*1: Trefusis Grove,
Falmouth, 1841, EAW
(TRU) (Paton 1969a:
696).
*2: St Breward, 1905,
RWS (TRU) (Paton 1969a:
696).
Grows on acidic soil (mineral, humic or peaty),
peat or peaty litter (e.g. on bases of Molinia caerulea or
fern tussocks), among living and moribund sphagna, thin soil
over rocks or occasionally on crumbling rocks; usually shaded,
sometimes in heavy shade (e.g. under overhanging banks, or
inside entrance to old mine adits), but also occurs in fully
insolated sites. Habitats include banks in deciduous and
conifer woodlands, swampy woodlands, groves of trees, Grey
Willow carr, scrub, old stone quarries, china clay pits and
spoil, banks beside roads, paths or tracks, banks on old mine
areas, on shaded 'hedges', on stream and ditch banks, on low
creek-side cliff, among granitic rocks, inside entrance to old
mine adit, peaty banks on heathland, mires, and above
sea-cliffs in rock-crevices or on banks and flushes.
Associates often include Calypogeia arguta, Cephalozia
bicuspidata, Diplophyllum albicans,
Lophocolea
bidentata, Mnium
hornum, Pellia
epiphylla, Pseudotaxiphyllum
elegans, Sphagnum
subnitens, less frequently Aneura pinguis, Calypogeia
muelleriana, Calypogeia
sphagnicola, Cephalozia connivens,
Cephalozia
lunulifolia, Cephaloziella
hampeana, Lophocolea fragrans,
Odontoschisma
sphagni, Plagiothecium denticulatum
var. denticulatum, Riccardia
chamedryfolia and Riccardia
latifrons.
Commonly with gemmae. Two records c.fr.: capsules
immature 5, dehiscing 4.