2019) GBIF Occurrence Download. https://doi.org/10.15468/dl.kcq0t5
(Accessed 14 April 2019).
GBIF (2019b) GBIF.org (17 April 2019) GBIF Occurrence Download. https://doi.org/10.15468/dl.9wbho5
(Accessed 18 April 2019).
GBIF (2019c) Lolliguncula panamensis Berry, 1911, in GBIF Secretariat. GBIF Backbone Taxonomy.
https://www.gbif.org/species/2290355 (Accessed 3 April 2019).
GBIF (2020) GBIF Occurrence Search. https://www.gbif.org/occurrence/search (Accessed 21 December 2020).
Geiger DL & Jansen P (2004) New species of Australian Scissurellidae (Mollusca: Gastropoda: Vetigastropoda) with
remarks on Australian and Indo-Malayan species. Zootaxa, 714: 1–72.
Geiger DL & Owen B (2012) Abalone Worldwide Haliotidae. ConchBooks, Hackenheim, 156 pp.
Geiger DL & Poppe GT (2000) The Family Haliotidae. A Conchological Iconography. ConchBooks, Hackenheim,
218 pp.
Gemert L van (2003) Marine schelpen verzamelen op Ko Samui (Golf van Thailand). De Kreukel, 39: 95–109.
Gemert LJ van (2016) Note on the publications of Edmond Saurin (1904–1977) on recent Mollusca; with a list of
all his new pyramidellids (Gastropoda, Pyramidellidae). Basteria, 80: 133–138.
GISD (2015a) Species profile: Ilyanassa obsoleta. Global Invasive Species Database. Invasive Species Specialist
Group. http://www.iucngisd.org/gisd/species.php?sc=1321 (Accessed 6 May 2019).
GISD (2015b) Species profile: Ostrea edulis. Global Invasive Species Database. Invasive Species Specialist Group.
http://www.iucngisd.org/gisd/species.php?sc=798 (Accessed 6 May 2019).
Gittenberger A (2003) The wentletrap Epitonium hartogi spec. nov. (Gastropoda: Epitoniidae), associated with
bubble coral species, Plerogyra spec. (Scleractinia: Euphylliidae), off Indonesia and Thailand. Zoologische
Verhandelingen Leiden, 345: 139–150.
Gittenberger A & Gittenberger E (2005) A hitherto unnoticed adaptive radiation: Epitoniid species (Gastropoda:
Epitoniidae) associated with corals (Scleractinia). Contributions to Zoology, 74: 125–203.
Gittenberger A & Gittenberger E (2011) Cryptic, adaptive radiation of endoparasitic snails: Sibling species of
Leptoconchus (Gastropoda: Coralliophilidae) in corals. Organisms, Diversity and Evolution, 11: 21–41.
Glover EA & Taylor JD (2001) Systematic revision of Australian and Indo-Pacific Lucinidae (Mollusca: Bivalvia):
Pillucina, Wallucina and descriptions of two new genera and four new species. Records of the Australian
Museum, 53: 263–292.
Glover EA & Taylor JD (2005) Diversity of chemosymbiotic bivalves on coral reefs: Lucinidae (Mollusca, Bivalvia)
of New Caledonia and Lifou. Zoosystema, 29: 109–181.
Glover EA & Taylor JD (2016) Lucinidae of the Philippines: Highest known diversity and ubiquity of
chemosymbiotic bivalves from intertidal to bathyal depths (Mollusca: Bivalvia). Tropical Deep-Sea Benthos,
29 (Mémoires du Muséum national d’Histoire naturelle, 208): 65–234.
Glover EA, Taylor JD & Williams ST (2008) Mangrove–associated lucinid bivalves of the central Indo-West Pacific:
Review of the “Austriella” group with a new genus and species (Mollusca: Bivalvia: Lucinidae). In: Bieler R,
Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand. Proceedings of the
International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005, with contributions
on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 25–40.
Glover EA, Williams ST & Taylor JD (2016) Lucinid bivalves of Singapore and their relationships (Bivalvia:
Lucinidae). Raffles Bulletin of Zoology, Supplement 34: 539–565.
Golding RE (2014) Molecular phylogeny and systematics of Australian and East Timorese Stenothyridae
(Caenogastropoda: Truncatelloidea). Molluscan Research, 34: 102–126.
Gosliner TM, Behrens DW & Valdés Á (2008) Indo-Pacific Nudibranchs and Sea Slugs: A Field Guide to the World’s
Most Diverse Fauna. Sea Challengers Natural History Books & The California Academy of Sciences, San
Francisco, 426 pp.
Gosliner TM & Fahey SJ (2011) Previously undocumented diversity and abundance of cryptic species: A phylogenetic
analysis of Indo-Pacific Arminidae Rafinesque, 1814 (Mollusca: Nudibranchia) with descriptions of 20 new
species of Dermatobranchus. Zoological Journal of the Linnean Society, 161: 245–356.
Habe T (1965) Family Solenidae in Japan and its adjacent areas. Venus, 23: 188–197.
Habe T (1980) Mytilopsis sallei (Récluz, 1849), a newly introduced mussel to Japan. Chiribotan, 11: 41–42.
Haga T (2011) Pholadidae. In: Poppe GT (ed.) Philippine Marine Mollusks, Volume IV. ConchBooks, Hackenheim,
pp. 392–397, pls. 1191–1193.
Harry HW (1969) Anatomical notes on the mactrid bivalve, Reata plicatella Lamarck, 1818, with a review of the
genus Raeta and related genera. The Veliger, 12: 1–23.
Heiman EL (2010) Blasicrura interrupta (Gray, 1824) and related taxa. Triton, 21, Supplement 3: 1–18.
171
The Marine and Estuarine Molluscs of Thailand
Highsmith RC (1980) Burrowing by the bivalve mollusc Lithophaga curta in the living reef coral Montipora berryi
and a hypothesis of reciprocal larval recruitment. Marine Biology, 56: 155–162.
Higo S, Callomon P & Goto Y (1999) Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan. Elle
Scientific Publications, Osaka, Japan, 749 pp.
Hoegh-Guldberg O (2009) The Coral Triangle and Climate Change: Ecosystems, People, and Societies at Risk. WWF
Australia, Sydney, 228 pp.
Hong PN & San HT (1993) Mangroves of Vietnam. IUCN Wetlands Programme, Bangkok, Thailand, 173 pp.
Houart R (1992) The genus Chicoreus and related genera (Gastropoda: Muricidae) in the Indo-West Pacific.
Mémoires du Muséum national d’Histoire naturelle, Sér. A–Zoologie, 154: 1–188.
Houart R (2001) Chicoreus (Triplex) setionoi n. sp. (Gastropoda: Muricidae) from Arafura Sea, Pacific Ocean.
Novapex, 2: 145–148.
Houart R (2002) Comments on a group of small Morula s.s. species (Gastropoda: Muricidae: Rapaninae) from the
Indo-West Pacific with the description of two new species. Novapex, 3: 97–118.
Houart R (2004) Review of the recent species of Morula (Oppomorus), M. (Azumamorula) and M. (Habromorula)
(Gastropoda: Muricidae: Ergalataxinae). Novapex, 5: 91–130.
Houart R (2008) Description of a new species of Chicoreus (Triplex) Perry, 1811 (Gastropoda: Muricidae) from
Palawan, Philippine Islands. Novapex, 9: 165–170.
Houart R (2010) Description of a new species from Indonesia in the Murex scolopax group (Mollusca: Gastropoda:
Muricidae) and comments about Murex (Murex) ternispina Lamarck, 1822 from East Java. Novapex, 11: 29–35.
Houart R (2014) Living Muricidae of the World: Muricinae. ConchBooks, Harxheim, 197 pp.
Houart R (2016) On some Indo-West Pacific species of Favartia sensu stricto (Gastropoda: Muricidae:
Muricopsinae) with the description of three new species from the Indian Ocean and comments on related
species. Vita Malacologica, 15: 57–76.
Houart R (2017) Description of eight new species and one new genus of Muricidae (Gastropoda) from the IndoWest Pacific. Novapex, 18: 81–103.
Houart R & Delemarre J-L (2013) Note sur la présence de Indothais blanfordi (Melvill, 1893) (Muricidae:
Rapaninae) au Sénégal et en Guinée-Bissau et commentaires sur deux autres espèces d’Afrique Occidentale.
Xenophora, 142: 13–20.
Houart R, Fraussen K & Barbier J-P (2013) Description of a new species of Lataxiena (Gastropoda: Muricidae) from
India. Gloria Maris, 52: 1–10.
Houart R & Gori S (2011) Description of two new Favartia species (Gastropoda: Muricidae: Muricopsinae) from
Masirah Island, Oman, Arabian Peninsula. Novapex, 12: 39–45.
Houart R & Vilvens C (1997) Description of a new species of Drupa Röding, 1798 (Gastropoda: Muricidae:
Rapaninae) from the Western Indian Océan. Apex, 12: 125–131.
Houbrick RS (1978) The family Cerithiidae in the Indo-Pacific. Part 1: The genera Rhinoclavis, Pseudovertagus and
Clavocerithium. Monographs of Marine Mollusca, 1: 1–130.
Houbrick RS (1985) Genus Clypeomorus Jousseaume (Cerithiidae: Prosobranchia). Smithsonian Contributions to
Zoology, 403: 1–131.
Houbrick RS (1991) Systematic review and functional morphology of the mangrove snails Terebralia and
Telescopium (Potamididae; Prosobranchia). Malacologia, 33: 289–338.
Houbrick RS (1992) Monograph of the genus Cerithium Bruguiére in the Indo-Pacific (Cerithiidae: Prosobranchia).
Smithsonian Contributions to Zoology, 510: 1–211.
Hualkasin W, Tongchuai W, Chotigeat W & Phongdara A (2008) Phylogeography of ivory shell ( Babylonia
areolata) in the Gulf of Thailand revealed by COI gene structure and differentiation of shell color by ITS1
DNA. Songklanakarin Journal of Science and Technology, 30: 141–146.
Huang ZG & Morton B (1983) Mytilopsis sallei (Mollusca: Dreissenoidea) established in Victoria Harbour, Hong
Kong. Malacological Review, 16: 99–100.
Huber M (2010) Compendium of Bivalves. ConchBooks, Hackenheim, Germany, 901 pp.
Huber M (2015) Compendium of Bivalves 2. ConchBooks, Harxheim, Germany, 907 pp.
Huber M, Langleit A & Kreipl K (2015) Tellinidae. In: Huber M. Compendium of Bivalves 2. ConchBooks, Harxheim,
Germany, pp. 167–297, 564–746.
Hunon C, Hoarau A & Robin R (2009) Olividae (Mollusca, Gastropoda). A Complete Survey of the Recent Species of
the Genus Oliva. ConchBooks, Hackenheim; Xenophora, Paris, 256 pp.
Hylleberg J (1994) Indo-Pacific cockles (Bivalvia: Cardiidae). Part I. Generic diagnoses and an overview of species
with mention of taxonomic problems encountered in Thailand. Phuket Marine Biologic al Center Special
Publication, 13: 113–136.
Hylleberg J (2004) Lexical approach to Cardiacea. Records of taxa. Illustrated and annotated records of living and
fossil shells, with emphasis on the families Cardiidae and Lymnocardiidae (Mollusca: Bivalvia). Part 2: A–M and
Part 3: N–Z. Phuket Marine Biological Center Special Publication, 30: 352–939.
Hylleberg J (2009) Cardiidae (Mollusca: Bivalvia) in the collection of Statens naturhistoriske Museum, previously
the Zoological Museum, University of Copenhagen (ZMUC). Annotated and revised. Steenstrupia, 31: 103–324.
172
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Hylleberg J (2013) Phuket Marine Biological Center (PMBC) and Thai–Danish cooperation summarized by a former
Danish advisor, with emphasis on the early history. Phuket Marine Biological Center Special Publication, 32:
13–30.
Hylleberg J (2015) Revision and morphometric characteristics of the Vasticardium fidele group (Bivalvia:
Cardiidae) in the Indo-Pacific with description of three new species. NOVAPEX, 16: 81–94.
Hylleberg J & Aungtonya C (2013) Biodiversity survey in Chalong Bay and surrounding areas, east coast of Phuket
Island. Phuket Marine Biological Center Special Publication, 32: 81–112.
Hylleberg J & Kilburn RN (2003) Marine molluscs of Vietnam: Annotations, voucher material and species in need
of verification. In: Hylleberg J (ed.) Tropical Marine Mollusc Programme (TMMP). Tenth International Congress
& Workshop, Hanoi & Haiphong, Vietnam. Phuket Marine Biological Center Special Publication, 28: 1–300.
Hylleberg J & Knudsen J (2001) Translations into English of Lorenz Spengler’s papers on bivalves (1783–1798).
Part 4, 1792. The genera Pholas and Teredo. Phuket Marine Biological Center Research Bulletin, 25: 557–570.
Hylleberg J & Nateewathana A (1991a) Redescription of Idiosepius pygmaeus Steenstrup, 1881 (Cephalopoda:
Idiosepiidae) with mention of additional morphological characters. Phuket Marine Biological Center Research
Bulletin, 55: 33–42.
Hylleberg J & Nateewathana A (1991b) Morphology, internal anatomy, and biometrics of the cephalopod Idiosepius
biserialis Voss, 1962. A new record for the Andaman Sea. Phuket Marine Biological Center Research Bulletin,
56: 1–9.
Hylleberg J & Nateewathana A (1992) Description of Chicoreus ramosus shells, with notes on Chicoreus torrefactus.
Phuket Marine Biological Center Special Publication, 10: 109–112.
Iredale T (1936) Australian molluscan notes. No. 2. Records of the Australian Museum, 19: 267–340.
Iwasaki K & Yamamoto H (2014) Recruitment and population structure of the non -indigenous brackish-water
mytilid Xenostrobus securis (Lamarck, 1819) in the Kino River, Japan. Aquatic Invasions, 9: 479–487.
Jarayabhand P & Paphavasit N (1996) A review of the culture of tropical abalone with special reference to Thailand.
Aquaculture, 140: 159–168.
Jarayabhand P, Yom-La R & Popongviwat A (1998) Karyotypes of marine molluscs in the family Haliotidae found
in Thailand. Journal of Shellfish Research, 17: 761–764.
Jayachandran PR, Aneesh BP, Oliver PG, Philomina J, Jima M, Harikrishnan K & Nandan SB (2019) First record of
the alien invasive biofouling mussel Mytella strigata (Hanley, 1843) (Mollusca: Mytilidae) from Indian waters.
BioInvasions Records, 8: 828–837.
Jensen KR (1989) A new species of Cylindrobulla from Phuket, Thailand with a discussion of the systematic
affiliation of the genus. Phuket Marine Biological Center Research Bulletin, 52: 1–11.
Jensen KR (1992) Anatomy of some Indo-Pacific Elysiidae (Opisthobranchia: Sacoglossa: Ascoglossa), with a
discussion of the generic divisions and phylogeny. Journal of Molluscan Studies, 58: 257 –296.
Jensen KR (1996) Phylogenetic systematics and classification of the Sacoglossa (Mollusca, Gastropoda,
Opisthobranchia). Philosophical Transactions of the Royal Society of London B, 351: 91 –122.
Jensen KR (1998) Anatomy of some opisthobranch molluscs from Phuket, Thailand with a list of Opisthobranchia
recorded from Phuket. Phuket Marine Biological Center Special Publication, 18: 243–262.
