Bryophytes – Mosses, Liverworts & Hornworts


An Introduction to Bryophytes: the Mosses, Liverworts and Hornworts


There are around 1100 species of bryophyte in Britain and Ireland. Mosses, liverworts and hornworts comprises a major group of land plants that are evolutionarily quite distinct from the more familiar vascular plants. Only in the last few years has it become apparent that the mon-vascular bryophytes are probably a single group, all more closely related to each other. They are among the earliest land plants and play significant roles in ecosystems worldwide. Bryophytes are characterized by their small size, lack of true roots, stems, and leaves, and their reliance on water for reproduction.


Classification

Kingdom:
Subkingdom:
Phylum:
Phylum:
Phylum:

Plantae (Plants)
Viridiplantae (Green Plants)
Marchantiophyta (Liverworts)
Bryophyta (Mosses)
Anthocerotophyta (Hornworts)


Phylum Marchantiophyta (Liverworts)


Liverworts belong to the division Marchantiophyta and are characterized by their flattened, ribbon-like thallus or leafy structures. Unlike mosses, liverworts exhibit a more primitive morphology and lack specialized conducting tissues. Liverworts reproduce via spores or asexual gemmae, small reproductive structures that detach and develop into new individuals. Liverworts are commonly found in moist environments such as wetlands, stream banks, and shaded forests. They play ecological roles in nutrient cycling and provide habitat for microorganisms and invertebrates. Some liverwort species are used in traditional medicine for their medicinal properties.


Order Marchantiales

Great Scented Liverwort (Conocephalum conicum)
Great Scented Liverwort
Conocephalum conicum
Marchantia polymorpha (Marchantia polymorpha) subsp ruderalis
Common Liverwort
Marchantia polymorpha subsp ruderalis

Order Lunulariales

Crescent-cup Liverwort (Lunularia cruciata)
Crescent-cup Liverwort Lunularia cruciata

Phylum Bryophyta (Mosses)


Mosses are small, herbaceous plants that belong to the division Bryophyta. They are characterized by their leafy structures, known as gametophytes, which lack true roots, stems, or vascular tissues. Mosses reproduce via spores and have a unique life cycle involving alternation of generations between gametophyte and sporophyte stages. The gametophyte is the dominant phase, consisting of leafy stems with structures that produce eggs and sperm. After fertilization, the sporophyte grows on the gametophyte and produces spores through meiosis. Mosses are found in diverse habitats, including forests, wetlands, and alpine regions, where they contribute to soil formation and moisture retention. They are also used in horticulture and landscaping for their ornamental value.

Mosses can be divided into two groups, acrocarps which grow upright, and pleurocarps which have a prostrate habit.


Acrocarps


Acrocarpous mosses have an upright growth habit and grow slower than Pleurocarpous mosses.


Order Polytrichales

Bank Haircap Moss (Polytrichum formosum)
Bank Haircap Moss Polytrichum formosum
Catherine’s Moss (Atrichum undulatum)
Catherine’s Moss Atrichum undulatum

Order Dicranales

Redshank Moss Ceratodon purpureus
Redshank Moss Ceratodon purpureus
Intermediate Screw-moss (Syntrichia intermedia)
Intermediate Screw-moss Syntrichia montana
Common Pincushion (Dicranoweisia cirrata)
Common Pincushion Dicranoweisia cirrata
Wall Screw-moss (Tortula muralis)
Wall Screw-moss Tortula muralis
Lesser Bird’s-claw Beard-moss (Barbula convoluta)
Lesser Bird’s-claw Beard-moss
Streblotrichum convolutum var. convolutum

Order Bryales

Smaller Lattice-moss (Cinclidotus fontinaloides)
Swan’s-neck Thyme-moss Mnium hornum
Capillary Thread-moss Bryum capillare
Capillary Thread-moss Bryum capillare
Silver Moss Bryum argenteum
Silver Moss Bryum argenteum

Order Grimmiales

Grey-cushioned Grimmia Grimmia pulvinata
Grey-cushioned Grimmia Grimmia pulvinata
Thickpoint Grimmia (Schistidium crassipilum)
Thickpoint Grimmia Schistidium crassipilum

Order Funariales

Bonfire-moss Funaria hygrometrica
Bonfire-moss Funaria hygrometrica

Order Orthotrichales

White-tipped Bristle-moss (Orthotrichum diaphanum)
White-tipped Bristle-moss
Orthotrichum diaphanum
Lyell's Bristle-moss (Orthotrichum lyellii)
Wood Bristle-moss
Lewinskya affinis

Pleurocarps


Pleurocapous mosses have a prostrate growth habit and grow quicker than Acrocarpous mosses.


Order Hypnales

Common Feather-moss (Kindbergia praelonga)
Common Feather-moss Kindbergia praelonga
Pointed Spear-moss (Calliergonella cuspidata)
Pointed Spear-moss
Calliergonella cuspidata
Springy Turf-moss (Rhytidiadelphus squarrosus)
Springy Turf-moss Rhytidiadelphus squarrosus
Rough-stalked Feather-moss Brachythecium rutabulum
Rough-stalked Feather-moss
Brachythecium rutabulum

All photographs copyright © Peter Hillman
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