Jensen KR (2007) The opisthobranch molluscs collected during the Thai-Danish Bioshelf project in the Andaman
Sea. Phuket Marine Biological Center Research Bulletin, 68: 21–30.
Jensen KR & Knudsen J (2001) 100 years anniversary for the first Danish marine biological expedition to Thailand.
Phuket Marine Biological Center Special Publication, 25: 517–525.
Jensen KR, Kohnert P, Bendell B & Schrödl M (2014) Life on a leaf: 3D–reconstruction and description of a new
limapontiid sacoglossan (Gastropoda: Heterobranchia: ‘Opisthobranchia’) living on the seagrass Halophila ovalis.
Journal of Molluscan Studies, 80: 624–641.
Jereb P & Roper CFE (eds.) (2005) Cephalopods of the world. An annotated and illustrated catalogue of cephalopod
species known to date. Volume 1. Chambered nautiluses and sepioids. FAO Species Catalogue for Fishery
Purposes, 1: 1–262.
Jereb P & Roper CFE (eds.) (2010) Cephalopods of the world. An annotated and illustrated catalogue of cephalopod
species known to date. Volume 2. Myopsid and oegopsid squids. FAO Species Catalogue for Fishery Purposes,
4: 1-605.
Jereb P & Roper CFE (eds.) (2014) Cephalopods of the world. An annotated and illustrated catalogue of cephalopod
species known to date. Volume 2. Myopsid and oegopsid squids. FAO Species Catalogue for Fishery Purposes,
2: 1–605.
Jereb P, Roper CFE, Norman MD & Finn JK (eds.) (2014) Cephalopods of the world. An annotated and illustrated
catalogue of cephalopod species known to date. Volume 3. Octopods and vampire squids. FAO Species
Catalogue for Fishery Purposes, 3: 1–352.
Jespersen Å, Lützen J & Nielsen C (2004) On three species and two new genera (Montacutella and Brachiomya) of
galeommatoid bivalves from the irregular sea urchin Brissus latecarinatus with emphasis on their reproduction.
Zoologischer Anzeiger, 243: 3–19.
Jindalikit J & Sereeruk K (2005) Distribution and spawning grounds of cuttlefish in the upper Gulf of Thailand.
Phuket Marine Biological Center Research Bulletin, 66: 275–282.
173
The Marine and Estuarine Molluscs of Thailand
Johnson CR & Chapman RDO (2007) Seaweed invasions: Introduction and scope. Botanica Marina, 50: 321–325.
Jung P & Abbott RT (1967) The genus Terebellum (Gastropoda: Strombidae). Indo-Pacific Mollusca, 1: 445–454.
Kaas P, van Belle RA & Strack HL (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6.
Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1–5. Koninklijke
Brill NV, Leiden, 463 pp.
Kalyanasundaram N (1975) Studies on the biology of Mytilopsis sallei (Recluz), an important marine fouling
mollusc. Bulletin of the Department of Marine Science, University of Cochin, 7: 658–693.
Kano Y & Kase T (2003) Systematics of the Neritilia rubida complex (Gastropoda: Neritiliidae): Three amphidromous
species with overlapping distributions in the Indo-Pacific. Journal of Molluscan Studies, 69: 273–284.
Kasetsart University (2000) Andaman Coastal Resources Park Honorary. Kasetsart University Research and
Development Institute Aksornsiam Printing, Bangkok, pp. 55–59.
Kasetsart University (2006) Biodiversity survey and information system project: Final report. Volume 3. Marine
and Coastal Biodiversity and Ecology. Forest Research Center, Faculty of Forestry, Kasetsart University,
Bangkok, pp. 11–117.
Kasetsart University (2010) Project on Biodiversity Inventory and Information Management in the Biodiversity
Hotspot of Ranong, Phangnga, Krabi and Phuket Provinces. Final Report Presented to the Office of Natural
Resources and Environmental Policy and Planning, Volume 5: Coastal and Marine Ecosystems. Faculty of
Fisheries, Kasetsart University, Bangkok, 1013 pp.
Kasetsart University Research and Development Institute (KURDI), Chulabhorn Research Institute & Third Naval
Area Command Royal Thai Fleet Royal Thai Navy (2004) Chulabhorn Marine Park Project 36 Handbook of
Marine Fauna and Flora in Coral Reefs, Surin and Similan Islands. Phuket Printing, Phuket, 109 pp.
Keawtawee T, Songsangjinda P, Sangnoi Y & Uppabullung A (2018) The current situation and environmental
conditions of green mussel farming in the Gulf of Thailand. IOP Conference Series: Earth and Environmental
Science, 137: 012093. https://doi.org/10.1088/1755-1315/137/1/012093 (Accessed 25 May 2019).
Khogkhaoa C, Hayashizakib K, Tuntiprapasa P & Prathepa A (2017) Changes in seagrass communities along the
runoff gradient of the Trang River, Thailand. ScienceAsia, 43: 339–346.
Kilburn RN, Fedosov AE & Olivera BM (2012) Revision of the genus Turris Batsch, 1789 (Gastropoda: Conoidea:
Turridae) with the description of six new species. Zootaxa, 3244: 1–58.
Kilburn R & Hylleberg J (1998) Gastropods and bivalves from Thailand; with taxonomic notes and records. Phuket
Marine Biological Center Special Publication, 18: 311–316.
Kittiwattanawong K, Nugranad J & Srisawat T (2001) High genetic divergence of Tridacna squamosa living at the
west and the east coasts of Thailand. Phuket Marine Biological Center Special Publication, 25: 343–347.
Kleeman KH (1983) Catalogue of recent and fossil Lithophaga (Bivalvia). Journal of Molluscan Studies, Supplement
12: 1–46.
Kleeman KH (1984) Lithophaga (Bivalvia) from dead coral from the Great Barrier Reef, Australia. Journal of
Molluscan Studies, 50: 192–230.
Kleeman K (2007) Catalogue and bibliography of recent and fossil Botula (Bivalvia: Mytilidae). Zootaxa, 1508: 1–48.
Klinbunga S, Khamnamtong B, Puanglarp N, Jarayabhand P, Yoosukh W & Menasveta P (2005) Molecular taxonomy
of cupped oysters (Crassostrea, Saccostrea, and Striostrea) in Thailand based on COI, 16S, and 18S rDNA
polymorphism. Marine Biotechnology, 7: 306–317.
Knudsen J & Jensen KR (2001) Translations into English of Lorenz Spengler’s papers on Bivalvia (1783–1798).
Part 3. 1788: Pholas siamensis with notes on the type specimen. The oldest marine bivalve with a type locality
in the Gulf of Thailand. Phuket Marine Biological Center Special Publication, 25: 547–556.
Kohama T, Montani S, Kajiwara Y & Yamada M (2001) Population dynamics of sessile bivalves Mytilus
galloprovincialis and Xenostrobus securis in hyper eutrophicated bay, Japan. Bulletin of the Japanese Society
for Fisheries Science, 67: 664–671.
Kohn AJ (1978) The Conidae (Mollusca: Gastropoda) of India. Journal of Natural History, 12: 295–335.
Kool HH (2003) Nassarius biendongensis spec. nov. from the South China Sea (Gastropoda, Prosobranchia,
Nassariidae). Basteria, 67: 43–46.
Kool HH (2006) Nassarius cadeei n. sp., a new species from South East Asia (Gastropoda: Nassariidae). Miscellanea
Malacologica, 2: 21–24.
Kool HH (2007) Nassarius garuda n. sp., a new deepwater species from the Indonesian Tanimbar and Kai Islands
and a review of the species N. crematus (Hinds, 1844), N. euglyptus (Sowerby III, 1914) and N. siquijorensis (A.
Adams, 1852) (Gastropoda: Buccinoidea: Nassariidae). Miscellanea Malacologica, 2: 87–92.
Kool HH (2008) On the identity of Nassarius vitiensis (Hombron & Jacquinot in Rousseau, 1854), N. rufus
(Dunker, 1847), N. kiiensis Kira, 1954, and N. caelatus (A. Adams, 1852) (Gastropoda: Nassariidae). Miscellanea
Malacologica, 3: 49–59.
Kool HH (2013a) Nassarius madurensis n. sp., a new nassariid species from south of Madura, Indonesia
(Gastropoda: Nassariidae). Miscellanea Malacologica, 6: 35–37.
Kool HH (2013b) Re-evaluation of several species in the family Nassariidae, considered synonyms of Nassarius
reeveanus (Dunker, 1847) by Cernohorsky (1984) and others (Gastropoda: Neogastropoda). Miscellanea
Malacologica, 6: 39–45.
174
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Kostina EE, Tsurpalo AP & Gulbin VV (2016) The species composition and distribution of microbenthic communities
in the intertidal zone of Vietnam. In: Adrianov AV & Lutataenko KA (eds.) Biodiversity of the Western Part of
the South China Sea. Dalnauka, Vladivostock, pp. 139–314.
Kosuge S, Roussy PH & Muangman P-P (1998) Report on the fauna of Thailand (1) with the description of a new
species (Columbellidae and Buccinidae). Bulletin of the Institute of Malacology of Tokyo, 3: 75–76.
Krijnen C, Delsaerdt A, Severijns N, Verhaeghe M & Vink R (2011) On the identity of Nerita undulata Gmelin, 1791,
previously known as Nerita grayana Récluz, 1843 (Gastropoda: Neritidae). Gloria Maris, 49: 125–140.
Kreipl K (1997) Recent Cassidae. Verlag Christa Hemmen, Wiesbaden, 151 pp.
Kronenberg GC (1998) Revision of Euprotomus Gill, 1870: 1. The systematic position of Strombus listen Gray, 1852
(Gastropoda: Prosobranchia: Strombidae). Vita Marina, 45: 1–6.
Kronenberg GC & Dekker H (1998) A new species of Cotonopsis Olsson, 1942, from an unexpected locality
(Gastropoda: Prosobranchia: Columbellidae). Vita Marina, 45: 11–16.
Kronenberg GC & Dekker H (1999) Cotonopsis vanwalleghemi Kronenberg & Dekker, 1998, a junior synonym of
Strombina phuketensis Kosuge, Roussy & Muangman, 1998, with some notes on the generic position and colour
pattern (Gastropoda Prosobranchia: Columbellidae). Vita Marina, 46: 69–72.
Kronenberg GC & Dharma B (2005) New distributional records for four species of Stromboidea (Mollusca:
Gastropoda) from Australasia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory,
21: 47–51.
Kronenberg GC, Liverani V & Dekker H (2009) On the identity of Strombus coniformis Sowerby II, 1842 (Gastropoda,
Strombidae), with additional notes on its distribution. Journal of Conchology, 39: 659–668.
Kurozumi T, Kosuge T & Tsuchiya M (1989) List of shallow-water marine molluscs in the Sichang Island, the Gulf
of Thailand. Galaxea, 8: 295–310.
Labbé A (1935) Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. gen. nov. sp. Bulletin de la Société
Zoologique de France, 60: 312–317.
Lam K & Morton B (2004) The oysters of Hong Kong (Bivalvia: Ostreidae and Gryphaeidae). The Raffles Bulletin of
Zoology, 52: 11–28.
Lam K & Morton B (2009) Oysters (Bivalvia: Ostreidae and Gryphaeidae) recorded from Malaysia and Singapore.
The Raffles Bulletin of Zoology, 57: 481–494.
Lamy E (1936) Révision des Mytilidae vivants du Muséum National d’Histoire Naturelle de Paris. Journal de
Conchyliologie, 80: 66–102; 107–198; 229–295.
Lamprell K & Whitehead T (1990) Eight new marine bivalves from Australia (Mollusca, Bivalvia). Journal of the
Malacological Society of Australasia, 11: 33–52.
LCP (2018) Laem Chabang Port Statistics. https://www.laemchabangportnew.com (Accessed 15 November 2018).
Leloux J & Wesselingh FP (2009) Types of Cenozoic Mollusca from Java in the Martin Collection of Naturalis.
National Museum of Natural History Naturalis, Leiden, NHM Technical Bulletin, 11: 1–765.
Lesueur CA (1821) Descriptions of several new species of cuttlefish. Journal of the Academy of Natural Sciences of
Philadelphia, 2: 86–101.
Li BQ, Li XZ & Kilburn RN (2010) Report on the genus Tritonoturris s. l., (Gastropoda: Conoidea) from the south
China Sea. Journal of Conchology, 40: 193–199.
Lim CF & Wee VTH (1992) Southeast Asian Conus: A Seashell Book. Seaconus Pte Ltd, Bangkok, 100 pp.
Lim JY, Tay TS, Lim CS, Lee SSC, Teo SLM & Tan KS (2018) Mytella strigata (Bivalvia: Mytilidae): An alien mussel
recently introduced to Singapore and spreading rapidly. Molluscan Research, 38: 170–186.
Lorenz F (1999) A western subspecies of Lyncina leucodon (Broderip 1832) (Mollusca: Gastropoda: Cypraeidae).
Schriften zur Malakozoologie, 13: 16–18.
Lorenz F (2018) Cowries. A Guide to the Gastropod Family Cypraeidae. ConchBooks, Hackenheim, 715 pp.
Lorenz F & Fehse D (2009) The Living Ovulidae. A Manual of the Families of Allied Cowries: Ovulidae,
Pediculariidae and Eocypraeidae. ConchBooks, Hackenheim, 651 pp.
Lützen J & Nielsen K (2005) Galeommatid bivalves from Phuket, Thailand. Zoological Journal of the Linnean
Society, 144: 261–308.
Lynge H (1909) Marine Lamellibranchia. The Danish Expedition to Siam 1899 –1900. Det Kongelige Danske
Videnskabernes Selskab Skrifter, Naturvidenskab og Mathematik Afdeling, 5: 99–299.
Malaquias MAE & Reid DG (2008) Systematic revision of the living species of Bullidae (Mollusca: Gastropoda:
Cephalaspidea), with a molecular phylogenetic analysis. Zoological Journal of the Linnean Society, 153:
453–543.
Man in 't Veld LA & Turck K de (1998) Contributions to the knowledge of Strombacea. 6. A revision of the subgenus
Laevistrombus Kira, 1955, including the description of a new species from the New Hebrides. Gloria Maris, 36:
73–107.
Matsukuma A & Habe T (1995) Systematic revision of living species of Meiocardia, Glossidae and Glossocardia,
Trapezidae (Bivalvia). In: Bouchet P (ed.) Résultats des Campagnes Musorstom, Volume 14. Mémoires du
Muséum national d’Histoire naturelle, 167: 75–106.
Mediodia DP, De Leon SMS, Anasco NC, Baylon CC (2017) Shell morphology and anatomy of the Philippine charru
mussel Mytella charruana (d’Orbigny, 1842). Asian Fisheries Science, 30: 185–194.
175
The Marine and Estuarine Molluscs of Thailand
Mehrotra R, Caballer Gutierrez M & Chavanich S (2017) On the genus Armina (Gastropoda: Heterobranchia:
Nudibranchia) in Thailand. Marine Biodiversity, 47: 549–559.
Mehrotra R & Scott CM (2017) Species inventory of sea slugs (Gastropoda: Heterobranchia) for Koh Tao, Thailand,
with 25 first records for Thai waters. Marine Biodiversity, 47: 549–550.
Meij SET Van Der, Moolenbeek RG & Dekker H (2010) The Lioconcha castrensis species group (Bivalvia: Veneridae),
with the description of two new species. Molluscan Research, 30: 117–124.
Merle D, Garrigues B & Pointier J-P (2011) Fossil and Recent Muricidae of the World. Part Muricinae. ConchBooks,
Hackenheim, 648 pp.
Merle D & Pacaud J-M (2003) New species of Eocithara Fischer, 1883 (Mollusca, Gastropoda, Harpidae) from the
Early Paleogene with phylogenetic analysis of the Harpidae. Geodiversitas, 26: 61–87.
Meyer E, Nilkerd B, Glover EA & Taylor JD (2008) Ecological importance of chemoautotrophic lucinid bivalves in a
peri-mangrove community in eastern Thailand. In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE
(eds.) Molluscs of Eastern Thailand. Proceedings of the International Marine Bivalve Workshop, Chanthaburi,
Thailand, August–September 2005, with contributions on other molluscan groups. Raffles Bulletin of Zoology,
Supplement 18: 41–55.
Middelfart P (1996) Egg capsules and early development of ten muricid gastropods from Thai waters. Phuket
Marine Biological Center Special Publication, 16: 103–130.
Middelfart P (1997) An illustrated checklist of Muricidae (Gastropoda: Prosobranchia) from the Andaman Sea,
Thailand. Phuket Marine Biological Center Special Publication, 17: 349–388.
Mikkelsen PS (1985) A comparison of two Florida populations of the coquina clam, Donax variabilis Say, 1822
(Bivalvia: Donacidae). 2. Growth rates. The Veliger, 27: 308–311.
Molnar JL, Gamboa RL, Revenga C & Spalding MD (2008) Assessing the global threat of invasive species to marine
biodiversity. Frontiers in Ecology and the Environment, 6: 485–492.
Monsecour K & Köhler F (2006) Annotated list of columbellid types held in the Malacological Collection of t he
Museum für Naturkunde, Berlin (Mollusca, Caenogastropoda, Columbellidae). Mitteilungen aus dem Museum
für Naturkunde in Berlin, Zoologische Reihe, 82: 282–306.
Monsecour K & Monsecour D (2000) A new Angaria from Thailand (Mollusca: Gastropoda). Gloria Maris, 38: 90–93.
Moolenbeek RG (1994) The Orbitestellidae (Gastropoda: Heterobranchia) of the Sultanate of Oman with
description of a new genus and two new species. Apex, 9: 5–10.
Moolenbeek RG (2009a) A common mytilid from the Sunda Shelf new to science Musculus panhai n. sp. (Bivalvia:
Mytilidae). Miscellanea Malacologica, 3: 113–116.
Moolenbeek RG (2009b) Hoenselaaria, a new genus with the description of a new species (Gastropoda: Eulimidae)
from the Indo-Pacific. Miscellanea Malacologica, 3: 71–75.
Morlet L (1889) Catalogue de coquilles recueillies, par M. Pavie, dans le Cambodge et le Royaume de Siam, et
description des espèces species nouvelles (1). Journal de Conchyliologie, 37: 121–199.
Morris S & Purchon RD (1981) The marine shelled Mollusca of West Malaysia and Singapore: Part 3, Bivalvia.
Journal of Molluscan Studies, 47: 322–327.
Morton B (1980) Mytilopsis sallei recorded from Hong Kong: An introduction by Vietnamese refugees? Malacological
Review, 13: 90–92.
Morton B (1989) Life-history characteristics and sexual strategy of Mytilopsis sallei (Bivalvia: Dreissenacea),
introduced into Hong Kong. Journal Zoology of London, 19: 469–485.
Morton B (2009) The functional morphology of Sinonovacula constricta with a discussion on the taxonomic status
of the Novaculininae (Bivalvia). Journal of Zoology, 202: 299–325.
Morton B, Lam K & Slack-Smith S (2003) First report of the European flat oyster Ostrea edulis, identified
genetically, from Oyster Harbour, Albany, south-western Western Australia. Molluscan Research, 23: 199–208.
Morton B & Leung KF (2015) Introduction of the alien Xenostrobus securis (Bivalvia: Mytilidae) into Hong Kong,
China: Interactions with and impacts upon native species and the earlier introduced Mytilopsis sallei (Bivalvia:
Dreissenidae). Marine Pollution Bulletin, 15: 134–142.
Morton B & Tan KS (2006) Brachidontes striatulus (Bivalvia: Mytilidae) introduced into Singapore. Raffles Bulletin
of Zoology, 54: 435–439.
MRGV (1984) Coastal Invertebrates of Victoria. Field Naturalists Club of Victoria Marine Research Group,
Melbourne, 170 pp.
Mynhardt G, Alejandrino A, Puslednik L, Corrales J & Serb JM (2014) Shell shape convergence masks biological
diversity in gliding scallops: Description of Ylistrum n. gen. (Pectinidae) from the Indo-Pacific Ocean. Journal of
Molluscan Studies, 80: 400–411.
Nabhitabhata J (2009) Checklist of Mollusca Fauna in Thailand. Office of Natural Resources and Environmental
Policy and Planning, Bangkok, 576 pp.
Nabhitabhata J & Mitrpanont T (2017) First photographic record of mimic octopus, Thaumoctopus mimicus
Norman and Hochberg, 2005 in the East Andaman Sea, East Indian Ocean. Phuket Marine Biological Center
Research Bulletin, 74: 45–50.
Nabhitabhata J, Nateewathana A & Sukhsangchan (2009a) Cephalopods. In: Nabhitabhata J. Checklist of Mollusca
Fauna in Thailand. Office of Natural Resources and Environmental Policy and Planning, Bangkok, pp. 256–277.
176
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Nabhitabhata J, Nilaphat P, Promboon P, Jaroongpattananon C, Nilaphat G & Reunreng A (2005) Performance of
simple large-scale cephalopod culture system in Thailand. Phuket Marine Biological Center Research Bulletin,
66: 337–350.
Nabhitabhata J & Sukhsangchan C (2007) New photographic record of “mimic octopus” in the Gulf of Thailand.
Phuket Marine Biological Center Research Bulletin, 68: 31–34.
Nabhitabhata J, Sukhsangchan C & Wongkamhaeng K (2009b) First record of two pelagic octopods, Argonauta argo
and Tremoctopus violaceus cf. gracilis from the Andaman Sea, Thailand. Vie et Milieu – Life and Environment,
59: 39–45.
Nabhitabhata J & Suwanamala J (2008) Reproductive behaviour and cross-mating of two closely related pygmy
squids: Idiosepius biserialis and Idiosepius thailandicus (Cephalopoda: Idiosepiidae). Journal of Marine
Biological Association of United Kingdom, 88: 987–993.
Nabhitabhata J, Tuanapaya S, Tongtherm K, Promdam R & Aungtonya C (2019) First record of seven neritic
cephalopods (Sepiidae, Loliginidae, Octopodidae) new to Thai waters with taxonomic list of Thai teuthofauna.
Phuket Marine Biological Center Research Bulletin, 76: 45–66.
Nabhitabhata J & Wongkamhaeng K (2012) New record of live specimens of emperor nautilus, Nautilus pompilius
from the East Andaman Sea, Thai Waters. Vie et Milieu, 62: 97–104.
Nateewathana A (1992) Taxonomic studies on loliginid squids (Cephalopoda: Loliginidae) from the Andaman Sea
coast of Thailand. Phuket Marine Biological Center Research Bulletin, 57: 1–40.
Nateewathana A (1995a) Taxonomic account of commercial and edible molluscs, excluding cephalopods, of
Thailand. Phuket Marine Biological Center Special Publication, 15: 93–116.
Nateewathana A (1995b) New record of oceanic squids from Thai waters, the Andaman Sea. Phuket Marine
Biological Center Research Bulletin, 60: 1–19.
Nateewathana A (1996) The Sepiidae (Cephalopoda) of the Andaman Sea, Thailand. Phuket Marine Biological
Center Special Publication, 16: 145–176.
Nateewathana A (1997a) The octopod fauna (Cephalopoda: Octopoda) of the Andaman Sea, Thailand. Phuket
Marine Biological Center Special Publication, 17: 407–452.
Nateewathana A (1997b) Two species of oceanic squids from the Andaman Sea, Indian Ocean. Phuket Marine
Biological Center Special Publication, 17: 453–464.
Nateewathana A (1997c) The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description of
Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication, 17: 465–481.
Nateewathana A (1998) A new record of cuttlefish Metasepia tullbergi (Apellof, 1886) (Sepiidae: Cephalopoda),
Gulf of Thailand. Phuket Marine Biological Center Special Publication, 18: 323–330.
Nateewathana A (2008) The Sepiidae (Cephalopoda) of Thailand. Phuket Marine Biological Research Bulletin, 69:
25–41.
Nateewathana A & Hylleberg J (1989) First record of oceanic squid, Thysanoteuthis rhombus Troschel, 1857
(Cephalopoda: Teuthoidea) in Thai waters. The Natural History Bulletin of the Siam Society, 37: 227–233.
Nateewathana A, Nabhitabhata J & Nilaphat P (2001) A new record of a bobtail squid, Euprymna hyllebergi
Nateewathana, 1997, in the Gulf of Thailand. Phuket Marine Biological Center Special Publication, 25: 501–506.
Nateewathana A & Norman M (1999) On three new species of ocellate octopuses (Cephalopoda: Octopoda) from
Thai waters. Phuket Marine Biological Center Special Publication, 19: 445–462.
Nateewathana A, Tantichodok P, Bussarawich S & Sirivejabandhu R (1981) Marine organisms in the Reference
Collection. Phuket Marine Biological Center Research Bulletin, 28: 43–86.
Negri MP, Sanfilippo R, Basso D, Rosso A & Geronimo SID (2014) Molluscan associations from the Pak Phanang
Bay (SW Gulf of Thailand) as a record of natural and anthropogenic changes. Continental Shelf Research, 84:
204–218.
Ngamprasertkit S (2018) Thailand Seafood Report. GAIN Report Number: TH8067. USDA Foreign Agricultural
Service, Global Agricultural Information Network, 19 pp.
Nielsen C (1976a) Notes on boring bivalves from Phuket, Thailand. Ophelia, 15: 141–148.
Nielsen C (1976b) An illustrated checklist of bivalves from PMBC beach with a reef-flat at Phuket, Thailand. Phuket
Marine Biological Center Research Bulletin, 9: 1–26.
NIMPCG (2009a) Marine Pests Monitoring Manual: Version 1. National Introduced Marine Pests Coordination
Group, Department of Agriculture, Fisheries and Forestry, Canberra, 142 pp.
NIMPCG (2009b) Australian Marine Pests Monitoring Guidelines: Version 1. National Introduced Marine Pests
Coordination Group, Department of Agriculture, Fisheries and Forestry, Canberra, 55 pp.
NIMPIS (2017) National Introduced Marine Pests Information System. http://data.daff.gov.au/marinepests/
(Accessed 25 December 2017).
Norman MD & Hochberg FG (2005) The current state of octopus taxonomy. Phuket Marine Biological Center
Research Bulletin, 66: 127–154.
Norman MD & Lu CC (1997) Redescription of the southern dumpling squid Euprymna tasmanica and a revision of
the genus Euprymna (Cephalopoda: Sepiolidae). Journal of the Marine Biological Association of the United
Kingdom, 77: 1109–1137.
177
The Marine and Estuarine Molluscs of Thailand
Norman MD, Nabhitabhata J & Lu CC (2016) An updated checklist of the cephalopods of the South China Sea. The
Raffles Bulletin of Zoology, Supplement 34: 566–592.
OBIS (2020) Ocean Biodiversity Information System. Intergovernmental Oceanographic Commission of UNESCO.
https://obis.org/ (Accessed 6 April 2020).
Ockelmann KW (1983) Descriptions of mytilid species and definition of the Dacrydiinae n. subfam. (Mytilacea Bivalvia). Ophelia, 22: 81–123.
ONEP (2007) Final Report: Biodiversity Survey and Information System of the Biodiversity Hotspots; Petburi and
Prachuap Khiri Khan Provinces. Office of Natural Resources and Environmental Policy and Planning, Bangkok,
1310 pp.
Office of Natural Resources (ONR) and Environmental Policy and Planning (EPP) (2006) Support Project for the
Strategic Plan for the Coastal Environment of the Andaman Sea. Faculty of Fisheries, Kasetsart University ,
Bangkok, pp. 5–249.
Okutani T (ed.) (2000) Marine Mollusks in Japan. Tokai University Press, Japan, 1375 pp.
Okutani T (2005) Past, present and future studies on cephalopod diversity in tropical west Pacific. Phuket Marine
Biological Center Research Bulletin, 66: 39–50.
Oliver PG, Hallan A, Jayachandran PR, Joseph P, Sanu VF & Nandan SB (2018) Taxonom y of myid bivalves from
fragmented brackishwater habitats in India, with a description of a new genus Indosphenia (Myida, Myoidea,
Myidae). ZooKeys, 799: 21–46.
Omar WBW & Kassim Z (2015) Biodiversity of mollusc in Setiu Wetlands by using molecular markers for
characterization and conservation. In: Mohamad F (ed.) Setiu Wetlands: Species, Ecosystems, and Livelihoods.
Universiti Malaysia Terengganu, Malaysia, pp. 27–37.
Oskars TR & Malaquias MAE (2019) A molecular phylogeny of the Indo-West Pacific species of Haloa sensu lato
gastropods (Cephalaspidea: Haminoeidae): Tethyan vicariance, generic diversity, and ecological specialization.
Molecular Phylogenetics and Evolution, 139: 106557.
Oskars TR, Too CC, Rees D, Mikkelsen PM, Willassen E & Malaquias MAE (2019) A molecular phylogeny of the
gastropod family Haminoeidae sensu lato (Heterobranchia: Cephalaspidea): A generic revision. Invertebrate
Systematics, 33: 426–472.
Ozawa T, Wei Y, Cuizhang F, Claremont M, Smith L & Reid DG (2015) Allopatry and overlap in a clade of snails
from mangroves and mud flats in the Indo-West Pacific and Mediterranean (Gastropoda: Potamididae:
Cerithideopsilla). Biological Journal of the Linnean Society, 114: 212–228.
PAT (2020) Annual Report. Port Authority of Thailand. http://www.port.co.th/cs/internet/internet/Annual_
Report.html (Accessed 5 May 2020).
Patamakanthin S (2001) A new Turbo species from Thailand, Andaman Sea (Gastropoda: Turbinidae). Of Sea and
Shore, 23: 192–195.
Patamakanthin S (2002) Description of a small new Haliotis species from Olango Island, Philippines. Of Sea and
Shore, 25: 4–8.
Perrone AS (2001) A new species of Nudibranchia of the genus Aldisa Bergh (Gastropoda, Opisthobranchia) from
Thailand. Basteria, 65: 105–116.
Phongsuwan N, Chankong A, Yamarunpatthana C, Chansang H, Boonprakob R, Petchkum nerd P, Thongtham N,
Paokantha S, Chanmethakul T, Panchaiyapoome P & Bundit O-A (2013) Status and changing patterns on coral
reefs in Thailand during the last two decades. Deep-Sea Research II, 96: 19–24.
Ponder WF (1983) A revision of the recent Xenophoridae of the world and of the Australian fossil species
(Mollusca, Gastropoda). Australian Museum Memoir, 17: 1–126 [Appendix by Ponder WF & Cooper J, pp. 67–74].
Ponder WF (1984) A review of the genera of the Rissoidae (Mollusca: Mesogastropoda: Rissoacea). Records of the
Australian Museum, Supplement 4: 1–221.
Ponder WF, Fukuda H & Hallan A (2014) A review of the family Clenchiellidae (Mollusca: Caenogastropoda:
Truncatelloidea). Zootaxa, 3872: 101–153.
Ponder WF & Keyzer R de (1992) A revision of the genus Diala (Gastropoda: Cerithioidea: Dialidae). Invertebrate
Taxonomy, 6: 1019–1075.
Ponder WF & Vokes EH (1988) A revision of the Indo-West Pacific fossil and recent species of Murex s.s. and
Haustellum (Mollusca: Gastropoda: Muricidae). Records of the Australian Museum, Supplement 8: 1–160.
Poorten JJ ter (1997) Acrosterigma sewelli (Prashad, 1932), a valid species from the central Indo-Pacific, compared
with Acrosterigma flava (Linnaeus, 1758) (Bivalvia, Cardiidae). Basteria, 61: 33–39.
Poorten JJ ter (2007) Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 13. Mollusca,
Bivalvia, Cardiidae. Zoologische Verhandelingen Leiden, 81: 259–301.
Poorten JJ ter (2009) The Cardiidae of the Panglao Marine Biodiversity Project 2004 and the Panglao 2005 Deep–
Sea Cruise with descriptions of four new species (Bivalvia). Vita Malacologica, 8: 9–96.
Poorten JJ ter (2013) Revision of the recent species of the genus Nemocardium Meek, 1876 (Bivalvia, Cardiidae),
with the descriptions of three new species. Basteria, 77: 45–73.
Poorten JJ ter & Gemert LJ van (2016) The genus Frigidocardium Habe, 1951 in the Red Sea (Bivalvia, Cardiidae).
Basteria, 80: 64–76.
Poppe GT (2017) Philippine Marine Mollusks. Volume V. ConchBooks, Hackenheim, 628 pp.
178
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Poppe GT & Groh K (2006) The Family Pectinidae. A Conchological Iconography. ConchBooks, Hackenheim, 401 pp.
Powell AWB (1964) The family Turridae in the Indo-Pacific. Part 1. The subfamily Turrinae. Indo-Pacific Mollusca,
1: 227–344.
Powell AWB (1969) The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo-Pacific
Mollusca, 2: 215–415.
Powell AWB (1973) The patellid limpets of the world (Patellidae). Indo-Pacific Mollusca, 3: 75–206.
Premcharoen S, Witirawat S & Tharapoom P (2016) Molluscan fauna in Bang Taboon Mangrove Estuary, Inner
Gulf of Thailand: Implications for conservation and sustainable use of coastal resources. MATEC Web of
Conferences, 60: 02003. https://doi.org/10.1051/matecconf/20166002003 (Accessed 13 November 2019).
Prezant RS, Sutcharit C, Chalermwat K, Kakhai N, Duangdee T & Dumrongrojwattana P (2008) Population study of
Laternula truncata (Bivalvia: Anomalodesmata: Laternulidae) in the mangrove sandflat of Kungkrabaen Bay,
Thailand with notes on L. cf. corrugata. In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.)
Molluscs of Eastern Thailand. Proceedings of the International Marine Bivalve Workshop, Chanthaburi, Thailand,
August–September 2005, with contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement
18: 57–73.
Printrakoon C, Wells FE & Chitramvong Y (2008) Distribution of molluscs in mangroves at six sites in the upper
Gulf of Thailand. In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand.
Proceedings of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005,
with contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 247–257.
Printrakoon C, Yeemin T & Valentich-Scott P (2016) Ecology of endolithic bivalve mollusks from Ko Chang,
Thailand. Zoological Studies, 55: 50.
Pumijumnong N (2014) Mangrove forests in Thailand. In: Faridah-Hanum I, Latiff A, Hakeem K & Ozturk M (eds.)
Mangrove Ecosystems of Asia. Springer, New York, pp. 62–79.
Ramakrishna & Dey A (2010) Annotated checklist of Indian marine molluscs (Cephalopoda, Bivalvia and
Scaphopoda) Part-1. Records of the Zoological Survey of India, Occasional Paper, 320: 1–357.
Raven H & Vermeulen JJ (2007) Notes on molluscs from NW Borneo and Singapore. 2. A synopsis of the Ellobiidae
(Gastropoda, Pulmonata). Vita Malacologica, 4: 29–62.
Raven JGM (2016) Notes on molluscs from NW Borneo. 3. A revision of Taurasia (Gastropoda, Muricidae) and
Preangeria (Gastropoda, Buccinidae) with comments on Semiricinula from NW Borneo. Vita Malacologica, 15:
77–104.
Raven JGM & Recourt P (2018) Notes on molluscs from NW Borneo. 4. Olivoidea (Gastropoda, Neogastropoda),
with the description of eight new species. Vita Malacologica, 17: 113–155.
Raybaudi Massilia L & Prati Musetti A (1995) A new species of Cassidae from Thailand: Phalium muangmani n.sp.
World Shells, 12: 14–17.
Rehder HA (1973) The family Harpidae of the world. Indo-Pacific Mollusca, 3: 207–274.
Reid DG (1986) The Littorinid Molluscs of Mangrove Forests in the Indo-Pacific Region. The Genus Littoraria.
British Museum of Natural History, London, 227 pp.
Reid DG (1989) Systematic revision of the recent species of Peasiella Nevill, 1885 (Gastropoda: Littorinidae), with
notes on the fossil species. The Nautilus, 103: 43–69.
Reid DG (1998) Additions and corrections to the taxonomy of the genus Peasiella Nevill, 1885 (Gastropoda:
Littorinidae). The Nautilus, 103: 6–33.
Reid DG (2001a) New data on the taxonomy and distribution of the genus Littoraria Griffith & Pidgeon, 1834
(Gastropoda: Littorinidae) in Indo-West Pacific mangrove forests. The Nautilus, 115: 115–139.
Reid DG (2001b) The genus Nodilittorina von Martens, 1897 (Gastropoda: Littorindae) in the Indo-Malayan region.
Phuket Marine Biological Center Special Publication, 25: 433–449.
Reid DG (2006) Comparative phylogeography and species boundaries in Echinolittorina snails in the central IndoWest Pacific. Journal of Biogeography, 33: 990–1006.
Reid DG (2014) The genus Cerithidea Swainson, 1840 (Gastropoda: Potamididae) in the Indo-West Pacific region.
Zootaxa, 3775: 1–60.
Reid DG & Claremont M (2014) The genus Cerithideopsis Thiele, 1929 (Gastropoda: Potamididae) in the Indo-West
Pacific region. Zootaxa, 3779: 61–80.
Reid DG & Ozawa T (2016) The genus Pirenella Gray, 1847 (= Cerithideopsilla Thiele, 1929) (Gastropoda:
Potamididae) in the Indo-West Pacific region and Mediterranean Sea. Zootaxa, 4076: 1–91.
Reid RGB (1990) Evoultionary implications of sulphide ozidizing symbioses in bivalves. In: Morton B (ed.)
Proceedings of a Memorial Symposium in Honour of Sir Maurice Yonge. Hong Kong University Press, Hong
Kong, pp. 127–140.
Rice MA, Rawson PD, Salinas AD & Rosario WR (2016) Identification and salinity tolerance of the western
hemisphere mussel Mytella charruana (d’Orbigny, 1842) in the Philippines. Journal of Shellfish Research, 35:
865–873.
Rilov G, Benayahu Y & Gasith A (2004) Prolonged lag in population outbreak of an invasive muss el: A shiftinghabitat model. Biological Invasions, 6: 347–364.
179
The Marine and Estuarine Molluscs of Thailand
Robba E, Di Geronimo I, Chaimanee N, Negri M & Sanfilippo R (2002) Holocene and recent shallow soft-bottom
mollusks from the northern Gulf of Thailand area: Bivalvia. Bollettino Malacologico, 38: 49–132.
Robba E, Di Geronimo I, Chaimanee N, Negri MP & Sanfilippo R (2003) Mollusk associations of the Northern Gulf
of Thailand and Holocene analogs from the Bangkok Clay. Proceedings of the First International Conference on
Paleontology of Southeast Asia. Mahasarakham University Journal, 22 (Special Issue 2003): 191–213.
Robba E, Di Geronimo I, Chaimanee N, Negri MP & Sanfilippo R (2004) Holocene and recent shallow soft-bottom
mollusks from the northern Gulf of Thailand area: Scaphopoda, Gastropoda, additions to Bivalvia. La Conchiglia,
35 (Supplement 309): 1–290.
Robba E, Di Geronimo I, Chaimanee N, Negri MP & Sanfilippo R (2007) Holocene and recent shallow soft-bottom
mollusks from the western Gulf of Thailand: Pak Phanang Bay and additions to Phetc haburi fauna. Bolletino
Malacologica, 43 (Speciale): 1–98.
Robertson R (1985) Archaeogastropod biology and the systematics of the genus Tricolia (Trochacea: Tricolidae)
in the Indo-West Pacific. Monographs of Marine Mollusca, 3: 1–103.
Röckel D, Korn W & Kohn AJ (1995) Manual of the Living Conidae. Volume 1: Indo-Pacific region. Verlag Christa
Hemmen, Wiesbaden, 517 pp.
Romero PE, Pfenninger M, Kano Y & Klussmann-Kolb A (2016) Molecular phylogeny of the Ellobiidae (Gastropoda:
Panpulmonata) supports independent terrestrial invasions. Molecular Phylogenetics and Evolution, 97: 43–54.
Roongratri M (1998) Re-examination of Loligo sumatrensis d’Orbigny, 1835 along the eastern coast of the Gulf of
Thailand. Thai Marine Fisheries Research Bulletin, 6: 11–16.
Rosenberg G (2010) Description of a new species of Prionovolva (Mollusca, Gastropoda, Ovulidae) from East Africa,
with reassessment of the composition of the genus. Proceedings of the Academy of Natural Sciences of
Philadelphia, 159: 39–66.
Rosenberg G, Moretzsohn F & García EF (2009) Gastropoda (Mollusca) of the Gulf of Mexico. In: Felder DL & Camp
DK (eds.) Gulf of Mexico. Origins, Waters, and Biota. Volume 1, Biodiversity. Texas A&M Press, College Station,
Texas, pp. 579–699.
Rosewater J (1961) The family Pinnidae in the Indo-Pacific. Indo-Pacific Mollusca, 1: 175–226.
Rosewater J (1965) The family Tridacnidae in the Indo-Pacific. Indo-Pacific Mollusca, 1: 347–396.
Rosewater J (1970) The family Littorinidae in the Indo-Pacific. Part I. The subfamily Littorininae. Indo-Pacific
Mollusca, 2: 417–506.
Rosewater J (1975) The marine commensal gastropod, Caledoniella montrouzieri (Prosobranchia: Hipponicacea)
in Thailand. The Nautilus, 89: 86.
Rottman M (1976) Euthecosomatous pteropods (Mollusca) in the Gulf of Thaila nd and the South China Sea:
Seasonal distribution and species associations. Naga Report, 4: 1–117.
Roussy HP (1987) A prize shell. Hawaiian Shell News, 35: 10.
Roussy H (1991a) Strombidae recorded in Thailand. Phuket Marine Biological Center Special Publication, 9: 20.
Roussy H (1991b) Family Pectinidae recorded – a checklist. Phuket Marine Biological Center Special Publication,
9: 21.
Rudman WB (2010) Sea Slug Forum. Australian Museum, Sydney. http://www.seaslugforum.net/ (Accessed 25
January 2019). [A number of entries were made over the years but are all listed as 2010 in this book.]
Ruhoff FA (1980) Index to the species of Mollusca introduced from 1850 to 1870. Smithsonian Contributions to
Zoology, 294: 1–640.
Sadeghi P & Loghmani M (2010) First record of Acanthopleura vaillantii (Mollusca: Polyplacophora) from Iran –
Chabahar Bay in the Oman Sea. Marine Biodiversity Records, 3: e7. https://doi.org/10.1017/S1755267209990984.
Saeedi H, Costello MJ & Cosel R von (2013) First report of anterior pallial tentacles in Solen dactylus (Bivalvia:
Solenidae) from the northern Persian Gulf, Iran. PLoS ONE, 8: e63487. https://doi.org/10.1371/journal.pone.
0063487.
Saito H (2017) Family Naticidae. In: Okutani T (ed.) Marine Mollusks in Japan. The Second Edition. Tokai University
Press, Japan, pp. 858–866.
Samakraman S, Williams GA & Ganmanee M (2010) Spatial and temporal variability of intertidal rocky shore
bivalves and gastropods in Sichang Island, east coast of Thailand. Publications of the Seto Marine Biological
Laboratory, Special Publication Series, 10: 35–46.
Sanpanich K (1998) An annotated check list of marine bivalves from Chonburi and Rayong Province, the east coast
of Thailand. Phuket Marine Biological Center Special Publication, 18: 297–306.
Sanpanich K (2011) Marine bivalves occurring on the east coast of the Gulf of Thailand. ScienceAsia, 37: 195–204.
Sanpanich K & Duangdee T (2009) The species diversity of marine molluscs at Nang-rong Beach, Jorake Island and
Juang Island, Chonburi Province. In: 4th Conference of RSPG Botanical and Biological Club. 20–23 October 2009,
Khao Khoew Open Zoo, Chonburi Province. Pp. 251–266.
Sanpanich K & Duangdee T (2013) The biodiversity of marine gastropods of Thailand in the late decade. Malaysian
Journal of Science, 32: 47–64.
Sanpanich K & Duangdee T (2018) A survey of marine mollusc diversity in the southern Mergui Archipelago,
Myanmar. Phuket Marine Biological Center Research Bulletin, 75: 45–60.
180
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Sanpanich K & Wells FE (2019) Mytella strigata (Hanley, 1843) emerging as an invasive marine threat in Southeast
Asia. BioInvasions Records, 8: 343–356.
Sanpanich K, Wells FE & Chitramvong Y (2004) Distribution of the family Littorinidae (Mollusca: Gastropoda) in
Thailand. Records of the Western Australian Museum, 22: 241–251.
Sartori AF, Printrakoon C, Mikkelsen PA & Bieler R (2008) Siphonal structure in the Veneridae (Bivalvia:
Heterodonta) with an assessment of its phylogenetic application and a review of venerids in the Gulf of Thailand.
In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand. Proceedings
of the International Marine Bivalve Workshop, Chanthaburi, Thaila nd, August–September 2005, with
contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 103–125.
Sato N, Awata S & Munehara H (2009) Seasonal occurrence and sexual maturation of Japanese pygmy squid
(Idiosepius paradoxus) at the northern limits of their distribution. ICES Journal of Marine Science, 66: 811–815.
Saurin E (1962) Pyramidellidae du Golfe de Thailande. Annales de la Faculté des Sciences (Saigon), 1961: 231–266.
Sawatna K, Champromarat A, Dumrongrojwattana P & Chiwapreecha B (2012) Diversity of marine microsnails on
Samaesarn Islands, Chonburi Province, Thailand. In: Proceedings of 50th Kasetsart University Annual Conference:
Science, January 31–February 2, 2012. Kasetsart University, Bangkok, Thailand, pp. 139–148.
Scarabino V (1995) Scaphopoda of the tropical Pacific and Indian Oceans, with description of 3 new genera and 42
new species. In: Bouchet P (ed.) Résultats des Campagnes Musorstom, Volume 14. Mémoires du Muséum
national d’Histoire naturelle, 167: 189–379.
Scarabino V (2008) New species and new records of scaphopods from New Caledonia. In: Héros V, Cowie RH &
Bouchet P (eds.) Tropical Deep-Sea Benthos 25. Mémoires du Muséum national d’Histoire naturelle, 196: 215–
268.
Schilder FA (1965) The cowrie fauna of Thailand. The Veliger, 8: 23–28.
Schultz PW & Huber M (2013) Revision of the worldwide recent Pinnidae and some remarks of fossil European
Pinnidae. Acta Conchyliorum, 13: 1–164.
Schwabe E (2006) Chitons collected during the Thai–Danish Bioshelf Surveys (1996–2000). Journal of the Zoological
Society of Wallacea, 2: 19–28.
Scott PH (1995) Infaunal bivalves of the Gulf of Thailand. Festivus, 27: 33–51.
Seidenfaden G, Smitinand T & Thorson G (1968) Report on the Fifth Thai–Danish Expedition 1966. The Natural
History Bulletin of the Siam Society, 22: 245–260.
Serène R (1937) Inventaire des Invertébrés marins de l’Indochine. Notes Institute Océanographique de la Indochine,
30: 1–83.
Signorelli JH & Printrakoon C (2020) The family Donacidae (Bivalvia: Tellinoidea) in Thai waters. Molluscan
Research, 40: 8–35.
Silina AV (2014) Ruditapes bruguieri (Bivalvia: Veneridae) at the coasts of Jeju Island. In: Sayenko EM & Lutaenko
KA. Abstracts of the Conference, Mollusks of the Eastern Asia and Adjacent Seas. October 6–8, 2014, Vladivostok,
Russia. Dalnauka, Vladivostok, pp. 94–97.
Simone LRL (2009) Anatomical description of Solen cf. exiguus Dunker from Thailand (Bivalvia: Solenidae). Archiv
für Molluskenkunde, 138: 113–122.
Simone LRL & Cunha CM (2008) Supplementary data for a recent revision of the genus Spinosipella (Bivalvia,
Septibranchia). Strombus, 15: 8–14.
Simone LRL & Simão do Amaral V (2008) Plicatulostrea, a new genus of Plicatulidae from Thailand (Bivalvia:
Pectinoidea). In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand.
Proceedings of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005,
with contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 127–135.
Simone LRL & Wilkinson S (2008) Comparative morphological study of some Tellinidae from Thailand (Bivalvia:
Tellinoidea). In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand.
Proceedings of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005,
with contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 151–190.
Smith BJ (1976) Revision of the recent species of the family Clavagellidae (Mollusca, Bivalvia). Journal of the
Malacological Society of Australia, 3: 187–207.
Sørensen AM (2012) Distribution, diversity and abundance patterns of intertidal benthic molluscs on rocky shores
(Phuket Island, Thailand). The Thai Natural History Museum Journal, 6: 113–128.
Spencer HG, Marshall BA & Willan RC (2009) Checklist of New Zealand living Mollusca. In: Gordon DP (ed.) New
Zealand Inventory of Biodiversity. Volume One. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia.
Canterbury University Press, Christchurch, pp. 196–219.
Sri-aroon P, Lohachit C & Harada M (2005) Brackish-water mollusks of Surat Thani Province, southern Thailand.
Southeast Asian Journal of Tropical Medicine and Public Health, 36 (Supplement 4): 180–188.
Sri-aroon P, Lohachit C, Harada M, Chusongsang P & Chusongsang Y (2006) Malacological survey in Phang-Nga
Province, southern Thailand, pre- and post-Indian Ocean tsunami. Southeast Asian Journal of Tropical Medicine
and Public Health, 36 (Supplement 3): 104–109.
Sterba GHW (2004) Olividae: A Collectors Guide (Mollusca, Neogastropoda). ConchBooks, Hackenheim, 172 pp.
181
The Marine and Estuarine Molluscs of Thailand
STRI (2019) Dendostrea frons (Linnaeus, 1758). Smithsonian Tropical Research Institute Bocas del Toro: Species
Database. https://biogeodb.stri.si.edu/bocas_database/search/species/332 (Accessed 21 November 2019).
Subba Rao NV & Dey A (2000) Catalogue of the marine molluscs of Andaman and Nicobar Islands. Records of the
Zoological Survey of India, Occasional Paper, 187: 1–323.
Sukhsangchan C & Kaewchaichalearnkit C (2013) First captured record of a mimic octopus (Thaumoctopus
mimicus) in Thailand. Phuket Marine Biological Center Research Bulletin, 72: 55–59.
Sukhsangchan C, Meksumpun S, Nabhitabhata J & Segawa S (2009) Distribution, biology, and stomach contents of
paper nautilus (Argonauta hians) in the Andaman Sea. ScienceAsia, 35: 315–319.
Sukhsangchan C & Nabhitabhata J (2007) Embryonic development of muddy paper nautilus, Argonauta hians
Lightfoot, 1786, from Andaman Sea, Thailand. Kasetsart Journal (Natural Science), 41: 531–538.
Sukhsangchan C, Nabhitabhata J & Meksumpun S (2008) Notes on the behaviour of the female muddy argonaut,
Argonauta hians Lightfoot, 1786 in captivity. Phuket Marine Biological Center Research Bulletin, 69: 55–59.
Suvatti C (1937) A Check-List of Aquatic Fauna in Siam (Excluding Fishes). Bureau of Fisheries, Bangkok, 116 pp.
Suvatti C (1938) Molluscs of Siam. Bureau of Fisheries, Bangkok, 91 pp.
Suwanmala J, Byern J von & Nabhitabhata J (2006) Observations of Idiosepius pygmaeus (Cephalopoda,
Idiosepiidae), the pygmy squid, at Klong Bangrong, Phuket Island, Thailand. Phuket Marine Biological Center
Research Bulletin, 67: 49–51.
Suzuki T, Nishihira M & Paphavasit N (2002) Size structure and distribution of Ovassiminea brevicula (Gastropoda)
in a Thai mangrove swamp. Wetlands Ecology and Management, 10: 265–271.
Swennen C (1996) Gymnodoris pattani, a new dorid nudibranch from Pattani Bay, Gulf of Thailand (Gastropoda,
Nudibranchia). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 15: 41–46.
Swennen C (1998) Two new gastropods, Elysia bangawaensis and E. siamensis from southern Thailand
(Opisthobranchia, Sacoglossa, Elysiidae). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 16: 33–40.
Swennen C (2001) Two new sacoglossans (Gasteropoda; Opisthobranchia) from Thailand. Beaufortia, 51: 75–81.
Swennen C (2007) Costasiella coronata, new species, and a revised diagnosis for the family Costasiellidae
(Mollusca: Opisthobranchia: Sacoglossa). Raffles Bulletin of Zoology, 55: 355–362.
Swennen C (2011) Large mangrove-dwelling Elysia species in Asia, with descriptions of two new species
(Gastropoda: Opistobranchia: Sacoglossa). Raffles Bulletin of Zoology, 59: 29–37.
Swennen C & Buatip S (2009) Aiteng ater, new genus, new species, an amphibious and insectivorous sea slug that
is difficult to classify [Mollusca: Gastropoda: Opisthobranchia: Sacoglossa(?): Aitengidae, new family]. Raffles
Bulletin of Zoology, 57: 495–500.
Swennen C & Buatip S (2012) Murphydoris singaporensis (Gastropoda: Nudibranchia: Goniodorididae) found in
Thailand; with external and first internal description of the specimens. Strombus, 19: 9–14.
Swennen C, Moolenbeek RG, Ruttanadakul N, Hobbelink H, Dekker H & Hajisamae S (2001) The molluscs of the
southern Gulf of Thailand. Thai Studies in Biodiversity, 4: 1–210.
Takenouchi K (1985) An analysis of shell character and distribution of the intertidal trochid, Monodonta labio
(Linné) (Gastropoda: Prosobranchia). Venus, 44: 110–122.
Tan KS & Goh L (eds.) (2016) The Comprehensive Marine Biodiversity Survey: Singapore Strait International
Workshop (2013). The Raffles Bulletin of Zoology, Supplement 34: 1–757.
Tan KS & Morton B (2008) The invasive Caribbean bivalve Mytilopsis sallei (Dreissenidae) introduced to Singapore
and Johor Bahru, Malaysia. Raffles Bulletin of Zoology, 54: 429–434.
Tan KS & Sigurdsson JB (1996) New species of Thais (Neogastropoda, Muricidae) from Singapore, with a redescription of Thais javanica (Philippi, 1848). Journal of Molluscan Studies, 62: 517–535.
Tan SK & Low MEY (2014) Singapore Mollusca: 5. The subfamily Planaxinae (Gastropoda: Caenogastropoda:
Cerithioidea: Planaxidae). Nature in Singapore, 7: 15–23.
Tan SK, Ng HE, Chan SY & Nguang LHS (2019) A review of the recent Agaronia Gray, 1839 (Caenogastropoda:
Olividae) of the Sundaic region, with description of a new species. Occasional Molluscan Papers, 7: 1–19.
Tan SK, Ng HE & Nguang LHS (2013) A new species of Phalium Link, 1807 (Gastropoda: Tonnoidea: Cassidae) from
the Sunda Shelf. The Raffles Bulletin of Zoology, 61: 507–514.
Tanamai S & Nabhitabhata J (2016) New records of marine cerithiid microgastropods (Prosobranchia: Cerithiidae)
from Thai waters. Phuket Marine Biological Center Research Bulletin, 73: 1–5.
Tantanasiriwong R (1978) An illustrated checklist of marine shelled gastropods from Phuket Island, adjacent
mainland and offshore island, western peninsular Thailand. Phuket Marine Biological Center Research Bulletin,
21: 1–63.
Tantanasiriwong R (1979) A checklist of marine bivalves from Phuket Island, adjacent mainland and offshore
island, western peninsular Thailand. Phuket Marine Biological Center Research Bulletin, 27: 1–15.
Taylor JD (2009) Mollusca. In: Paterson GLJ, Kendall MA & Aryuthaka C (eds.) Guide to the Common Offshore Fauna
of Laem Son National Park, Ranong, Thailand. Tsunami Impacts in Laem Song Project. Magic Publication,
Bangkok, pp. 56–76.
Taylor JD & Glover EA (2005) Cryptic diversity of chemosymbiotic bivalves: A systematic revision of worldwide
Anodontia (Mollusca: Bivalvia: Lucinidae). Systematics and Biodiversity, 3: 281–338.
182
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Taylor JD & Glover EA (2013) New lucinid bivalves from shallow and deeper water of the Indian and West Pacific
Oceans (Mollusca, Bivalvia, Lucinidae). ZooKeys, 326: 69–90.
Taylor JD, Williams ST, Glover EA & Dyal P (2007) A molecular phylogeny of heterodo nt bivalves (Mollusca:
Bivalvia: Heterodonta): New analyses of 18S and 28S rRNA genes. Zoologica Scripta, 36: 587–606.
Tëmkin I (2010) Molecular phylogeny of pearl oysters and their relatives (Mollusca, Bivalvia, Pterioidea). BMC
Evolutionary Biology, 10: 342.
Tëmkin I & Printrakoon C (2016) Morphology and taxonomy of Isognomon spathulatus (Reeve, 1858), a cryptic
bivalve from the mangroves of Thailand. Zootaxa, 4107: 141–174.
Tendal O, Aungtonya C, Bussarawit S (2014) Gear and sampling methods used during the Thai–Danish biodiversity
surveys 1996–2000 in the Andaman Sea. Phuket Marine Biological Center Special Publication, 32: 187–194.
Tenorio MJ, Poppe GT & Tagaro SP (2007) New Indo-Pacific Conidae with taxonomic and nomenclatural notes on
Conus recluzianus. Visaya, November 2007: 84–96.
Terranova SM, Lo Brutto S, Arculeo M & Mitton JB (2006) Population structure of Brachidontes pharaonis (P. Fisher,
1870) (Bivalvia, Mytilidae) in the Mediterranean Sea, and evolution of a novel mtDNA polymorphism. Marine
Biology, 150: 89–101.
Terryn Y (2007) A Collectors Guide to Recent Terebridae (Mollusca: Neogastropoda). ConchBooks, Hackenheim ;
Natural Art, Gent, Belgium, 57 pp.
Thamrongnawasawat T, Duangdee T & Sittithaweepat N (2008) Marine Mollusks in Thailand. Office of the Higher
Education Commission, Bangkok, 328 pp. [In Thai].
Tibiriçá Y & Malaquias MAE (2017) The bubble snails (Gastropoda, Heterobranchia) of Mozambique: An
overlooked biodiversity hotspot. Marine Biodiversity, 47: 791–811.
Torigoe K & Inaba A (2011) Revision on the classification of recent Naticidae. Bulletin of the Nishinomiya Shell
Museum, 7: 1–133.
Tuanapaya S & Nabhitabhata J (2016) Comparative external morphology of the pharaoh cuttlefish, Sepia pharaonis
Ehrenberg, 1831 from the Gulf of Thailand and Andaman Sea, of Peninsular Thailand. Phuket Marine Biological
Center Research Bulletin, 73: 15.
Tuaycharoen S (1999) Distribution of razor clams in the Gulf of Thailand. Phuket Marine Biological Center Special
Publication, 19: 173–176.
Tuaycharoen S & Matsukuma A (2001) Razor clams (Bivalvia: Solenidae) from the east and west coasts of Thailand.
Phuket Marine Biological Center Special Publication, 25: 377–386.
Tursch B, Germain L & Greifeneder D (1986) Studies on Olividae. III. Description of a novel subspecies: Oliva bulowi
phukentensis. Apex, 1: 71–87.
Tursch B & Greifeneder D (1996) The “Oliva miniacea complex”, with the description of a familiar, unnamed species
(Studies on Olividae. 25). Apex, 11: 1–49.
Tursch B & Machbaete Y (1995) The microstructure of the shell in the genus Oliva (Studies on Olividae. 24). Apex,
10: 61–78.
Tursch B, Missa O & Bouillon J (1992) Studies on Olividae. XVI. The taxonomic structure of Oliva oliva (auct.). Apex,
7: 3–22.
Ujino S & Matsukuma A (2013) Taxonomic revision of Serratina capsoides (Lamarck, 1818) (Bivalvia: Tellinidae),
an endangered species in Japan. Venus, 71: 175–189.
UNEP/GEF/SCS/RWG–SG.6/7 (2005) National Action Plan for Seagrass Thailand. Sixth Meeting of the Regional
Working Group for the Seagrass Sub-component of the UNEP/GEF Project: “Reversing Environmental
Degradation Trends in the South China Sea and Gulf of Thailand”, Bolinao, Philippines, 27th–30th September
2005. http://www.unepscs.org/remository/startdown/1670.html (Accessed 23 August 2020).
Upanoi T & Banchungmanee S (1999) Distribution and abundance of giant clams in the Adang-Rawi Island group,
the Andaman Sea, Thailand. Phuket Marine Biological Center Special Publication, 19: 243–251.
Valdés A & Barwick K (2005) First record of Akera Müller, 1776, from the eastern Pacific, with the description of
a new species. The Nautilus, 119: 43–49.
Valentich-Scott P & Dinesen GE (2004) Rock and coral boring Bivalvia (Mollusca) of the middle Florida Keys, USA.
Malacologia, 46: 339–354.
Valentich-Scott P & Tongkerd P (2008) Coral-boring bivalve molluscs of southeastern Thailand with the
description of a new species. Distribution of molluscs in mangroves at six sites in the upper Gulf of Thailand.
In: Bieler R, Chalermwat K, Mikkelsen P, Tan KS & Wells FE (eds.) Molluscs of Eastern Thailand. Proceedings
of the International Marine Bivalve Workshop, Chanthaburi, Thailand, August–September 2005, with
contributions on other molluscan groups. Raffles Bulletin of Zoology, Supplement 18: 191–216.
Vallejo B Jr, Conejar-Espedido J, Manubag L, Artiaga KCC, Damatac II AM, Imperial ICVJ, Itong TAB, Fontanilla IK &
Cao EP (2017) First record of the Charru mussel Mytella charruana d’Orbigny, 1846 (Bivalvia: Mytilidae) from
Manila Bay, Luzon, Philippines. Bioinvasions Records, 6: 49–55.
Vecchione M, Shea E, Bussarawit S, Anderson F, Alexeyev D, Lu CC, Okutani T, Roeleveld M, Chotiyaputta C, Roper
C, Jorgensen E & Sukramongkol N (2005) Systematics of Indo-West Pacific loliginids. Phuket Marine Biological
Center Research Bulletin, 66: 23–26.
183
The Marine and Estuarine Molluscs of Thailand
Verbinnen G, Segers L, Swinnen F, Kreipl K & Monsecour D (2016) Cassidae. An Amazing Family of Seashells.
ConchBooks, Harxheim, Germany, 251 pp.
Verhecken A (2011) The Cancellariidae of the PANGLAO Marine Biodiversity Project 2004 and the PANGLAO 2005
and AURORA 2007 deep sea cruises in the Philippines, with description of six new species (Neogastropoda,
Cancellarioidea). Vita Malacologica, 9: 1–60.
Vidal J (1993) Variability of Acrosterigma elongatum, a polytypic species (Mollusca, Cardiidae). Journal of the
Malacological Society of Australia, 14: 41–58.
Vidal J (1994) A review of the genus Fulvia Gray, 1853 (Mollusca, Cardiidae). Apex, 9: 93–118.
Vidal J (1997a) Large Trachycardiinae from the Indo-West Pacific: The group of Vasticardium orbita (Broderip &
Sowerby, 1833) (Mollusca, Cardiidae). Molluscan Research, 18: 11–32.
Vidal J (1997b) Taxonomic revision of the Indo-Pacific Vasticardium flavum species group (Bivalvia, Cardiidae).
Zoosystema, 19: 233–253.
Vidal J (1999a) Reflections on the distribution of polytypic species of the genus Vasticardium (Mollusca, Cardiidae)
from the Indo-Pacific. Phuket Marine Biological Center Special Publication, 19: 353–362.
Vidal J (1999b) Taxonomic review of the elongated cockles: Genera Trachycardium, Vasticardium and Acrosterigma
(Mollusca, Cardiidae). Zoosystema, 21: 259–335.
Vidal J (2000) Genus Vepricaurium Iredale, 1929 (Bivalvia, Cardiidae) with a description of a new species from
Thailand Vepricardium albohamatum Hylleberg & Vidal. Phuket Marine Biological Center Special Publication,
21: 447–464.
Vilvens C (2009) New species and new records of Calliostomatidae (Gastropoda: Trochoidea) from New Caledonia
and Solomon Islands. Novapex, 10: 125–163.
Vongpanich V (1996) The Arcidae of Thailand. Phuket Marine Biological Center Special Publication, 16: 177–192.
Vongpanich V (1999) A rectification of an erroneous record of Anadara mosambica (Bianconi, 1850) from the
Gulf of Thailand. Phuket Marine Biological Center Special Publication, 19: 395–396.
Vongpanich V (2000) Family Mactridae (Mollusca: Bivalvia) in Thai waters. Phuket Marine Biological Center
Special Publication, 21: 483–498.
Vongpanich V (2001) The order Thecosomata (Mollusca: Subclass Opisthobranchia) in the Andaman Sea. Phuket
Marine Biological Center Special Publication, 25: 493–500.
Vongpanich V & Matsukuma A (2004) Family Noetiidae in Thailand. Phuket Marine Biological Center Research
Bulletin, 65: 31–44.
Vos C (2007) Family Tonnidae. A Conchological Iconography. ConchBooks, Hackenheim, 188 pp.
Voskuil RPA & Onverwagt WJH (1991) Studies on Cardiidae. 4. The taxonomy of the genus Trachycardium (Part 1)
with descriptions of three new species. Vita Marina, 42: 56–72.
Voss NA, Vecchione M, Toll RB & Sweeney MJ (1998) Systematics and biology of cephalopod s. Smithsonian
Contributions to Zoology, 586: 1–276.
Wakefield A (2010) A revision of the genus Cryptospira Hinds, 1844 (Caenogastropoda: Marginellidae). Novapex,
11: 1–55.
Wangkulangkul K (2018) Comments on restricted distribution of Mytilopsis adamsi Morrison, 1946, a non-native
false mussel in the Songkhla Lagoon System, southern Thailand. Limnology, 19: 151–156.
Wangkulangkul K & Lheknim V (2008) The occurrence of an invasive alien mussel Mytilopsis adamsi Morrison,
1946 (Bivalvia: Dreissenidae) in estuaries and lagoons of the lower south of the Gulf of Thailand with comments
on their establishment. Aquatic Invasions, 3: 325–330.
Wangkulangkul K & Lkheknim V (2010) Variability in recruitment in an introduced mussel, Mytilopsis adamsi
Morrison, 1946, in Haad-kaew Lagoon, Songkhla Lake Basin, South Thailand (Abstract). World Congress of
Malacology, Phuket, Thailand 18−24 July 2010. Tropical Natural History, Supplement 3: 84.
Warén A (1981) Revision of the genera Apicalia A. Adams and Stilapex lredale and description of two new genera
(Mollusca, Prosobranchia, Eulimidae). Zoologica Scripta, 10: 133–154.
Wattayakorn G (2006) Environmental issues in the Gulf of Thailand. In: Wolanski E (ed.) The Environment in Asia
Pacific Harbours. Springer, Dordrecht, pp. 249–259.
Way K & Purchon RD (1981) The marine shelled Mollusca of West Malaysia and Singapore: Part 2. Polyplacophora
and Gastropoda. Journal of Molluscan Studies, 47: 313–321.
Weil A, Brown L & Neville B (1999) The Wentletrap Book. Guide to the Recent Epiton iidae of the World. Arti
Grafiche La Moderna, Rome, 244 pp.
Wells FE (2005) Molluscs of the Phuket region, Thailand. In: Allen GR & Stone GS (eds.) Rapid Assessment Survey
of Tsunami-Affected Reefs of Thailand. Final Technical Report. New England Aquarium, Boston, pp. 57–68,
111–119.
Wells FE, McDonald JI & Huisman JM (2009) Introduced Marine Species in Western Australia. WA Department of
Fisheries, Perth, 97 pp.
Wells FE, Walker D & Jones DS (2003) The Marine Flora and Fauna of Dampier, Western Australia. Western
Australian Museum, Perth, 587 pp.
Willan RC (1993) Taxonomic revision of the family Psammobiidae (Bivalvia: Tellinoidea) in the Australian and
New Zealand region. Records of the Australian Museum, Supplement 18: 1–132.
184
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Willan RC, Liu LL & Chiu YW (2007) Review of recent Psammobiidae (Bivalvia: Tellinoidea) from Taiwan. Journal
of the Fisheries Society of Taiwan, 34: 397–412.
Willan RC & Poppe P (2010) Doridomorphidae & Tritoniidae. In: Poppe GT. Philippine Marine Mollusks, Volume
III. ConchBooks, Hackenheim, p. 392, pl. 887.
Willan RC, Russell BC, Murfet NB, Moore KL, McEnnulty FR, Horner SK, Hewitt CL, Dally GM, Campbell ML &
Bourke ST (2000) Outbreak of Mytilopsis sallei (Récluz, 1849) (Bivalvia: Dreissenidae) in Australia. Molluscan
Research, 20: 25–30.
Wilson BR (1993–1994) Australian Marine Shells. Prosobranch Gastropods. Odyssey Publishing, Kallaroo,
Western Australia, 778 pp.
Wium-Andersen G (1977) Marine Nerita species from Phuket and their chromosome numbers. Phuket Marine
Biological Center Research Bulletin, 15: 1–9.
WoRMS Editorial Board (2019) World Register of Marine Species. Flanders Marine Institute, Belgium.
http://www.marinespecies.org (Accessed 23 August 2020).
Yang H, Sturmer LN & Baker S (2019) Molluscan Shellfish Aquaculture and Production. Publication #FA191.
University of Florida IFAS Extension. Electronic Data Information Source. https://edis.ifas.ufl.edu/fa191
(Accessed 25 May 2019).
Yeemin T, Sudara S, Krairapanond N, Silpsoonthorn C, Ruengsawang N & Asa S (2001) Country Report: Thailand.
International Coral Reef Initiative (ICRI). Regional ICRI Workshop for East Asia, Cebu, Philippines, 13 pp.
http://www.reefbase.org/resource_center/publication/main.aspx?refid=12388 (Accessed 23 August 2020).
Yonow N (1996) Systematic revision of the family Phyllidiidae in the Indian Ocean province: Part 1
(Opisthobranchia: Nudibranchia: Doridoidea). Journal of Conchology, 35: 483–516.
Yonow N (2011) Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 15. The suborder
Doridina (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia). Zoologische Mededelingen, 85: 905–956.
Yonow N (2012) Opisthobranchs from the western Indian Ocean, with descriptions of two new species and ten
new records (Mollusca, Gastropoda). ZooKeys, 197: 1–129.
Yoosukh W & Duangdee T (1999) Living oyster in Thailand. Phuket Marine Biological Center Special Publication,
19: 363–370.
Yoosukh W & Jitkaew M (1997) Marine wood-boring bivalves (Pholadacea) in the Gulf of Thailand. Phuket Marine
Biological Center Special Publication, 17: 401–405.
Yoosukh W & Matsukuma A (2001) Taxonomic study on Meretrix (Mollusca: Bivalvia) from Thailand. Phuket
Marine Biological Center Special Publication, 25: 451–460.
Yoosukh W & Sukhsangchan C (2000) Electrophoretic studies on tuberculated oysters from Ranong Province,
Thailand. Phuket Marine Biological Center Special Publication, 21: 477–481.
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ACKNOWLEDGEMENTS
We are pleased to acknowledge the extensive work undertaken on the marine and estuarine molluscs of Thailand
by countless researchers over many years. The present book is a summary of their work. John Taylor of the Natural
History Museum, London, kindly examined an earlier version of the lucinid section and added unpublished results
from a survey of marine molluscs in Ranong. Andreia Salvador of the Natural History Museum, London, provided
access to the museum’s extensive molluscan library. In Perth, Lisa Kirkendale and Corey Whisson of the Western
Australian Museum also provided access to important references. Three reviewers made excellent comments on
the draft manuscript and we appreciate their assistance. Ng Ting Hui and Cai Peixuan of the National University of
Singapore provided essential advice in preparing the manuscript for publication.
HOW TO CITE THIS BOOK
Cite this book as:
Wells FE, Sanpanich K, Tan SK & Duangdee T (2021) The Marine and Estuarine Molluscs of Thailand. Lee Kong
Chian Natural History Museum, National University of Singapore, Singapore, 195 pp. http://doi.org/10.26107/
LKCNHM-EBOOK-2021-0001 (Accessed <date>).
186
F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
APPENDIX
Appendix 1. Species recorded in the literature from Thailand that are rejected or are considered to be doubtful.
Class Bivalvia
Family Anatinellidae
Raeta meridionalis Tate, 1889. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
(2011). Justification: Geography. Harry (1969) lists this species as being of uncertain status, but it is a southern
Australian species, with a type locality of Middle Harbour, South Australia.
Family Arcidae
Anadara trapezia (Deshayes, 1840). Gulf of Thailand. Reported by: Nabhitabhata (2009). Justification: Geography.
An eastern Australian species (ALA, 2020; AMS, 2020).
Mosambicarca mosambicana (Bianconi, 1856). Reported as: Anadara mosambicana (Bianconi, 1850). Gulf
of Thailand. Reported by: Vongpanich (1996), Aungtonya et al. (1999), Nabhitabhata (2009). Justification:
Taxonomy. Vongpanich (1999) corrected the identification to Anadara sp. but did not identify it to species.
Scaphula pinna Benson, 1856. Gulf of Thailand, Andaman Sea. Reported by: Vongpanich (1996), Carpenter &
Niem (1998), Sanpanich (1998), Aungtonya et al. (1999), Robba et al. (2002, 2003), Wells (2005), Kasetsart
University (2006), Duangdee (2007), Printrakoon et al. (2008), Nabhitabhata (2009), Galli (2015). Justification:
WoRMS Editorial Board (2019) lists this as a freshwater species.
Family Donacidae
Donax acuticarinatus Sowerby, 1866. Gulf of Thailand. Reported by: Bertin (1881), Serène (1937), Suvatti
(1938), Nabhitabhata (2009). Justification: Geography. A junior synonym of the eastern Pacific Donax carinatus
Hanley, 1843 (Signorelli & Printrakoon, 2020).
Donax variabilis Say, 1822. Andaman Sea. Reported by: BEDO (2017a). Justification: Geography. East coast of the
United States (Mikkelsen, 1985; Adamkewicz, 1989).
Family Galeommatidae
Scintilla dubia (Deshayes, 1856). Also reported as: Scintilla duia (Deshayes, 1856). Andaman Sea. Reported by:
Lützen & Nielsen (2005), Hylleberg & Aungtonya (2013), BEDO (2017a). Justification: Geography. Coan & Scott
(2012) noted that the record from Phuket by Lützen & Nielsen (2005) is probably in error. The holotype from
Ecuador is figured in Coan & Scott (2012).
Family Lucinidae
Anodontia edentula (Linnaeus, 1758). Also reported as: Lucina (Thyasira) edentula Linnaeus, 1758. Gulf of
Thailand, Andaman Sea. Reported by: Lynge (1909), Nateewathana et al. (1981), Bussarawit (1995), Carpenter
& Niem (1998), Aungtonya et al. (1999), Swennen et al. (2001), Robba et al. (2002), Duangdee (2004), Duangdee
et al. (2007, 2009, 2011), Nabhitabhata (2009), BEDO (2017a). Justification: Taxonomy. Taylor & Glover (2005)
recommended abandoning the use of Anodontia edentula as there is no Linnean type and the description is
brief and could be any species. Most references are to Pegophysema philippiana (Reeve, 1850).
Anodontia pila (Reeve, 1850). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), BEDO (2017a).
Justification: Taxonomy. Anodontia pila was synonymised with the Atlantic Anodontia alba Link, 1807, and the
species is actually Cryptophysema vesicula (Gould, 1850) (see Taylor & Glover, 2005).
Cardiolucina crassilirata (Tate, 1887). Reported as: Linga crassilirata (Tate, 1887). Gulf of Thailand, Andaman
Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011), Duangdee et al. (2013b). Justification: Geography.
A southern Australian species (ALA, 2019).
Ctena gunnamatta (Iredale, 1930). Reported as: Epicodakia gunnamatta Iredale, 1930. Andaman Sea. Reported
by: Nabhitabhata (2009), Duangdee et al. (2011), Sanpanich (2011). Justification: Geography. The type locality
for this species is Gunnamatta Bay in southern Sydney, New South Wales, Australia, at a latitude of 38°S, a
temperate region. ALA (2019) does not have any records of the species.
Epicodakia consettiana Iredale, 1930. Reported as: Codakia minima Tension Woods, 1875. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009). Justification: Geography. Codakia minima Tension Woods,
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1875, was described from Tasmania. Iredale (1930) provided a new name, Epicodakia consettiana Iredale,
1930, when recording the species from New South Wales. It is a southern Australian species (ALA, 2019).
Lucinoma heroica (Dall, 1901). Andaman Sea. Reported by: BEDO (2017a). Justification: Geography. A deepwater
species from the Pacific coast of Mexico (J. Taylor, pers. comm.).
Family Mactridae
Mactra pura Reeve, 1854. Gulf of Thailand. Reported by: BEDO (2017a). Justification: Geography. A southeastern
Australian species (MRGV, 1984).
Mactra cf. westralis Lamprell & Whitehead, 1990. Gulf of Thailand. Reported by: Gemert (2003). Justification:
Geography. Endemic to Western Australia (Lamprell & Whitehead, 1990).
Family Mesodesmatidae
Paphies australis (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017a). Justification: Geography. Endemic
to New Zealand (ALA, 2020).
Family Mytilidae
Botula fusca (Gmelin, 1791). Gulf of Thailand. Reported by: GBIF (2019b). Justification: Geography. A Florida Keys
species (Valentich-Scott & Dinesen, 2004).
Brachidontes pharaonis (Fischer, 1870). Gulf of Thailand. Reported by: Swennen et al. (2001), Nabhitabhata
(2009). Justification: Taxonomy. Brachidontes pharaonis is native to the Red Sea. The Thai references are likely
to be a misidentification of Brachidontes variabilis (Krauss, 1848).
Brachidontes rostratus (Dunker, 1857). Andaman Sea. Reported by: Nateewathana et al. (1981), BEDO (2017a).
Justification: Geography. A southeastern Australian species (ALA, 2020).
Family Mytilidae
Lithophaga antillarum (d’Orbigny, 1853). Reported as: Lithophaga straminea Dunker, 1880. Gulf of Thailand.
Reported by: Lynge (1909), Nabhitabhata (2009). Justification: Geography. Occurs in the Caribbean Sea and
Gulf of Mexico (WoRMS Editorial Board, 2019).
Modiolaria opifex Philippi, 1841. Gulf of Thailand. Reported by: Lynge (1909), Nabhitabhata (2009). Justification:
Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019). It is listed in Kleeman (1983), but
the only reference is Lynge (1909).
Musculus discors (Linnaeus, 1767). Gulf of Thailand. Reported by: GBIF (2019b). Justification: Geography. Occurs
on both sides of the North Atlantic and along the North Pacific coast of North America (OBIS, 2020).
Musculus varicosus (Gould, 1861). Gulf of Thailand. Reported by: Nabhitabhata (2009), Sanpanich (2011).
Justification: Geography. A southeastern Australian species (ALA, 2020).
Family Noetidae
Striarca lamyi Koperberg, 1931. Reported as: Arca lamyi (Koperberg, 1931). Andaman Sea. Reported by: BEDO
(2017a). Justification: Cannot be confirmed through WoRMS Editorial Board (2019); it is shown as a fossil by
Fossilworks (2020).
Family Ostreidae
Crassostrea rhizophorae (Guilding, 1828). Andaman Sea. Reported by: BEDO (2017a). Justification: Geography.
Occurs in the Caribbean Sea and Gulf of Mexico (WoRMS Editorial Board, 2019).
Dendostrea frons (Linnaeus, 1758). Gulf of Thailand. Reported by: Gemert (2003). Justification: Geography. Occurs
in the Caribbean Sea and Gulf of Mexico (WoRMS Editorial Board, 2019; STRI, 2019).
Ostrea edulis (Linnaeus, 1758). Gulf of Thailand. Reported by: Suvatti (1937, 1938), Duangdee et al. (2011).
Justification: Geography. Status uncertain. Not listed by Bussarawit & Cedhagen (2010) in their revision of Thai
oysters. However, the species has been introduced for aquaculture in a number of countries and Yang et al.
(2019) list a production of four tonnes per year in Thailand.
Family Pectinidae
Delectopecten vitreus (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017a). Justification: Geography. North
Atlantic species (GBIF, 2020).
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Mesopeplum fenestratum (Hedley, 1901). Gulf of Thailand. Reported by: Nabhitabhata (2009), Sanpanich
(2011). Justification: Geography. Southern Australian species (Dijkstra & Beu, 2018; ALA, 2020).
Mimachlamys funebris (Reeve, 1853). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich (2011). Justification: Geography. Northern Australia (Dijkstra & Beu, 2018; ALA, 2020).
Mimachlamys inuate (Linnaeus, 1758). Andaman Sea. Reported by: Nabhitabhata (2009). Justification:
Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019), or other websites.
Mimachlamys scabricostata (G. B. Sowerby III, 1915). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich
(2011). Justification: Geography. Northern Australia (Dijkstra & Beu, 2018; ALA, 2020).
Mirapecten tuberosus Dijkstra & Kilburn, 2001. Andaman Sea. Reported by: GBIF (2019a). Justification:
Geography. Western Indian Ocean (Dijkstra & Kilburn, 2001).
Pecten fumatus Reeve, 1852. Andaman Sea. Reported by: BEDO (2017a). Justification: Geography. Southern
Australia, Tasman Sea and New Zealand (Dijkstra & Marshall, 1997; Dijkstra & Beu, 2018).
Talochlamys pulleineana (Tate, 1887). Reported as: Mimachlamys famigerator (Iredale, 1925). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich (2011). Justification: Geography. Southern
Australian species (Dijkstra & Beu, 2018; ALA, 2019).
Family Pholadidae
Barnea australasiae (G. B. Sowerby II, 1849). Reported as: Pholas australasiae Sowerby, 1849. Gulf of Thailand.
Reported by: Morlet (1889), Lynge (1909), Suvatti (1938), Nabhitabhata (2009). Justification: Geography.
Southern Australian species (ALA, 2019).
Pholas dactylus Linnaeus, 1758. Andaman Sea. Reported by: Kasetsart University (2006). Justification: Geography.
Northwestern Atlantic species (WoRMS Editorial Board, 2019).
Family Psammobiidae
Asaphis deflorata (Linnaeus, 1758). Also reported as: Venus deflorata Linnaeus, 1758. Gulf of Thailand, Andaman
Sea. Reported by: Lynge (1909), Suvatti (1938), Aungtonya et al. (1999), Nabhitabhata (2009), BEDO (2017a).
Justification: Geography. Caribbean species. It is likely all records refer to Asaphis violascens (Willan, 1993).
Gari modesta (Deshayes, 1855). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich (2011). Justification:
Geography. Southern Australia (ALA, 2019).
Plebidonax deltoides (Lamarck, 1818). Reported as: Donax deltoides Lamarck, 1818. Andaman Sea. Reported by:
Carpenter & Niem (1998), Nabhitabhata (2009). Justification: Geography. Southeastern Australia (MRGV, 1984).
Psammosolen esaratus Philippi. Gulf of Thailand. Reported by: Suvatti (1938), Nabhitabhata (2009). Justification:
Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019) or other internet sites.
Solenotellina emingina Deshayes. Gulf of Thailand. Reported by: Suvatti (1938), Nabhitabhata (2009). Justification:
Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019) or other internet sites.
Family Tellinidae
Pseudarcopagia botanica Hedley, 1918. Gulf of Thailand. Reported by: Sanpanich (2011). Justification: Geography.
Southern Australia (ALA, 2019).
Family Veneridae
Dosinia victoriae (Gatliff & Gabriel, 1914). Gulf of Thailand, Andaman Sea. Reported by: Kasetsart University
(2006), Duangdee (2007), Nabhitabhata (2009). Justification: Geography. Southern Australia (Cotton, 1961).
Placamen placidum (Philippi, 1844). Andaman Sea. Reported by: GBIF (2019a). Justification: Geography.
Southeastern Australia (ALA, 2020).
Polititapes aureus (Gmelin, 1791). Andaman Sea. Reported by: BEDO (2017a). Justification: Geography.
Northeastern Atlantic Ocean, Mediterranean Sea and Black Sea (OBIS, 2020).
Paphia (Ruditapes) gibba Gmelin, 1791. Gulf of Thailand. Reported by: Suvatti (1937), Nabhitabhata (2009).
Justification: Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019) or an alternative method.
Paphia (Ruditapes) indica Gmelin, 1791. Gulf of Thailand. Reported by: Suvatti (1937, 1938), Nabhitabhata (2009).
Justification: Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019), but WoRMS Editorial
Board (2019) does list Tapes indica G. B. Sowerby II, 1852, accepted as Ruditapes philippinarum (Adams & Reeve,
1850).
Venus dysira Chemnitz. Gulf of Thailand. Reported by: Suvatti (1938), Nabhitabhata (2009). Justification:
Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019).
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Family Vesicomyidae
Calyptogena magnifica Boss & R. D. Turner, 1980. Andaman Sea. Reported by: BEDO (2017a). Justification:
Geography. Galapagos Rift bivalve (Reid, 1990).
Class Cephalopoda
Family Enteroctopodidae
Enteroctopus dofleini (Wülker, 1910). Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. This
is a temperate North Pacific species. No record in the tropics or the tropical Indian Ocean (Norman et al., 2016).
Family Idiosepiidae
Alloteuthis subulata (Lamarck, 1798). Gulf of Thailand. Reported by: GBIF (2019b). Justification: Geography.
East Atlantic species (GBIF, 2020).
Idiosepius paradoxus (Ortmann, 1888). Andaman Sea. Reported by: GBIF (2019a). Justification: Geography.
Asian North Pacific (Sato et al., 2009).
Family Loliginidae
Lolliguncula (Lolliguncula) panamanensis Berry, 1911. Gulf of Thailand. Reported by: Nabhitabhata (2009),
BEDO (2017d), Nabhitabhata et al. (2019). Justification: Geography. West coast of Central America (GBIF, 2019).
Family Octopodidae
Abdopus cf. abaculus (Norman & Sweeney, 1997). Reported as: Octopus cf. abaculus Norman & Sweeney, 1997.
Andaman Sea. Reported by: Aungtonya et al. (2014). Justification: Geography. Known only from its type locality,
the Philippines (Norman et al., 2016).
Family Ommastrephidae
Dosidicus gigas (d’Orbigny [in 1834–1847], 1835). Andaman Sea. Reported by: GBIF (2019a). Justification:
Geography. East Pacific species (GBIF, 2020). Imported frozen from South America for canning and re-exported.
They are also sold in many Thai supermarkets.
Family Sepiadariidae
Sepioloidea lineolata (Quoy & Gaimard, 1832). Andaman Sea. Reported by: GBIF (2019a). Justification: Geography.
Australian species (ALA, 2020).
Family Sepiidae
Onykia carriboea Lesueur, 1821. Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017d), Nabhitabhata et al.
(2019). Justification: Geography. Gulf of Mexico, Northeastern Atlantic Ocean (Spain and Portugal) and the
Mediterranean coast of Spain (WoRMS Editorial Board, 2019).
Sepia apama Gray, 1849. Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. Southern Australia
(ALA, 2020).
Sepia officinalis Linnaeus, 1758. Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. East Atlantic
and Mediterranean Sea (GBIF, 2020).
Sepia smithi Hoyle, 1885. Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. Northern Australia
(ALA, 2020).
Sepiella ornata (Rang, 1837). Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. East Atlantic
(GBIF, 2020).
Class Gastropoda
Family Aeolidiidae
Cerberilla incola Burn, 1974. Gulf of Thailand, Andaman Sea. Reported by: Chavanich et al. (2013). Justification:
Geography. Southeastern Australia (ALA, 2019).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Family Amphibolidae
Salinator fragilis (Lamarck, 1822). Gulf of Thailand. Reported by: Negri et al. (2014). Justification: Geography.
Southern Australia (GBIF, 2020).
Family Calyptraeidae
Calyptraea scabris. No Thai locality provided. Reported by: THNHM. Justification: Taxonomy. Listed by THNHM
without author and date, this species could not be verified through WoRMS Editorial Board (2019) and is not
shown on any other websites.
Family Cassidae
Semicassis faurotis (Jousseaume, 1888). Reported as: Phalium faurotis. No Thai locality provided. Reported by:
THNHM. Justification: Geography. Western Indian Ocean (Abbott, 1968).
Family Cerithiidae
Cerithium corallinum G. B. Sowerby II, 1855. Gulf of Thailand. Reported by: Morlet (1889). Justification: Taxonomy.
Cannot be confirmed through WoRMS Editorial Board (2019) or Ruhoff (1980). Possibly a misspelling of
Cerithium coralium Kiener, 1841.
Family Chilodontaidae
Herpetopoma aspersum (Philippi, 1846). Gulf of Thailand, Andaman Sea. Reported by: Duangdee et al. (2011),
Sanpanich & Duangdee (2013). Justification: Geography. Southern Australia (Wilson, 1993).
Hybochelus triangularus A. Adams. Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al.
(1981), BEDO (2017b). Justification: Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019)
or Ruhoff (1980).
Family Chromodorididae
Goniobranchus rufomaculatus (Pease, 1871). Reported as: Chromodoris histrio Bergh, 1878. Andaman Sea.
Reported by: Jensen (2007), Nabhitabhata (2009). Justification: Taxonomy. Jensen (2007) states the Thai
specimens are either Chromodoris tumulifera Collingwood, 1881, or Chromodoris bimaensis Bergh, 1905.
Family Conidae
Conus senator Linnaeus, 1758. Gulf of Thailand. Reported by: Morlet (1889). Justification: Taxonomy. WoRMS
Editorial Board (2019) states: “unaccepted (Nomenclaturally invalid name, placed on the Official Index of
Invalid and Rejected Specific Names in Zoology (ICZN Opinion n°753)”.
Conus victoriae Reeve, 1843. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2013).
Justification: Geography. Endemic to northwestern Australia (Wilson, 1993).
Family Costellariidae
Vexillum japonicum A. Adams, 1864. Andaman Sea. Reported by: BEDO (2017b). Justification: Geography. Japan
(GBIF, 2020).
Family Cypraeidae
Notocypraea comptonii (Gray, 1847). Gulf of Thailand. Reported by: Sanpanich & Duangdee (2009, 2013).
Justification: Geography. Temperate Australia (Wilson, 1993).
Notocypraea pulicaria (Reeve, 1846). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2009, 2013). Justification: Geography. Temperate Australia (Wilson, 1993).
Family Dendrodorididae
Dendrodoris senegalensis Bouchet, 1975. Gulf of Thailand, Andaman Sea. Reported by: Thamrongnawasawat
et al. (2008), Nabhitabhata (2009). Justification: Geography. West coast of Africa (Bouchet, 1975).
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The Marine and Estuarine Molluscs of Thailand
Family Drilliidae
Paradrillia serena (P. J. Fischer, 1927). Gulf of Thailand. Reported by: Gemert (2003). Justification: Fossil species
(Leloux & Wesselingh, 2009).
Family Epitoniidae
Boreoscala blainei (Clench & R. D. Turner, 1953). Andaman Sea. Reported by: BEDO (2017b). Justification:
Geography. Florida (Rosenberg et al., 2009).
Family Eulimidae
Sticteulima wareni Engl, 1997. Andaman Sea. Reported by: GBIF (2019c). Justification: Geography. Described
from the Canary Islands in the Atlantic Ocean (Engl, 1997).
Family Fasciolariidae
Goniofusus spectrum (A. Adams & Reeve, 1848). Reported as: Fusinus spectrum (A. Adams & Reeve, 1848).
Andaman Sea. Reported by: GBIF (2019a). Justification: Geography. Goniofusus spectrum is a Panamic species
(Callomon & Snyder, 2008). The Thai species is possibly Fusinus stannum Callomon & Snyder, 2008.
Latirus pulchellus (Reeve, 1847). Gulf of Thailand. Reported by: Gemert (2003). Justification: Taxonomy.
Could not be verified through WoRMS Editorial Board (2019), but WoRMS Editorial Board (2019) lists the
Mediterranean Fusinus pulchellus (Philippi, 1840).
Family Ficidae
Ficus subintermedia (d’Orbigny, 1852). No Thai locality provided. Reported by: THNHM. Justification: Fossil
species (WoRMS Editorial Board (2018).
Family Haliotidae
Haliotis iris Gmelin, 1791. Andaman Sea. Reported by: Galli (2015). Justification: Geography. New Zealand
(Spencer et al., 2009).
Family Littorinidae
Echinolittorina australis (Gray, 1826). Also reported as: Nodilittorina nodosa (Gray, 1839). Gulf of Thailand,
Andaman Sea. Reported by: Nabhitabhata (2009), GBIF (2019a, 2019b). Justification: Geography. Endemic to
Western Australia (Wilson, 1993; Reid, 2006).
Echinolittorina feejeensis (Reeve, 1857). Gulf of Thailand, Andaman Sea. Reported by: Sanpanich et al. (2004),
Sanpanich & Duangdee (2013). Justification: Geography. Tropical Pacific centred on Fiji (Reid, 2006).
Echinolittorina miliaris (Quoy & Gaimard, 1833). Gulf of Thailand, Andaman Sea. Reported by: GBIF (2019b).
Justification: Geography. Reid (2007) discusses the complex use of the name Echinolittorina miliaris but
considers it to be endemic to Ascension Island in the Atlantic Ocean. Echinolittorina miliaris was not recorded
by Sanpanich et al. (2004).
Echinolittorina millegrana (Philippi, 1848). Reported as: Nodilittorina millegrana (Philippi, 1848). Andaman
Sea. Reported by: Rosewater (1970), Tantanasiriwong (1978), Nateewathana et al. (1981), Bussarawit (1995),
Subba Rao & Dey (2000), Nabhitabhata (2009), Kostina et al. (2016), BEDO (2017b), GBIF (2019a, 2019b).
Justification: Taxonomy. Rosewater (1970) considered Echinolittorina reticulata to be a synonym of
Echinolittorina millegrana, but Reid (2006) separated the two species, and restricted Echinolittorina millegrana
to the northwestern Indian Ocean and Red Sea. The Thai records of Echinolittorina millegrana follow Rosewater
(1970) and are probably all Echinolittorina reticulata.
Echinolittorina trochoides (Gray, 1839). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Swennen et al.
(2001), Duangdee (2004), Sanpanich et al. (2004), Wells (2005), ONR & EPP (2006), Reid (2006), Kasetsart
University (2007), Thamrongnawasawat et al. (2008), Nabhitabhata (2009). Justification: Taxonomy. Reid (2006)
discusses the complex use of the name Echinolittorina trochoides (Gray, 1839) and considers it to probably be
a synonym of Echinolittorina malaccana (Philippi, 1847).
Littoraria filosa (G. B. Sowerby I, 1832). Reported as: Littorina filosa G. B. Sowerby I, 1832. Gulf of Thailand.
Reported by: Morlet (1889). Justification: Geography. Reid (1986) shows the distribution of this species as
northern Australia, southern Indonesia and southern New Guinea. Littoraria filosa was not recorded by
Sanpanich et al. (2004).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Littoraria philippiana (Reeve, 1857). Gulf of Thailand. Reported by: Morlet (1889). Justification: Geography.
Reid (1986) shows the distribution of this species as the east coast of northern Australia and southern New
Guinea. Littoraria philippiana was not recorded by Sanpanich et al. (2004).
Family Marginellidae
Cryptospira glauca Jousseaume, 1875. Andaman Sea. Reported by: BEDO (2017b). Justification: Geography.
Cryptospira glauca is restricted to the Bay of Bengal (northeast Sri Lanka and east coast of India). The Thai
records are likely to be Cryptospira marchii (Wakefield, 2010).
Rivomarginella morrisoni Brandt, 1968. Gulf of Thailand. Reported by: Galli (2015). Justification: WoRMS
Editorial Board (2019) lists this as a freshwater species.
Family Muricidae
Ergalatax nodulosus (Pease, 1869). Andaman Sea. Reported by: Bussarawit (1991). Justification: Taxonomy.
Could not be confirmed through WoRMS Editorial Board (2019). Ruhoff (1980) lists Engina nodulosus Pease,
1869 as the only species with that species name described by Pease but this is a columbellid, not a muricid.
Purpura panama (Röding, 1798). Gulf of Thailand, Andaman Sea. Reported by: Carpenter & Niem (1998),
Nabhitabhata (2009). Justification: Geography. WoRMS Editorial Board (2019) shows this as a species from
the east coast of southern Africa.
Thais undata. Gulf of Thailand. Reported by: THNHM. Justification: Taxonomy. Unable to confirm through
WoRMS Editorial Board (2019). Possibly Purpura undata Lamarck, 1822, which is now recognised as Reishia
bitubercularis (Lamarck, 1822), which is present in Thailand.
Family Nacellidae
Cellana conciliata Iredale, 1940. Gulf of Thailand. Reported by: Sanpanich & Duangdee (2013). Justification:
Geography. Shown by Powell (1973) and Wilson (1993) as an eastern Australian species from Queensland.
Cellana grata (Gould, 1859). Gulf of Thailand, Andaman Sea. Reported by: KUMF, Duangdee (2004), Duangdee
et al. (2009), Nabhitabhata (2009), Sanpanich & Duangdee (2013). Justification: Geography. Shown by Powell
(1973) as a species from Japan and Korea.
Family Nassariidae
Nassarius arbicularis (Linnaeus). Andaman Sea. Reported by: Nateewathana et al. (1981). Justification:
Taxonomy. Unable to confirm through WoRMS Editorial Board (2019). Possibly a misspelling of Nassarius
arcularius (Linnaeus, 1758).
Tritia obsoleta (Say, 1822). Reported as: Ilyanassa obsoleta (Say, 1822). Gulf of Thailand, Andaman Sea. Reported
by: Nabhitabhata (2009), Sanpanich & Duangdee (2013). Justification: Geography. East coast of the United
States (GBIF, 2019).
Family Naticidae
Euspira yokoyamai (Kuroda & Habe, 1952). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2013). Justification: Geography. Japan (Torigoe & Inaba, 2011).
Natica luculenta Iredale, 1929. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich &
Duangdee (2013). Justification: Geography. Eastern Australia (Torigoe & Inaba, 2011).
Natica marochiensis (Gmelin, 1791). Reported as: Natica (Natica) luridis. Gulf of Thailand. Reported by: THNHM.
Justification: Geography. Western Atlantic (Florida to Brazil and eastern Atlantic ) (Rosenberg et al., 2009;
(Torigoe & Inaba, 2011).
Natica simplex G. B. Sowerby III, 1897. Reported as: Natica simplex (Röding, 1798). Gulf of Thailand. Reported
by: Nabhitabhata (2009). Justification: Geography. South Africa (Torigoe & Inaba, 2011).
Natica unifasciata Lamarck, 1822. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
& Duangdee (2009, 2013). Justification: Geography. Tropical West America (Torigoe & Inaba, 2011).
Notocochlis antoni (Philippi, 1851). Reported as: Natica antoni Philippi. Andaman Sea. Reported by:
Nateewathana et al. (1981). Justification: Geography. Mozambique (Torigoe & Inaba, 2011).
Polinices putealis Garrard, 1961. Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
& Duangdee (2013), BEDO (2017b). Justification: Geography. Eastern Australia (Torigoe & Inaba, 2011).
Polinices sagamiensis Pilsbry, 1904. Andaman Sea. Reported by: Nateewathana et al. (1981). Justification:
Geography. Japan (Torigoe & Inaba, 2011).
Sigatica bathyraphe (Pilsbry, 1911). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2013). Justification: Geography. Japan (Torigoe & Inaba, 2011).
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The Marine and Estuarine Molluscs of Thailand
Sinum concavum (Lamarck, 1822). Andaman Sea. Reported by: BEDO (2017b). Justification: Geography. West
Africa (Torigoe & Inaba, 2011).
Sinum japonicum (Lischke, 1872). Reported as: Sigaretus japonicum Lischke, 1872. Gulf of Thailand. Reported
by: Morlet (1889). Justification: Geography. Japan (Torigoe & Inaba, 2011).
Sinum maculatum (Say, 1831). Andaman Sea. Reported by: BEDO (2017b). Justification: Geography. Western
Atlantic (North Carolina to Brazil) (Torigoe & Inaba, 2011).
Sinum zonale (Quoy & Gaimard, 1832). Andaman Sea. Reported by: Nateewathana et al. (1981). Justification:
Geography. Southern Australia (Torigoe & Inaba, 2011; ALA, 2020).
Stigmaulax cayennensis (Récluz, 1850). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2013). Justification: Geography. Western Atlantic (West Indies to Brazil) (Torigoe &
Inaba, 2011).
Stigmaulax sulcatus (Born, 1778). No Thai locality provided. Reported by: Subba Rao & Dey (2000). Justification:
Geography. Western Atlantic (West Indies to Brazil) (Torigoe & Inaba, 2011).
Tanea zelandica (Quoy & Gaimard, 1832). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009),
Sanpanich & Duangdee (2013). Justification: Geography. South Pacific (New Caledonia to New Zealand)
(Torigoe & Inaba, 2011).
Family Neritidae
Vitta virginea (Linnaeus, 1758). No Thai locality provided. Reported by: Galli (2015). Justification: Geography.
Gulf of Mexico and the Caribbean Sea (WoRMS Editorial Board (2018).
Family Olividae
Agaronia ispidula (Linnaeus, 1758). Reported as: Agaronia plicaria Lamarck, 1811. Gulf of Thailand. Reported
by: Sanpanich & Duangdee (2013). Justification: Listed as a fossil by WoRMS Editorial Board (2019).
Oliva inspidula Fischer, 1807. Gulf of Thailand, Andaman Sea. Reported by: THNHM, Morlet (1889),
Nabhitabhata (2009). Justification: Taxonomy. Cannot be confirmed through WoRMS Editorial Board (2019)
or other websites. Possibly a misspelling of Oliva ispidula Linnaeus, 1758, not Fischer, 1807.
Family Patellidae
Scutellastra barbara (Linnaeus, 1758). Gulf of Thailand. Reported by: Sanpanich & Duangdee (2013). Justification:
Geography. A South African species (Powell, 1973).
Family Pleurobranchidae
Berthella africana Pruvot-Fol, 1953. Andaman Sea. Reported by: Thamrongnawasawat et al. (2008), Nabhitabhata
(2009). Justification: Geography. Northwest coast of Africa (WoRMS Editorial Board [2018]).
Family Pyramidellidae
Oscilla perfelix (Nomura, 1938). Gulf of Thailand, Andaman Sea. Reported by: BEDO (2017e). Justification:
Geography. Japan (Okutani, 2000; GBIF, 2020).
Family Pyrenidae
Euplica bidentata (Menke, 1843). Gulf of Thailand, Andaman Sea. Reported by: Nabhitabhata (2009), Sanpanich
& Duangdee (2013). Justification: Geography. Southwestern Australia (Wilson, 1994).
Family Terebridae
Duplicaria bernardii (Deshayes, 1857). Andaman Sea. Reported by: BEDO (2017b). Justification: Geography.
Northern Australia (Wilson, 1994).
Family Trochidae
Austrocochlea concamerata (Wood, 1828). No Thai locality provided. Reported by: THNHM. Justification:
Geography. Southern Australia (Wilson, 1993).
Austrocochlea constricta (Lamarck, 1822). Reported as: Austrocochlea obtusa. No Thai locality provided.
Reported by: THNHM. Justification: Geography. Southern Australia (Wilson, 1993).
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F. E. Wells, K. Sanpanich, S. K. Tan & T. Duangdee
Clanculus comarilis Hedley, 1912. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2013).
Justification: Geography. Northern Australia (Wilson, 1993; GBIF, 2020).
Clanculus consobrinus Tate, 1893. Gulf of Thailand, Andaman Sea. Reported by: Sanpanich & Duangdee (2009,
2013). Justification: Geography. Southern Australia (Wilson, 1993; GBIF, 2020).
Family Turbinellidae
Syrinx aruanus (Linnaeus, 1758). No Thai locality provided. Reported by: THNHM. Justification: Geography.
Northwestern Australia (Wilson, 1993) and eastern Indonesia (Dharma, 1988).
Family Turbinidae
Astralium rotularium (Lamarck, 1822). Gulf of Thailand. Reported as: Astraea rotularia. Reported by: THNHM.
Justification: Geography. Endemic to northern Australia (Wilson, 1993; GBIF, 2020).
Family Turridae
Gemmula hastula (Reeve, 1843). Andaman Sea. Reported by: BEDO (2017b). Justification: Geography. Persian
Gulf and Arabian Sea (Powell, 1964).
Gemmula monilifera (Pease, 1860). Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana et al.
(1981), BEDO (2017b). Justification: Taxonomy. Powell (1964) states that Gemmula gilchristi (Sowerby, 1902)
is frequently misidentified as Gemmula monilifera (Pease, 1860). Although he considers it to be a well-known
South African species, he does record Gemmula gilchristi from the Andaman Islands.
Family Turritellidae
Neohaustator fortilirata (G. B. Sowerby III, 1914). Andaman Sea. Reported by: BEDO (2017b). Justification:
Geography. Japan (WoRMS Editorial Board, 2019; GBIF, 2020).
Neohaustator tsushimaensis Kotaka, 1951. Andaman Sea. Reported by: Tantanasiriwong (1978), Nateewathana
et al. (1981), Bussarawit (1995), Nabhitabhata (2009), BEDO (2017b). Justification: Geography. Cannot be
confirmed through WoRMS Editorial Board (2019). It is shown as a Japanese species by GBIF (2020).
Turritella ligar Deshayes, 1843. Gulf of Thailand. Reported by: THNHM. Justification: Geography. West Africa
(Bernard, 1984).
Family Volutidae
Amoria damonii Gray, 1864. Andaman Sea. Reported by: Duangdee et al. (2013a). Justification: Geography.
Western Australia (Wilson, 1994).
Class Polyplacophora
Family Chitonidae
Acanthopleura vaillantii Rochebrune, 1882. Andaman Sea. Reported by: GBIF (2019a). Justification: Geography.
Middle East (Sadeghi & Loghmani, 2010).
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